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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L979	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Myotis zenatius	Myotis zenatius	Myotis zenatius	N/A	Myotis zenatius	Myotis zenatius	Myotis zenatius	Myotis zenatius	Myotis zenatius		[HMW] Myotis zenatius Ibanez et al . in Juste et al, 2019 , “Mizou Cave, Tetouan, Moraceo (ca. 35°30' 42"N , 5°19'33"W ), at an altitude of 330 m a.s.1.” Subgenus Myotis ; myotis species group. See M. nattereri . The name M. escalerai cabrerae was recently proposed by E. Coraman and colleagues in 2019 based on populations that are now included under this species; the name is considered a nomen nudum as the publication did not include a proper description for the name. Monotypic.; [batnames2022] Çoraman et al. (2019) failed to meet the requirements for electronic publication of the name cabrerae (see Ruedi et al. 2019). Juste et al. (2019) electronically published the name zenatius for this species on 30 January 2019, satisfying all the ICZN (1999) requirements of electronic publication.; [MDD2022] recently described; includes the recently described name cabrerae, which is considered unavaialble due to publication in which it was described not adhering to the ICZN Code; [batnames2023] Çoraman et al. (2019) failed to meet the requirements for electronic publication of the name cabrerae (see Ruedi et al. 2019). Juste et al. (2019) electronically published the name zenatius for this species on 30 January 2019, satisfying all the ICZN (1999) requirements of electronic publication.; [MDD2023] recently described; includes the recently described name cabrerae, which is considered unavaialble due to publication in which it was described not adhering to the ICZN Code; [MDD2025_2.0] recently described; includes the recently described name cabrerae, which is considered unavailable due to publication in which it was described not adhering to the ICZN Code; [batnames2025_1.7] Çoraman et al. (2019) failed to meet the requirements for electronic publication of the name cabrerae (see Ruedi et al. 2019). Juste et al. (2019) electronically published the name zenatius for this species on 30 January 2019, satisfying all the ICZN (1999) requirements of electronic publication.; [MDD2025_2.2] recently described; includes the recently described name cabrerae, which is considered unavailable due to publication in which it was described not adhering to the ICZN Code												zenatius 	zenatius - cabrerae	zenatius, cabrerae		zenatius 	zenatius - cabrerae	zenatius, cabrerae	cabrerae, zenatius	zenatius	zenatius - cabrerae	cabrerae Çoraman, F. Mayer, & Dietz in Çoraman, Dietz, Hempel, Ghazaryan, Levin, Presetnik, Zagmajster, & F. Mayer, 2019 [nomen nudum]|zenatius Ibáñez, Juste, Salicini, Puechmaille, & Ruedi in Juste, Ruedi, Puechmaille, Salicini, & Ibáñez, 2019						N/A																																								NA																											4C3D87E8FF216A9EFA53922E1913B7B0	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	980	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF216A9EFA53922E1913B7B0.xml	Myotis zenatius	Vespertilionidae	Myotis	zenatius	Ibáñez et al.	2019	Murin zénéte @fr | Zenati-Mausohr @de | Ratonero zenate @es | Zenati Bat @en	Myotis zenatius Ibanez et al . in Juste et al, 2019 , “Mizou Cave, Tetouan, Moraceo (ca. 35°30' 42"N , 5°19'33"W ), at an altitude of 330 m a.s.1.” Subgenus Myotis ; myotis species group. See M. nattereri . The name M. escalerai cabrerae was recently proposed by E. Coraman and colleagues in 2019 based on populations that are now included under this species; the name is considered a nomen nudum as the publication did not include a proper description for the name. Monotypic.	NW Africa, including Morocco , Algeria ; also possibly in Tunisia .	Head—-body 44-48 mm , tail 38-42 mm , ear 15-3-17- 3 mm , hindfoot 7-9-8-7, forearm 38- 2—40 mm . The Zenati Myotis is very similar to the other species in the M. nattereri species complex, which makesits identification complicated. As in Escalera’s Myotis ( M. escalerai ), plagiopatagium is inserted in mid-metatarsus, making it clearly distinguishable from the Cryptic Myotis (M. crypticus) and Natterer’s Myotis ( M. nattereri ), which have wings attached to bases oftoes. Line of hair at edge of uropatagium is similar to that of Escalera’s Myotis , with two lines of bristles, one pointing outward and the another pointing inward, with thicker aspect than in the Cryptic Myotis and Natterer’s Myotis . Dorsal and ventral fur color is similar among all these species, but dorsum is generally much darker on the Zenati Myotis . Hairs also seem distinctly tricolored: black bases, brown middles, and white tips. Other morphological traits are common among all species in this species complex: relatively long ears, with several folds on outer edges that make them foldable; long, pointed, and straight tragus surpassing one-half the ear length;hairless face; narrow, long, and pointed muzzle; comparatively short feet, less than one-half the tibia length; and naked wings,ears, and tail membrane. Uropatagium is also enclosed by S-shaped long calcar. Skull is small and delicate with high braincase and strongly concave forehead regions; there are no sagittal or occipital crests; very similar to Escalera’s Myotis , although it can be distinguished byits broader and more globose braincase, generally more delicate skull structure, and slightly smaller dimensions (except for wider postorbital constriction). P? is more than one-half height and about three-quarters crown area of P* and is within tooth row; lower molars are myotodont.	Known from northern slopes of the Riff to dry mountains and several different lowland habitats from sea level up to relatively high mountains.	The Zenati Myotis is probably a gleaner, with the capacity to hover, and probably forages in dense cluttered habitats and close to the ground. It probably hunts arthropods on surfaces of leaves or directly from the ground.	The Zenati Myotis breeds in caves.	Along with Escalera’s Myotis, Zenati Myotis seem to be strictly cave dwelling.	No information.	Not assessed on The [UCN Red List. Zenati Myotis is one of the rarest bats in the Mediterranean region.	Coraman et al. (2019) | Juste et al. (2019) | Puechmaille, Allegrini et al. (2012) | Ruedi et al. (2019) | Salicini et al. (2011, 2012, 2013)	https://zenodo.org/record/6399013/files/figure.png	493. Zenati Myotis Myotis zenatius French: Murin zénéte / German: Zenati-Mausohr / Spanish: Ratonero zenate Other common names: Zenati Bat Taxonomy. Myotis zenatius Ibanez et al . in Juste et al, 2019 , “Mizou Cave, Tetouan, Moraceo (ca. 35°30' 42"N , 5°19'33"W ), at an altitude of 330 m a.s.1.” Subgenus Myotis ; myotis species group. See M. nattereri . The name M. escalerai cabrerae was recently proposed by E. Coraman and colleagues in 2019 based on populations that are now included under this species; the name is considered a nomen nudum as the publication did not include a proper description for the name. Monotypic. Distribution. NW Africa, including Morocco , Algeria ; also possibly in Tunisia . Descriptive notes. Head—-body 44-48 mm , tail 38-42 mm , ear 15-3-17- 3 mm , hindfoot 7-9-8-7, forearm 38- 2—40 mm . The Zenati Myotis is very similar to the other species in the M. nattereri species complex, which makesits identification complicated. As in Escalera’s Myotis ( M. escalerai ), plagiopatagium is inserted in mid-metatarsus, making it clearly distinguishable from the Cryptic Myotis (M. crypticus) and Natterer’s Myotis ( M. nattereri ), which have wings attached to bases oftoes. Line of hair at edge of uropatagium is similar to that of Escalera’s Myotis , with two lines of bristles, one pointing outward and the another pointing inward, with thicker aspect than in the Cryptic Myotis and Natterer’s Myotis . Dorsal and ventral fur color is similar among all these species, but dorsum is generally much darker on the Zenati Myotis . Hairs also seem distinctly tricolored: black bases, brown middles, and white tips. Other morphological traits are common among all species in this species complex: relatively long ears, with several folds on outer edges that make them foldable; long, pointed, and straight tragus surpassing one-half the ear length;hairless face; narrow, long, and pointed muzzle; comparatively short feet, less than one-half the tibia length; and naked wings,ears, and tail membrane. Uropatagium is also enclosed by S-shaped long calcar. Skull is small and delicate with high braincase and strongly concave forehead regions; there are no sagittal or occipital crests; very similar to Escalera’s Myotis , although it can be distinguished byits broader and more globose braincase, generally more delicate skull structure, and slightly smaller dimensions (except for wider postorbital constriction). P? is more than one-half height and about three-quarters crown area of P* and is within tooth row; lower molars are myotodont. Habitat. Known from northern slopes of the Riff to dry mountains and several different lowland habitats from sea level up to relatively high mountains. Food and Feeding. The Zenati Myotis is probably a gleaner, with the capacity to hover, and probably forages in dense cluttered habitats and close to the ground. It probably hunts arthropods on surfaces of leaves or directly from the ground. Breeding. The Zenati Myotis breeds in caves. Activity patterns. Along with Escalera’s Myotis, Zenati Myotis seem to be strictly cave dwelling. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The [UCN Red List. Zenati Myotis is one of the rarest bats in the Mediterranean region. Bibliography. Coraman et al. (2019), Juste et al. (2019), Puechmaille, Allegrini et al. (2012), Ruedi et al. (2019), Salicini et al. (2011, 2012, 2013).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis zenatius	Myotis	Unassigned-Myotis	zenatius	Ibáñez, Juste, Salicini, Puechmaille & Ruedi	2019	0	Acta Chiropterol.	20(2):295	Zenati Myotis	 cabrerae Çoraman, Mayer & Dietz, 2019 [nomen nudum]	Morocco, Tetouan, Mizou Cave	Morocco, Algeria, possibly Tunisia	Not listed (new species)	Not Evaluated	Çoraman et al. (2019) failed to meet the requirements for electronic publication of the name cabrerae (see Ruedi et al. 2019). Juste et al. (2019) electronically published the name zenatius for this species on 30 January 2019, satisfying all the ICZN (1999) requirements of electronic publication.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis zenatius	23	Zenati Myotis	Zenati Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	zenatius	Ibáñez, Juste, Salicini, Puechmaille, & Ruedi in Juste, Ruedi, Puechmaille, Salicini, & Ibáñez	2019	0	Myotis_zenatius	Juste, J., Ruedi, M., Puechmaille, S. J., Salicini, I., & Ibáñez, C. (2018). Two new cryptic bat species within the Myotis nattereri species complex (Vespertilionidae, Chiroptera) from the Western Palaearctic. Acta Chiropterologica, 20(2), 295.	https://bioone.org/journals/acta-chiropterologica/volume-20/issue-2/15081109ACC2018.20.2.001/Two-New-Cryptic-Bat-Species-within-the-Myotis-nattereri-Species/10.3161/15081109ACC2018.20.2.001.full	EBD 29831		"Mizou Cave, Tetouan, Morocco (ca. 35°30'42"N, 5°19'33"W), at an altitude of 330 m a.s.l."	35.512	-5.326	zenatius Ibáñez, Juste, Salicini, Puechmaille, & Ruedi in Juste, Ruedi, Puechmaille, Salicini, & Ibáñez, 2019|cabrerae Çoraman, F. Mayer, & Dietz in Çoraman, Dietz, Hempel, Ghazaryan, Levin, Presetnik, Zagmajster, & F. Mayer, 2019 [unavailabe name]	recently described; includes the recently described name cabrerae, which is considered unavaialble due to publication in which it was described not adhering to the ICZN Code	Juste, J., Ruedi, M., Puechmaille, S. J., Salicini, I., & Ibáñez, C. (2018). Two new cryptic bat species within the Myotis nattereri species complex (Vespertilionidae, Chiroptera) from the Western Palaearctic. Acta Chiropterologica, 20(2), 285-300.|Çoraman, E., Dietz, C., Hempel, E., Ghazaryan, A., Levin, E., Presetnik, P., ... & Mayer, F. (2019). Reticulate evolutionary history of a Western Palaearctic Bat Complex explained by multiple mt DNA introgressions in secondary contacts. Journal of Biogeography, 46(2), 343-354.|Ruedi, M., Puechmaille, S. J., Ibáñez, C., & Juste, J. (2019). Unavailable names in the Myotis nattereri species complex. Journal of Biogeography, 46(9), 2145-2146.	Morocco|Algeria|Tunisia?	Africa	Palearctic	NA	0	0	0	Myotis_zenatius	0	unmatched	NA	1																																			Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	zenatius	Ibáñez, Juste, Salicini, Puechmaille & Ruedi	2019	0	Acta Chiropterol.	20(2):295	Zenati Myotis	 cabrerae Çoraman, Mayer & Dietz, 2019 [nomen nudum]	Morocco, Tetouan, Mizou Cave	Morocco, Algeria, possibly Tunisia	Not listed (new species)	Not Evaluated	Çoraman et al. (2019) failed to meet the requirements for electronic publication of the name cabrerae (see Ruedi et al. 2019). Juste et al. (2019) electronically published the name zenatius for this species on 30 January 2019, satisfying all the ICZN (1999) requirements of electronic publication.	Myotis zenatius	1005488	23	Zenati Myotis	Zenati Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	zenatius	Ibáñez, Juste, Salicini, Puechmaille, & Ruedi in Juste, Ruedi, Puechmaille, Salicini, & Ibáñez	2019	0	Myotis_zenatius	Juste, J., Ruedi, M., Puechmaille, S. J., Salicini, I., & Ibáñez, C. (2018). Two new cryptic bat species within the Myotis nattereri species complex (Vespertilionidae, Chiroptera) from the Western Palaearctic. Acta Chiropterologica, 20(2), 295.	https://bioone.org/journals/acta-chiropterologica/volume-20/issue-2/15081109ACC2018.20.2.001/Two-New-Cryptic-Bat-Species-within-the-Myotis-nattereri-Species/10.3161/15081109ACC2018.20.2.001.full	EBD 29831		"Mizou Cave, Tetouan, Morocco (ca. 35°30'42"N, 5°19'33"W), at an altitude of 330 m a.s.l."	35.51167	-5.32583	zenatius Ibáñez, Juste, Salicini, Puechmaille, & Ruedi in Juste, Ruedi, Puechmaille, Salicini, & Ibáñez, 2019|cabrerae Çoraman, F. Mayer, & Dietz in Çoraman, Dietz, Hempel, Ghazaryan, Levin, Presetnik, Zagmajster, & F. Mayer, 2019 [unavailabe name]	recently described; includes the recently described name cabrerae, which is considered unavaialble due to publication in which it was described not adhering to the ICZN Code	Juste, J., Ruedi, M., Puechmaille, S. J., Salicini, I., & Ibáñez, C. (2018). Two new cryptic bat species within the Myotis nattereri species complex (Vespertilionidae, Chiroptera) from the Western Palaearctic. Acta Chiropterologica, 20(2), 285-300.|Çoraman, E., Dietz, C., Hempel, E., Ghazaryan, A., Levin, E., Presetnik, P., ... & Mayer, F. (2019). Reticulate evolutionary history of a Western Palaearctic Bat Complex explained by multiple mt DNA introgressions in secondary contacts. Journal of Biogeography, 46(2), 343-354.|Ruedi, M., Puechmaille, S. J., Ibáñez, C., & Juste, J. (2019). Unavailable names in the Myotis nattereri species complex. Journal of Biogeography, 46(9), 2145-2146.				Morocco|Algeria|Tunisia?	Africa	Palearctic	NA	0	0	0	Myotis_zenatius	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_zenatius	1005488	23	Zenati Myotis	Zenati Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	zenatius	Ibáñez, Juste, Salicini, Puechmaille, & Ruedi in Juste, Ruedi, Puechmaille, Salicini, & Ibáñez	0	Myotis zenatius	Juste, J., Ruedi, M., Puechmaille, S.J., Salicini, I. and Ibáñez, C. 2019-02-14. Two new cryptic bat species within the _Myotis nattereri_ species complex (Vespertilionidae, Chiroptera) from the western Palaearctic. Acta Chiropterologica 20(2):285-300.	https://doi.org/10.3161/15081109ACC2018.20.2.001	EBD 29831	holotype		"Mizou Cave, Tetouan, Morocco (ca. 35°30'42"N, 5°19'33"W), at an altitude of 330 m a.s.l."	35.51167	-5.32583	recently described; includes the recently described name cabrerae, which is considered unavailable due to publication in which it was described not adhering to the ICZN Code	Juste, J., Ruedi, M., Puechmaille, S. J., Salicini, I., & Ibáñez, C. (2018). Two new cryptic bat species within the Myotis nattereri species complex (Vespertilionidae, Chiroptera) from the Western Palaearctic. Acta Chiropterologica, 20(2), 285-300.|Çoraman, E., Dietz, C., Hempel, E., Ghazaryan, A., Levin, E., Presetnik, P., ... & Mayer, F. (2019). Reticulate evolutionary history of a Western Palaearctic Bat Complex explained by multiple mt DNA introgressions in secondary contacts. Journal of Biogeography, 46(2), 343-354.|Ruedi, M., Puechmaille, S. J., Ibáñez, C., & Juste, J. (2019). Unavailable names in the Myotis nattereri species complex. Journal of Biogeography, 46(9), 2145-2146.				Morocco|Algeria|Tunisia?	Africa	Palearctic	NE	0	0	0	Myotis_zenatius	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	zenatius	Ibáñez, Juste, Salicini, Puechmaille & Ruedi	2019	0	Acta Chiropterol.	20(2):295	Zenati Myotis	cabrerae Çoraman, Mayer & Dietz, 2019 [nomen nudum]	Morocco, Tetouan, Mizou Cave	Morocco, Algeria, possibly Tunisia	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	Not Evaluated	Çoraman et al. (2019) failed to meet the requirements for electronic publication of the name cabrerae (see Ruedi et al. 2019). Juste et al. (2019) electronically published the name zenatius for this species on 30 January 2019, satisfying all the ICZN (1999) requirements of electronic publication.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis zenatius; Myotis zenatius; Myotis zenatius; Myotis zenatius; cabrerae; zenatius; cabrerae; Murin zénéte; Zenati-Mausohr; Ratonero zenate; Zenati Bat; Zenati Myotis; Zenati Bat; Zenati Myotis; M. zenatius