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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L978	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis	Myotis yumanensis		[MSW2] Subgenus Leuconoe. See comments under leibii and lucifugus.; [MSW3] An older name for this species may be subulatus Say, 1823; see Glass and Baker (1968). Those authors recommended that subulatus should be supressed, but see Hall (1981), who used subulatus for the species we recognize as leibii. See also comments under leibii and lucifugus. Apparently closely related to velifer; see Ruedi and Mayer (2001).; [HMW] Vespertilio yumanensis H. Allen, 1864 , “Fort Yuma.” Restricted by G. S. Miller, Jr. in 1924 to “Old Fort Yuma, Imperial County, California , USA , on right bank of Colorado River, opposite present town of Yuma, Arizona .” Subgenus Pizonyx; albescens species group. Six subspecies recognized.; [batnames2022] An older name for this species may be subulatus Say, 1823; see Glass and Baker (1968). Those authors recommended that subulatus should be supressed, but see Hall (1981), who used subulatus for the species we recognize as leibii . Seealso comments under leibii and lucifugus . Apparently closely related to velifer ; see Ruedi and Mayer (2001).; [IUCN] Myotis yumanensis is apparently closely related to velifer .; [batnames2023] An older name for this species may be subulatus Say, 1823; see Glass and Baker (1968). Those authors recommended that subulatus should be supressed, but see Hall (1981), who used subulatus for the species we recognize as leibii . Seealso comments under leibii and lucifugus . Apparently closely related to velifer ; see Ruedi and Mayer (2001).; [batnames2025_1.7] An older name for this species may be subulatus Say, 1823; see Glass and Baker (1968). Those authors recommended that subulatus should be supressed, but see Hall (1981), who used subulatus for the species we recognize as leibii. Seealso comments under leibii and lucifugus. Apparently closely related to velifer; see Ruedi and Mayer (2001).						durangae. Iambi, lutosus, obscurus, oxalis, phasma, saturatus, sociabilis.	saturatus, oxalis, sociabilis, yumanensis, lambi, lutosus	yumanensis, lambi, lutosus, oxalis, saturatus, sociabilis	durangae, macropus, obscurus, phasma	yumanensis, lambi, lutosus, oxalis, saturatus, sociabilis		yumanensis, lambi, lutosus, oxalis, saturatus, sociabilis	yumanensis - durangae, macropus, obscurus, phasma	yumanensis, macropus, obscurus, saturatus, durangae, sociabilis, lutosus, phasma, lambi, oxalis	Myotis yumanensis is apparently closely related to velifer .	yumanensis, lambi, lutosus, oxalis, saturatus, sociabilis	yumanensis - durangae, macropus, obscurus, phasma	yumanensis, macropus, obscurus, saturatus, durangae, sociabilis, lutosus, phasma, lambi, oxalis 	subulatus, yumanensis, macropus, obscurus, saturatus, durangae, durangoae, sociabilis, lutosus, phasma, lambi, oxalis 	lambi, lutosus, oxalis, saturatus, sociabilis, yumanensis	yumanensis - durangae, macropus, obscurus, phasma	subulatus (Say in James, 1823)|yumanensis (H. Allen, 1864)|macropus (H. Allen, 1866) [preoccupied]|obscurus (H. Allen, 1866)|saturatus G. S. Miller, 1897|durangae J. A. Allen, 1903|durangoae Trouessart, 1904 [incorrect subsequent spelling]|sociabilis H. W. Grinnell, 1914|lutosus G. S. Miller & G. M. Allen, 1928|phasma G. S. Miller & G. M. Allen, 1928|lambi Benson, 1947|oxalis Dalquest, 1947		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Yuma myotis	British Columbia – C USA – C Mexico	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis yumanensis	U.S.A., California, Imperial Co., Old Fort Yuma.	H. Allen	1864	Smithson. Misc. Coll., 7:58.	Distribution: Ranging from southwestern Canada to central Mexico.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Yuma myotis	British Columbia – C USA – C Mexico	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	H. Allen	1864	Smithson. Misc. Coll., 7:58.	Subgenus Leuconoe. See comments under leibii and lucifugus.	Hidalgo, Morelos and Baja California (Mexico) north to British Columbia (Canada), east to Montana and W Texas (USA).	USA, California, Imperial Co., Old Fort Yuma.		H. ALLEN	1864	Size fairly small (forearm length, 32-38 mm; condylobasal length, 12-14 mm). Braincase of medium height. Middle upper premolar in toothrow. Fur not glossy.	Distribution: Ranging from southwestern Canada to central Mexico.	Six subspecies are currently recognized:	M. y. saturatus (southwestern Canada to southern California, mostly in coastal regions), M. y. oxalis (a small area in central California), M. y. sociabilis (southwestern Canada to northern California, mostly in interior regions), M. y. yumanensis (southwestern United States and most of northwestern Mexico), M. y. lambi (a small area in central Baja California), M. y. lutosus (central Mexico).	108	species	M. yumanensis	H. ALLEN	1864	Leuconoe	subgenus	Myotis yumanensis				Size fairly small (forearm length, 32-38 mm; condylobasal length, 12-14 mm). Braincase of medium height. Middle upper premolar in toothrow. Fur not glossy.	Six subspecies are currently recognized:		75. M. yumanensis (H. ALLEN 1864) [albescens group].	75	_M. y. lambi_ Benson, 1947; _M. y. lutosus_ Miller & Allen, 1928; _M. y. oxalis_ Dalquest, 1947; _M. y. saturatus_ Miller, 1897; _M. y. sociabilis_ Grinnell, 1914; _M. y. yumanensis_ (Allen, 1864) (synonyms: _durangae_ Allen, 1903, _macropus_ (Allen, 1866), _obscurus_ (Allen, 1866), _phasma_ Miller & Allen, 1928)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis yumanensis	Myotis		yumanensis	H. Allen	y	1864		Smithson. Misc. Coll.	7		58		Yuma Myotis	USA, California, Imperial Co., Old Fort Yuma.	Hidalgo, Morelos and Baja California (Mexico) north to British Columbia (Canada), east to Montana and W Texas (USA).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	durangae J. A. Allen, 1903; macropus H. Allen, 1866 [not Gould, 1854]; obscurus H. Allen, 1866; phasma Miller and G. M. Allen, 1928; lambi Benson, 1947; lutosus Miller and G. M. Allen, 1928; oxalis Dalquest, 1947; saturatus Miller, 1897 [not Kuzyakin, 1934]; sociabilis H. W. Grinnell, 1914.	An older name for this species may be subulatus Say, 1823; see Glass and Baker (1968). Those authors recommended that subulatus should be supressed, but see Hall (1981), who used subulatus for the species we recognize as leibii. See also comments under leibii and lucifugus. Apparently closely related to velifer; see Ruedi and Mayer (2001).	4C3D87E8FF4B6AF4FF5190C81657B1A9	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	938	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF4B6AF4FF5190C81657B1A9.xml	Myotis yumanensis	Vespertilionidae	Myotis	yumanensis		1864	Murin de Yuma @fr | Yuma-Mausohr @de | Ratonero de Yuma @es	Vespertilio yumanensis H. Allen, 1864 , “Fort Yuma.” Restricted by G. S. Miller, Jr. in 1924 to “Old Fort Yuma, Imperial County, California , USA , on right bank of Colorado River, opposite present town of Yuma, Arizona .” Subgenus Pizonyx; albescens species group. Six subspecies recognized.	M.y.yumanensisH.Allen,1864—WUSAandWMexicofromextremeW&SNe-vada,portionsofUtahandColorado,SthroughmostofBajaCaliforniaandtoCDurango,EtopanhandleofOklahomaandStoDelRio,Texas. M.y.lambiBenson,1947—NWMexico,knownonlyfromSanIgnacio,BajaCalifornia. M. y. lutosus G. S. Miller & G. M. Allen, 1928 — C Mexico from Sinaloa and S Zacatecas to Morelos . M.y.oxalisDalquest,1947—WUSA,inCentralValleyofCalifornia. M.y.saturatusG.S.Miller1897—SWCanadainCBritishColumbiaandthroughcoastalandcentralrangesofWestCoasttoMexicanborder. M. y. sociabilis H. W. Grinnell, 1914 — from SW Canada in SE British Columbia and W Montana W to C Washington and Oregon , and then S to NE California in W USA .	Head-body ¢. 48-49 mm , tail 27-40 mm , ear 11-15 mm , hindfoot 6-9 mm , forearm 32-37- 9 mm ; weight 4-7 g . Males are significantly smaller than females in some external and cranial measurements. Size of the Yuma Myotis is clinal from small in the south-western part of the distribution ( Sonora and southern Baja Peninsula) to large in north-eastern Utah and north-western Colorado . Fur is short (dorsal fur 5-6 mm ) and woolly. Dorsal hairs are bicolored, with dark brown bases and gray to cinnamon-brown or pale tan tips. Ventral hairs are paler, with dark brown bases and whitish to buffy tips. Color varies geographically, tending to be light in arid parts of the distribution and darker in forested areas. Color is dull brown in central part of the distribution, dark brown in the north-west, pale in the south-west, and dark brown in the Mexican highlands. Ears are comparatively short and extend beyond nares when laid forward. Tragus is slim and straight, with bluntly rounded tip, and basal lobe is large and somewhat semicircular in shape. Membranes, ears, and skin are usually pale brown to gray. Plagiopatagium is attached to foot at base of toes; uropatagium has sparse hairs, nearly reaching knees; scattered hairs on posterior edge of uropatagium do not constitute a fringe. Calcar is straight and long, extends ¢.60% the length between hindfoot and tail, lacks a keel, and terminates in minute lobule. Skull is moderate in size (greatest length of skull 12:9-14- 4 mm ); rostrum is broad, and palate is short and broad; forehead has abrupt slope; braincase is globose; sagittal crest is absent; and lambdoidal crests are absent or, when present, very low. Molars are dilambdodonts, with crowns longer in proportion to width. P? is aligned in tooth row. Chromosomal complement has 2n = 44 and FN = 50, with three large pairs and one small pair of metacentric and 17 pairs of acrocentric autosomes. X-chromosome is medium submetacentric, and Y-chromosome is small submetacentric or acrocentric.	Juniper and riparian woodlands and desert regions in coniferous forests, deciduous forests, grasslands, and treeless thorn scrublands, generally at low elevations but up to ¢. 3350 m . The Yuma Myotis is exclusively associated with open water, especially with relatively large bodies of permanent water, regardless of habitat type.	The Yuma Myotis forages over water and above riparian vegetation. It catches insects in the air during flight or by trawling in water surface using feet and uropatagium. It can also capture prey on surfaces of leaves, trunks, and rocks. Diet includes coleopterans and soft-bodied insects, such as dipterans, isopterans, lepidopterans, ephemeropterans, trichopterans, orthopterans, homopterans, hymenopterans, and neuropterans, and arachnids. Adults eat greater proportions of Trichoptera and Ephemeroptera and fewer Diptera , and subadults eat proportionally more Diptera . Diet appears to vary with insect availability rather than differences in prey selection, suggesting opportunistic feeding.	Males Yuma Myotis captured in June-August showed signs of spermatogenesis. Mating mostly occurs in autumn, and females storage sperm all winter until spring when ovulation and fertilization occur. Peak of pregnancies varies geographically: May-July in British Columbia; April-June until the 1990s and more recentlyJune-August in California; May-June in New Mexico , Oklahoma, and Mexico ; and July in Idaho and Colorado. Females give birth to one young, and during birth, they sit upright and catch newborn in their uropatagia. At birth, neonates are c.20% of their mothers’ weights. Females form nursery colonies in caves, abandoned buildings, or anywhere else that has high and stable temperatures. Males are usually absent from maternity colonies and remain alone while young are reared. Yuma Myotis attain reproductive maturity at a younger age than others North American Myotis .	Yuma Myotis emerge from diurnal roosts at dusk and forage in open areas over streams and ponds and over land in cluttered or uncluttered areas or close to trees. Activity is mostly nocturnal, starting a few minutes before sunset, usually peaking within two hours, and then decreasing, sometimes with second activity peak before dawn. Yuma Myotis often fed just a few inches above the surface of water, repeatedly flying regular routes. They flew up and down rivers and streams in relatively straight patterns and in circular patterns over ponds and small lakes. The Yuma Myotis roosts in caves, limestone and sandstone cliff crevices, hollow trees, loose pieces of bark, cracks in dead trees, abandoned mud nests of swallows, abandoned mines, bridges, attics, and other man-made buildings. Diurnal roosts are short distances from water and where light is dim. Yuma Myotis often use different roosting places each night. They use moderate-length, steeply swept FM calls while forage. Starting frequencies are 59-72 kHz that sweep abruptly down to 45-50 kHz. Calls rarely exceed 70 kHz and have distinctive lazy S-shape. Echolocation parameters vary geographically. In Arizona , maximum frequencies are 55-7-72-7 kHz, and durations are 1-8—4-6 milliseconds. In New Mexico , maximum frequencies are 60-2-78-8 kHz, minimum frequencies are 44-4-48-6 kHz, and durations are 2-2—4-2 milliseconds. Average frequency with maximum energy of search-phase echolocation calls in British Columbia was 55-7 kHz, lowest frequency was 38-5 kHz, and highest frequency was 87-6 kHz.	In California , the Yuma Myotis moved an average of 2 km (range 0-59- 3-5 km ) from roosts to capture locations, and in British Columbia , they moved c. 4 km from maternity roosts to foraging areas. In Arizona , it was reported to be a common summer resident but rare in winter. It was considered a temporary resident in some areas, presumably migrating to avoid very low winter temperatures. Nevertheless, observation of individuals in winter at known summer roosts in New Mexico suggests that it hibernates and does not migrate long distances. It can hibernate in low temperatures and generally is not active in winter. Yuma Myotis can form aggregations of more than 10,000 individuals. Roosts can be shared with other bat species, such as Little Brown Myotis ( M. lucifugus ).	Classified as Least Concern on The IUCN Red List. The Yuma Myotis is widespread and presumably has a large population.	Adams (2004) | Allen (1864) | Baker & Patton (1967) | Barbour & Davis (1969) | Benson (1947) | Braun et al. (2015) | Brigham et al. (1992) | Dalquest (1947) | Davis & Barbour (1970b) | Evelyn et al. (2004) | Fenton & Morris (1976) | Fenton et al. (1980) | Hall (1981) | Miller (1897 1924) | Nagorsen & Brigham (1993) | Solari (2019) | Vargas & Briones (2014) | Vaughan (1980) | Wilson & Ruff (1999)	https://zenodo.org/record/6398763/files/figure.png	396. Yuma Myotis Myotis yumanensis French: Murin de Yuma / German: Yuma-Mausohr / Spanish: Ratonero de Yuma Taxonomy. Vespertilio yumanensis H. Allen, 1864 , “Fort Yuma.” Restricted by G. S. Miller, Jr. in 1924 to “Old Fort Yuma, Imperial County, California , USA , on right bank of Colorado River, opposite present town of Yuma, Arizona .” Subgenus Pizonyx; albescens species group. Six subspecies recognized. Subspecies and Distribution. M.y.yumanensisH.Allen,1864—WUSAandWMexicofromextremeW&SNe-vada,portionsofUtahandColorado,SthroughmostofBajaCaliforniaandtoCDurango,EtopanhandleofOklahomaandStoDelRio,Texas. M.y.lambiBenson,1947—NWMexico,knownonlyfromSanIgnacio,BajaCalifornia. M. y. lutosus G. S. Miller & G. M. Allen, 1928 — C Mexico from Sinaloa and S Zacatecas to Morelos . M.y.oxalisDalquest,1947—WUSA,inCentralValleyofCalifornia. M.y.saturatusG.S.Miller1897—SWCanadainCBritishColumbiaandthroughcoastalandcentralrangesofWestCoasttoMexicanborder. M. y. sociabilis H. W. Grinnell, 1914 — from SW Canada in SE British Columbia and W Montana W to C Washington and Oregon , and then S to NE California in W USA . Descriptive notes. Head-body ¢. 48-49 mm , tail 27-40 mm , ear 11-15 mm , hindfoot 6-9 mm , forearm 32-37- 9 mm ; weight 4-7 g . Males are significantly smaller than females in some external and cranial measurements. Size of the Yuma Myotis is clinal from small in the south-western part of the distribution ( Sonora and southern Baja Peninsula) to large in north-eastern Utah and north-western Colorado . Fur is short (dorsal fur 5-6 mm ) and woolly. Dorsal hairs are bicolored, with dark brown bases and gray to cinnamon-brown or pale tan tips. Ventral hairs are paler, with dark brown bases and whitish to buffy tips. Color varies geographically, tending to be light in arid parts of the distribution and darker in forested areas. Color is dull brown in central part of the distribution, dark brown in the north-west, pale in the south-west, and dark brown in the Mexican highlands. Ears are comparatively short and extend beyond nares when laid forward. Tragus is slim and straight, with bluntly rounded tip, and basal lobe is large and somewhat semicircular in shape. Membranes, ears, and skin are usually pale brown to gray. Plagiopatagium is attached to foot at base of toes; uropatagium has sparse hairs, nearly reaching knees; scattered hairs on posterior edge of uropatagium do not constitute a fringe. Calcar is straight and long, extends ¢.60% the length between hindfoot and tail, lacks a keel, and terminates in minute lobule. Skull is moderate in size (greatest length of skull 12:9-14- 4 mm ); rostrum is broad, and palate is short and broad; forehead has abrupt slope; braincase is globose; sagittal crest is absent; and lambdoidal crests are absent or, when present, very low. Molars are dilambdodonts, with crowns longer in proportion to width. P? is aligned in tooth row. Chromosomal complement has 2n = 44 and FN = 50, with three large pairs and one small pair of metacentric and 17 pairs of acrocentric autosomes. X-chromosome is medium submetacentric, and Y-chromosome is small submetacentric or acrocentric. Habitat. Juniper and riparian woodlands and desert regions in coniferous forests, deciduous forests, grasslands, and treeless thorn scrublands, generally at low elevations but up to ¢. 3350 m . The Yuma Myotis is exclusively associated with open water, especially with relatively large bodies of permanent water, regardless of habitat type. Food and Feeding. The Yuma Myotis forages over water and above riparian vegetation. It catches insects in the air during flight or by trawling in water surface using feet and uropatagium. It can also capture prey on surfaces of leaves, trunks, and rocks. Diet includes coleopterans and soft-bodied insects, such as dipterans, isopterans, lepidopterans, ephemeropterans, trichopterans, orthopterans, homopterans, hymenopterans, and neuropterans, and arachnids. Adults eat greater proportions of Trichoptera and Ephemeroptera and fewer Diptera , and subadults eat proportionally more Diptera . Diet appears to vary with insect availability rather than differences in prey selection, suggesting opportunistic feeding. Breeding. Males Yuma Myotis captured in June-August showed signs of spermatogenesis. Mating mostly occurs in autumn, and females storage sperm all winter until spring when ovulation and fertilization occur. Peak of pregnancies varies geographically: May-July in British Columbia; April-June until the 1990s and more recentlyJune-August in California; May-June in New Mexico , Oklahoma, and Mexico ; and July in Idaho and Colorado. Females give birth to one young, and during birth, they sit upright and catch newborn in their uropatagia. At birth, neonates are c.20% of their mothers’ weights. Females form nursery colonies in caves, abandoned buildings, or anywhere else that has high and stable temperatures. Males are usually absent from maternity colonies and remain alone while young are reared. Yuma Myotis attain reproductive maturity at a younger age than others North American Myotis . Activity patterns. Yuma Myotis emerge from diurnal roosts at dusk and forage in open areas over streams and ponds and over land in cluttered or uncluttered areas or close to trees. Activity is mostly nocturnal, starting a few minutes before sunset, usually peaking within two hours, and then decreasing, sometimes with second activity peak before dawn. Yuma Myotis often fed just a few inches above the surface of water, repeatedly flying regular routes. They flew up and down rivers and streams in relatively straight patterns and in circular patterns over ponds and small lakes. The Yuma Myotis roosts in caves, limestone and sandstone cliff crevices, hollow trees, loose pieces of bark, cracks in dead trees, abandoned mud nests of swallows, abandoned mines, bridges, attics, and other man-made buildings. Diurnal roosts are short distances from water and where light is dim. Yuma Myotis often use different roosting places each night. They use moderate-length, steeply swept FM calls while forage. Starting frequencies are 59-72 kHz that sweep abruptly down to 45-50 kHz. Calls rarely exceed 70 kHz and have distinctive lazy S-shape. Echolocation parameters vary geographically. In Arizona , maximum frequencies are 55-7-72-7 kHz, and durations are 1-8—4-6 milliseconds. In New Mexico , maximum frequencies are 60-2-78-8 kHz, minimum frequencies are 44-4-48-6 kHz, and durations are 2-2—4-2 milliseconds. Average frequency with maximum energy of search-phase echolocation calls in British Columbia was 55-7 kHz, lowest frequency was 38-5 kHz, and highest frequency was 87-6 kHz. Movements, Home range and Social organization. In California , the Yuma Myotis moved an average of 2 km (range 0-59- 3-5 km ) from roosts to capture locations, and in British Columbia , they moved c. 4 km from maternity roosts to foraging areas. In Arizona , it was reported to be a common summer resident but rare in winter. It was considered a temporary resident in some areas, presumably migrating to avoid very low winter temperatures. Nevertheless, observation of individuals in winter at known summer roosts in New Mexico suggests that it hibernates and does not migrate long distances. It can hibernate in low temperatures and generally is not active in winter. Yuma Myotis can form aggregations of more than 10,000 individuals. Roosts can be shared with other bat species, such as Little Brown Myotis ( M. lucifugus ). Status and Conservation. Classified as Least Concern on The IUCN Red List. The Yuma Myotis is widespread and presumably has a large population. Bibliography. Adams (2004), Allen (1864), Baker & Patton (1967), Barbour & Davis (1969), Benson (1947), Braun et al. (2015), Brigham et al. (1992), Dalquest (1947), Davis & Barbour (1970b), Evelyn et al. (2004), Fenton & Morris (1976), Fenton et al. (1980), Hall (1981), Miller (1897 1924), Nagorsen & Brigham (1993), Solari (2019), Vargas & Briones (2014), Vaughan (1980), Wilson & Ruff (1999).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis yumanensis	Myotis	Unassigned-Myotis	yumanensis	H. Allen	1864	1	Smithson. Misc. Coll.	7:58	Yuma Myotis	 durangae J. A. Allen, 1903; macropus H. Allen, 1866 [not Gould, 1854]; obscurus H. Allen, 1866; phasma Miller and G. M. Allen, 1928; <b> lambi </b> Benson, 1947; <b> lutosus </b> Miller and G. M. Allen, 1928; <b>oxalis</b> Dalquest, 1947; <b> saturatus </b> Miller, 1897 [not Kuzyakin, 1934]; <b> sociabilis </b> H. W. Grinnell, 1914.	USA, California, Imperial Co., Old Fort Yuma.	Hidalgo, Morelos and Baja California (Mexico) north to British Columbia (Canada), east to Montana and W Texas (USA).	Not listed.	Least Concern	An older name for this species may be subulatus Say, 1823; see Glass and Baker (1968). Those authors recommended that subulatus should be supressed, but see Hall (1981), who used subulatus for the species we recognize as leibii . Seealso comments under leibii and lucifugus . Apparently closely related to velifer ; see Ruedi and Mayer (2001).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis yumanensis	23	Yuma Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	yumanensis	H. Allen	1864	1	Vespertilio_yumanensis	Allen, H. (1864). Monograph of the Bats of North America. Smithsonian Miscellaneous Collections, 7(1), 58.	https://www.biodiversitylibrary.org/page/8817039#page/98/mode/1up	USNM 5367, USNM 6019, USNM 6020, USNM 6021 [syntypes]		"Fort Yuma." Restricted by G. S. Miller, Jr. in 1924 to "Old Fort Yuma, Imperial County, California, USA, on right bank of Colorado River, opposite present town of Yuma, Arizona."			yumanensis (H. Allen, 1865)|macropus (H. Allen, 1866) [preoccupied]|obscurus (H. Allen, 1866)|saturatus G. S. Miller, 1897|durangae J. A. Allen, 1903|sociabilis H. W. Grinnell, 1914|lutosus G. S. Miller & G. M. Allen, 1928|phasma G. S. Miller & G. M. Allen, 1928|lambi Benson, 1947|oxalis Dalquest, 1947	NA	NA	Canada|United States|Mexico	North America	Nearctic	LC	0	0	0	Myotis_yumanensis	0	sciname match	Myotis_yumanensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14213	Myotis yumanensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	yumanensis	(H. Allen, 1864)	Myotis yumanensis is apparently closely related to velifer .	20000000	Myotis yumanensis	Least Concern		2019	2018-06-27 00:00:00 UTC	3.1	English	This species is listed as Least Concern in because of its wide distribution, presumed large population (locally common), large number of roosts in natural and human-made structures, occurs in pristine and altered habitats, it is present ;in a number of protected areas, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	This species is found in a variety of habitats, ranging from juniper and riparian woodlands to thorn scrubs and desert regions near open water (Ceballos and Oliva 2005, Braun et al. 2015). One is almost guaranteed to find this species wherever there are rivers, streams, ponds, lakes, etc. In fact, it is more closely associated with water than any other North American species of bat (Barbour and Davis 1969). ; It is generally considered to be an inhabitant of lower elevations and riparian situations, often in otherwise arid country, but usually associated with relatively large bodies of permanent water (Braun et al. 2015). When not near water over which to forage, these animals can be found in the thousands roosting in caves, attics, abandoned buildings, mines, underneath bridges, and other similar structures. Little is known about the migration of this species. However, it has been recorded in Texas as well as in its normal range during the winter season (Allen 1994). This bat is a very efficient insectivorous feeder that begins foraging at dusk and usually finishes two hours after sunset (Barbour and Davis 1969), this species is considered a 'trawler' (an aerial feeder) by its flight and morphometric characteristics (Fenton and Bogdanowicz 2002). Its diet includes beetles and soft-bodied insects such as flies, termites, moths, and mayflies (Braun et al. 2015).	There are no major threats throughout the species' range. However, ;distribution, roost requirements, and threats in winter are poorly known.	This bat is locally common. ;Total adult population size is unknown but undoubtedly exceeds 100,000. Hundreds of roosts exist, and these often include 100's to 1000's of individuals, colonies of M. yumanensis have been reported numbering as high as 10,000 individuals in crevices at Davis Dam on the Colorado River (Adams 2004, Braun et al.  2015).	Stable	Myotis yumanensis occurs in North America as far north as British Columbia, Canada southward through the western United States (Washington, Oregon, Idaho, Utah, Colorado, Nevada, California, New Mexico, and Arizona) and eastward to Montana, Oklahoma, and Texas, reaching as far south as Hidalgo, Morelos, and Michoacán, México (Simmons 2005, Braun et al.  2015).		Terrestrial	Overall, the species appears to be subject to ongoing slow reduction of available roost sites, localized detrimental effects of roost disturbance by humans, and degradation of foraging habitats. At the moment, it has not been listed in any conservation measure by the Mexican Government (Ceballos and Oliva 2005). The species occurs in several protected areas through its geographic distribution.	Nearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	yumanensis	H. Allen	1864	1	Smithson. Misc. Coll.	7:58	Yuma Myotis	 durangae J. A. Allen, 1903; macropus H. Allen, 1866 [not Gould, 1854]; obscurus H. Allen, 1866; phasma Miller and G. M. Allen, 1928; <b> lambi </b> Benson, 1947; <b> lutosus </b> Miller and G. M. Allen, 1928; <b>oxalis</b> Dalquest, 1947; <b> saturatus </b> Miller, 1897 [not Kuzyakin, 1934]; <b> sociabilis </b> H. W. Grinnell, 1914.	USA, California, Imperial Co., Old Fort Yuma.	Hidalgo, Morelos and Baja California (Mexico) north to British Columbia (Canada), east to Montana and W Texas (USA).	Not listed.	Least Concern	An older name for this species may be subulatus Say, 1823; see Glass and Baker (1968). Those authors recommended that subulatus should be supressed, but see Hall (1981), who used subulatus for the species we recognize as leibii . Seealso comments under leibii and lucifugus . Apparently closely related to velifer ; see Ruedi and Mayer (2001).	Myotis yumanensis	1005487	23	Yuma Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	yumanensis	H. Allen	1864	1	Vespertilio_yumanensis	Allen, H. (1864). Monograph of the Bats of North America. Smithsonian Miscellaneous Collections, 7(1), 58.	https://www.biodiversitylibrary.org/page/8817039#page/98/mode/1up	USNM 5367, USNM 6019, USNM 6020, USNM 6021 [syntypes]		"Fort Yuma." Restricted by G. S. Miller, Jr. in 1924 to "Old Fort Yuma, Imperial County, California, USA, on right bank of Colorado River, opposite present town of Yuma, Arizona."			yumanensis (H. Allen, 1865)|macropus (H. Allen, 1866) [preoccupied]|obscurus (H. Allen, 1866)|saturatus G. S. Miller, 1897|durangae J. A. Allen, 1903|sociabilis H. W. Grinnell, 1914|lutosus G. S. Miller & G. M. Allen, 1928|phasma G. S. Miller & G. M. Allen, 1928|lambi Benson, 1947|oxalis Dalquest, 1947	NA	NA			USA(WA,OR,ID,MT,WY,CA,NV,UT,AZ,CO,NM,OK,TX)	Canada|United States|Mexico	North America	Nearctic	LC	0	0	0	Myotis_yumanensis	0	sciname match	Myotis_yumanensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_yumanensis	1005487	23	Yuma Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	yumanensis	H. Allen	1	Vespertilio yumanensis	Allen, H. 1864-06. Monograph of the bats of North America. Smithsonian Miscellaneous Collections 7(1):1-85.	https://www.biodiversitylibrary.org/page/8817120	USNM:MAMM:5367, USNM:MAMM:6019, USNM:MAMM:6020, USNM:MAMM:6021	syntypes		"Fort Yuma." Restricted by G. S. Miller, Jr. in 1924 to "Old Fort Yuma, Imperial County, California, USA, on right bank of Colorado River, opposite present town of Yuma, Arizona."			NA	NA			USA(WA,OR,ID,MT,WY,CA,NV,UT,AZ,CO,NM,OK,TX)	Canada|United States|Mexico	North America	Nearctic	LC	0	0	0	Myotis_yumanensis	0	sciname match	Myotis_yumanensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	yumanensis	H. Allen	1864	1	Smithson. Misc. Coll.	7:58	Yuma Myotis	durangae J. A. Allen, 1903; macropus H. Allen, 1866 [not Gould, 1854]; obscurus H. Allen, 1866; phasma Miller and G. M. Allen, 1928; lambi Benson, 1947; lutosus Miller and G. M. Allen, 1928; oxalis Dalquest, 1947; saturatus Miller, 1897 [not Kuzyakin, 1934]; sociabilis H. W. Grinnell, 1914.	USA, California, Imperial Co., Old Fort Yuma.	Hidalgo, Morelos and Baja California (Mexico) north to British Columbia (Canada), east to Montana and W Texas (USA).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14213/22068335/' target='_blank'>Least Concern</a>	An older name for this species may be subulatus Say, 1823; see Glass and Baker (1968). Those authors recommended that subulatus should be supressed, but see Hall (1981), who used subulatus for the species we recognize as leibii. Seealso comments under leibii and lucifugus. Apparently closely related to velifer; see Ruedi and Mayer (2001).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis yumanensis; Myotis yumanensis; Myotis yumanensis; Myotis yumanensis; Myotis yumanensis; Myotis yumanensis; yumanensis; lambi; lutosus; oxalis; saturatus; sociabilis; durangae; macropus; obscurus; phasma; yumanensis; lambi; lutosus; oxalis; saturatus; sociabilis; lambi; lutosus; oxalis; saturatus; sociabilis; durangae; macropus; obscurus; phasma; yumanensis; macropus; obscurus; saturatus; durangae; sociabilis; lutosus; phasma; lambi; oxalis; Murin de Yuma; Yuma-Mausohr; Ratonero de Yuma; Yuma Myotis; Yuma Myotis; Yuma Myotis; M. yumanensis