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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L960	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis sicarius	Myotis sicarius	Myotis sicarius	Myotis sicarius	Myotis sicarius	Myotis sicarius	Myotis sicarius	Myotis sicarius	Myotis sicarius	Myotis sicarius	Myotis sicarius	Myotis sicarius	Myotis sicarius	Myotis sicarius	Myotis sicarius		[MSW2] Subgenus Myotis.; [MSW3] Reviewed by Bates and Harrison (1997).; [HMW] Myotis sicarius Thomas, 1915 , India , “Northern Sikim [= Sikkim ],” India . Subgenus Myotis ; daubentonii species group (7 species). See M. daubentonii . Monotypic.; [batnames2022] Reviewed by Bates and Harrison (1997).; [batnames2023] Reviewed by Bates and Harrison (1997).; [batnames2025_1.7] Reviewed by Bates and Harrison (1997).														sicarius				sicarius	sicarius			sicarius O. Thomas, 1915		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Nepal, Sikkim; ref. 4.76	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis sicarius	India, N. Sikkim.	Thomas	1915	J. Bombay Nat. Hist. Soc., 23:608.	Distribution: Confined to northeastern India.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Nepal, Sikkim	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1915	J. Bombay Nat. Hist. Soc., 23:608.	Subgenus Myotis.	Sikkim (NE India).	India, N Sikkim.		THOMAS	1915	Size fairly large (forearm length, 48-55 mm; condylobasal length, 17-19 mm). Rostrum shortened and narrowed. Middle upper premolar reduced and displaced medially from the toothrow. Anterior and middle lower incisors four-cusped.	Distribution: Confined to northeastern India.	No subspecies.		101	species	M. sicarius	THOMAS	1915	Myotis	subgenus	Myotis sicarius				Size fairly large (forearm length, 48-55 mm; condylobasal length, 17-19 mm). Rostrum shortened and narrowed. Middle upper premolar reduced and displaced medially from the toothrow. Anterior and middle lower incisors four-cusped.	No subspecies.		5. M. sicarius (THOMAS 1915) [myotis group].	5	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis sicarius	Myotis		sicarius	Thomas		1915		J. Bombay Nat. Hist. Soc.	23		608		Mandelli's Mouse-eared Myotis	India, N Sikkim.	Sikkim (NE India); Nepal.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		Reviewed by Bates and Harrison (1997).	4C3D87E8FF2E6A91FF8297591F9EB3D5	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	975	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF2E6A91FF8297591F9EB3D5.xml	Myotis sicarius	Vespertilionidae	Myotis	sicarius	Thomas	1915	Murin du Sikkim @fr | Mandelli-Wasserfledermaus @de | Ratonero de Sikkim @es | Mandelli's Mouse-eared Myotis @en | Sikkim Myotis @en	Myotis sicarius Thomas, 1915 , India , “Northern Sikim [= Sikkim ],” India . Subgenus Myotis ; daubentonii species group (7 species). See M. daubentonii . Monotypic.	N India ( Sikkim , West Bengal ) and C Nepal (Annapurna Conservation Area, Kathmandu Valley, Kaski district, Lalitpur district).	Head-body 50-56 mm , tail 44-54 mm , ear 17- 2-19 mm , hindfoot 85- 11 mm , forearm 48-1-54- 6 mm ; weight 9-611 g . Fur of Mandelli’s Myotis is dense, relatively short, and very soft. Dorsum is uniform chocolate-brown to dark ferruginous brown. Underparts are paler, with ginger hair tips and darker hair bases; belly is yellow-shaded. Forehead and muzzle are covered with hairs; ears, areas around eyes, and nostrils are essentially naked and dark. Ears are dark, with bluntly rounded tips. Anterior border of ear is evenly rounded and convex; posterior border has long shallow concavity beneath tip. Basal lobeis large and rounded. Tragus is tall and narrow, c.50% of pinna height. Feet are small but about equal to one-half the length of tibiae. Third metacarpal scarcely exceeds fourth and fifth. Wing membranes and uropatagium are naked, semi-translucent, and uniformly dark brown to black. Membranes are attached to base offirst phalanx of outer metatarsal. Narrow postcalcarial lobe is present. Baculum is small and saddle-shaped, with blunt rounded tip. Skull is relatively small, rostrum is long and robust, dorsal profile exhibits only slight post-nasal depression, and braincase is relatively shallow. Zygomata are short and widest posteriorly. Sagittal crest and lambdoid crests are weakly developed. Supraoccipital is nearly vertical and lambda just forms the most posterior part of the skull. Braincase is small and scarcely exceeds rostrum in surface area when viewed from above. P* and P, are relatively small. P, has crown area between one-quarter and one-third that of P, Condylo-canine lengths are 16-9-17- 7 mm ; maxillary tooth row lengths are 7-1-7- 5 mm .	Montane forests on hillsides and in valleys. Mandelli’s Myotis was collected over a river at Godvari in the Kathmandu Valley at ¢. 1350 m and over Bhurungdi River c. 3 m above waterin a partly forested area in Annapurna Himal ( Nepal ) at ¢. 1500 m .	No information.	Two female Mandelli’s Myotis captured in May in Annapurna Himal ( Nepal ) were heavily pregnant, each with full-term embryo in right uterine horn.	Mandelli’s Myotis is nocturnal. Individuals caught in Nepal (Annapurna Himal) emitted relatively high-intensity FM echolocation calls compared with other species of Myotis .	No information.	Classified as Vulnerable on The IUCN Red List. Mandelli’s Myotis occurs in less than 2000 km ?, with all known individuals at fewer than ten locations. Extent and quality ofits habitat continues to decline.	Acharya et al. (2010) | Agrawal et al. (1992) | Bates & Harrison (1997) | Corbet & Hill (1992) | Csor ba et al. (1999) | Findley (1972) | Koopman (1994) | Molur & Srinivasulu (2008d) | Molur et al. (2002) | Nguyen Truong Son et al. (2013) | Pearch (2011) | Ruedi, Csorba et al. (2015) | Ruedi, Stadelmann et al. (2013) | Simmons (2005) | Srinivasulu & Srinivasulu (2012) | Stadelmann, Jacobs et al. (2004) | Thapa, Shrestha et al. (2012) | Thomas (1915e) | Wroughton (1916)	https://zenodo.org/record/6398977/files/figure.png	479. Mandelli’s Myotis Myotis sicarius French: Murin du Sikkim / German: Mandelli-Wasserfledermaus / Spanish: Ratonero de Sikkim Other common names: Mandelli's Mouse-eared Myotis , Sikkim Myotis Taxonomy. Myotis sicarius Thomas, 1915 , India , “Northern Sikim [= Sikkim ],” India . Subgenus Myotis ; daubentonii species group (7 species). See M. daubentonii . Monotypic. Distribution. N India ( Sikkim , West Bengal ) and C Nepal (Annapurna Conservation Area, Kathmandu Valley, Kaski district, Lalitpur district). Descriptive notes. Head-body 50-56 mm , tail 44-54 mm , ear 17- 2-19 mm , hindfoot 85- 11 mm , forearm 48-1-54- 6 mm ; weight 9-611 g . Fur of Mandelli’s Myotis is dense, relatively short, and very soft. Dorsum is uniform chocolate-brown to dark ferruginous brown. Underparts are paler, with ginger hair tips and darker hair bases; belly is yellow-shaded. Forehead and muzzle are covered with hairs; ears, areas around eyes, and nostrils are essentially naked and dark. Ears are dark, with bluntly rounded tips. Anterior border of ear is evenly rounded and convex; posterior border has long shallow concavity beneath tip. Basal lobeis large and rounded. Tragus is tall and narrow, c.50% of pinna height. Feet are small but about equal to one-half the length of tibiae. Third metacarpal scarcely exceeds fourth and fifth. Wing membranes and uropatagium are naked, semi-translucent, and uniformly dark brown to black. Membranes are attached to base offirst phalanx of outer metatarsal. Narrow postcalcarial lobe is present. Baculum is small and saddle-shaped, with blunt rounded tip. Skull is relatively small, rostrum is long and robust, dorsal profile exhibits only slight post-nasal depression, and braincase is relatively shallow. Zygomata are short and widest posteriorly. Sagittal crest and lambdoid crests are weakly developed. Supraoccipital is nearly vertical and lambda just forms the most posterior part of the skull. Braincase is small and scarcely exceeds rostrum in surface area when viewed from above. P* and P, are relatively small. P, has crown area between one-quarter and one-third that of P, Condylo-canine lengths are 16-9-17- 7 mm ; maxillary tooth row lengths are 7-1-7- 5 mm . Habitat. Montane forests on hillsides and in valleys. Mandelli’s Myotis was collected over a river at Godvari in the Kathmandu Valley at ¢. 1350 m and over Bhurungdi River c. 3 m above waterin a partly forested area in Annapurna Himal ( Nepal ) at ¢. 1500 m . Food and Feeding. No information. Breeding. Two female Mandelli’s Myotis captured in May in Annapurna Himal ( Nepal ) were heavily pregnant, each with full-term embryo in right uterine horn. Activity patterns. Mandelli’s Myotis is nocturnal. Individuals caught in Nepal (Annapurna Himal) emitted relatively high-intensity FM echolocation calls compared with other species of Myotis . Movements, Home range and Social organization. No information. Status and Conservation. Classified as Vulnerable on The IUCN Red List. Mandelli’s Myotis occurs in less than 2000 km ?, with all known individuals at fewer than ten locations. Extent and quality ofits habitat continues to decline. Bibliography. Acharya et al. (2010), Agrawal et al. (1992), Bates & Harrison (1997), Corbet & Hill (1992), Csor ba et al. (1999), Findley (1972), Koopman (1994), Molur & Srinivasulu (2008d), Molur et al. (2002), Nguyen Truong Son et al. (2013), Pearch (2011), Ruedi, Csorba et al. (2015), Ruedi, Stadelmann et al. (2013), Simmons (2005), Srinivasulu & Srinivasulu (2012), Stadelmann, Jacobs et al. (2004), Thapa, Shrestha et al. (2012), Thomas (1915e), Wroughton (1916).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis sicarius	Myotis	Unassigned-Myotis	sicarius	Thomas	1915	0	J. Bombay Nat. Hist. Soc.	1.3806	Mandelli's Mouse-eared Myotis	None.	India, N Sikkim.	Sikkim (NE India); Nepal.	Not listed.	Vulnerable	Reviewed by Bates and Harrison (1997).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis sicarius	23	Mandelli's Myotis	Mandelli's Mouse-eared Myotis|Sikkim Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	sicarius	O. Thomas	1915	0	Myotis_sicarius	Thomas, O. (1915). Scientific Results form the Mammal Survey No. 10: The Indian Bats assigned to the genus Myotis. The journal of the Bombay Natural History Society, 23, 608.	https://www.biodiversitylibrary.org/item/95912#page/772/mode/1up	BM 1891.10.7.56		India, "Northern Sikim [= Sikkim]," India.			sicarius O. Thomas, 1915	NA	NA	Nepal|India	Asia	Indomalaya	VU	0	0	0	Myotis_sicarius	0	sciname match	Myotis_sicarius	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14202	Myotis sicarius	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	sicarius	Thomas, 1915		20000000	Myotis sicarius	Vulnerable	B2ab(iii)	2019	2018-08-31 00:00:00 UTC	3.1	English	This bat is endemic to South Asia, where it is only known from 6-7 locations in India and Nepal. Its area of occupancy is less than 2,000 km² and its habitat is declining due to deforestation, generally resulting from logging operations and the conversion of land for agricultural. It is confirmed as Vulnerable under criterion B2ab(iii).	This species is found in montane forests on hill sides and in valleys (Molur et al. 2002).	The habitat where this bat occurs is threatened by deforestation, generally resulting from logging operations and the conversion of land for agricultural and other uses, especially in Nepal (Molur et al. 2002).	There is little information available on the population of this species.	Unknown	This species is endemic to South Asia. Currently it is known from India (Sikkim and West Bengal) and Nepal (Central) (Molur et al. 2002, Srinivasulu and Srinivasulu 2012). Its extent of occurrence is estimated to be 68,954 km², however, the area of occupancy has been estimated as less than 2,000 km² based on the likely availability in habitat in the locations it has been recorded in. In Nepal, it has been recently reported from Bajrabarahi at an altitude of 1,485 m asl. in Khatmandu area (Thapa et al. 2012).		Terrestrial	Although there are no direct conservation measures in place, the species has been recorded from the Annapurna Conservation Area in Nepal. Surveys are needed and habitat management in Nepal is recommended (Molur et al. 2002).	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	sicarius	Thomas	1915	0	J. Bombay Nat. Hist. Soc.	1.380556	Mandelli's Mouse-eared Myotis	None.	India, N Sikkim.	Sikkim (NE India); Nepal.	Not listed.	Vulnerable	Reviewed by Bates and Harrison (1997).	Myotis sicarius	1005472	23	Mandelli's Myotis	Mandelli's Mouse-eared Myotis|Sikkim Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	sicarius	O. Thomas	1915	0	Myotis_sicarius	Thomas, O. (1915). Scientific Results form the Mammal Survey No. 10: The Indian Bats assigned to the genus Myotis. The journal of the Bombay Natural History Society, 23, 608.	https://www.biodiversitylibrary.org/item/95912#page/772/mode/1up	BM 1891.10.7.56		India, "Northern Sikim [= Sikkim]," India.			sicarius O. Thomas, 1915	NA	NA				Nepal|India	Asia	Indomalaya	VU	0	0	0	Myotis_sicarius	0	sciname match	Myotis_sicarius	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_sicarius	1005472	23	Mandelli's Myotis	Mandelli's Mouse-eared Myotis|Sikkim Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	sicarius	O. Thomas	0	Myotis sicarius	Thomas, O. 1915-05-25. Scientific results of the mammal survey. No. X. Journal of the Bombay Natural History Society 23(4):607-613.	https://www.biodiversitylibrary.org/page/30454075	BMNH:Mamm:1891.10.7.56	holotype	https://data.nhm.ac.uk/object/88ce5fa6-0cdd-4107-80ce-65a873de3451	India, "Northern Sikim [= Sikkim]," India.			NA	GyÅ‘rössy, D., Tu, V. T., Csorba, G., Thapa, S., Estók, P., Földvári, G., & Görföl, T. (2024). The grey zone of taxonomy—The case of the Sikkim Myotis (Chiroptera: Vespertilionidae: Myotis sicarius), first recorded from Southeast Asia. Vertebrate Zoology, 74, 737-749.				Nepal|India|Vietnam	Asia	Indomalaya	VU	0	0	0	Myotis_sicarius	0	sciname match	Myotis_sicarius	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	sicarius	Thomas	1915	0	J. Bombay Nat. Hist. Soc.	1.380556	Mandelli's Mouse-eared Myotis	None.	India, N Sikkim.	Sikkim (NE India); Nepal.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14202/22063965/' target='_blank'>Vulnerable</a>	Reviewed by Bates and Harrison (1997).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis sicarius; Myotis sicarius; Myotis sicarius; Myotis sicarius; Myotis sicarius; Myotis sicarius; sicarius; Murin du Sikkim; Mandelli-Wasserfledermaus; Ratonero de Sikkim; Mandelli's Mouse-eared Myotis; Sikkim Myotis; Mandelli's Myotis; Mandelli's Mouse-eared Myotis; Sikkim Myotis; Mandelli's Mouse-eared Myotis; Mandelli's Mouse-eared Myotis; M. sicarius