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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L959	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Myotis brandti [synonym of]	N/A	Myotis brandti [synonym of]	Myotis sibiricus	Myotis sibiricus	Myotis sibiricus	Myotis sibiricus	Myotis sibiricus	Myotis sibiricus	Myotis sibiricus	Myotis sibiricus	Myotis sibiricus		[HMW] Vespertilio mystacinus sibiricus Kastschenko, 1905 , Tomsk , Russia . Subgenus Pizonyx; brandtii species group. Myotis sibiricus is traditionally considered a subspecies of M. brandi under the name M. b. gracilis , which is now considered a synonym of M. sibiricus . Myotis gracilis has been variously recognized as a distinct species, but M. sibiricus 1s the oldest available name and thus has nomenclatural priority. Myotis sibiricus appears to be sister species to M. brandtii , but they are very tae Both species are very peculiarly embedded in the New World clade of Myotis , although the exact position of M. brandtii and M. sibiricus with respect to the New World Myotis is still not completely clear. Monotypic.; [batnames2022] Distinct from brandtii; see Kruskop et al. 2012, who suggested that sibiricus is the correct name for this species, rather than gracilis , a suggestion we have adopted here although additional work is required to clarify the relationships among sibiricus , gracilis , and specimens from Japan. See review in Kruskop et al. (2012).; [MDD2022] split from M. brandtii; includes gracilis; [IUCN] This species belongs to the subgenus Aeorestes Fitzinger, 1870 (Stadelmann et al. 2007). Most close to Myotis brandtii and treated formerly within it (Corbet 1978, Pavlinov et al. 1995, Tiunov 1997, Simmons 2005). Known geographic hiatus between western and eastern populations of M. brandtii s. lato runs somewhat W of the Ob’ River. Animals captured east of the Ob’ are genetically identical to Far East M. gracilis and differ from M. brandtii s. str. (Kruskop et al. 2007, 2008, 2012; Zhigalin et al. 2016, 2019).; [batnames2023] Distinct from brandtii; see Kruskop et al. 2012, who suggested that sibiricus is the correct name for this species, rather than gracilis , a suggestion we have adopted here although additional work is required to clarify the relationships among sibiricus , gracilis , and specimens from Japan. See review in Kruskop et al. (2012).; [MDD2023] split from M. brandtii; includes gracilis; [MDD2025_2.0] split from M. brandtii; includes gracilis; [batnames2025_1.7] Distinct from brandtii; see Kruskop et al. 2012, who suggested that sibiricus is the correct name for this species, rather than gracilis, a suggestion we have adopted here although additional work is required to clarify the relationships among sibiricus, gracilis, and specimens from Japan. See review in Kruskop et al. (2012).; [MDD2025_2.2] split from M. brandtii; includes gracilis													sibiricus	sibiricus, gracilis	This species belongs to the subgenus Aeorestes Fitzinger, 1870 (Stadelmann et al. 2007). Most close to Myotis brandtii and treated formerly within it (Corbet 1978, Pavlinov et al. 1995, Tiunov 1997, Simmons 2005). Known geographic hiatus between western and eastern populations of M. brandtii s. lato runs somewhat W of the Ob’ River. Animals captured east of the Ob’ are genetically identical to Far East M. gracilis and differ from M. brandtii s. str. (Kruskop et al. 2007, 2008, 2012; Zhigalin et al. 2016, 2019).	sibiricus 	sibiricus - gracilis	sibiricus, gracilis	sibiricus, gracilis	sibiricus	sibiricus - gracilis 	sibiricus (Kastschenko, 1905)|gracilis Ognev, 1927						N/A																																								NA																											4C3D87E8FF436AFCFA52936D161AB625	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	946	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF436AFCFA52936D161AB625.xml	Myotis sibiricus	Vespertilionidae	Myotis	sibiricus	Kastschenko	1905	Murin de Sibérie @fr | Sibirien-Mausohr @de | Ratonero de Siberia @es | Siberian Myotis @en	Vespertilio mystacinus sibiricus Kastschenko, 1905 , Tomsk , Russia . Subgenus Pizonyx; brandtii species group. Myotis sibiricus is traditionally considered a subspecies of M. brandi under the name M. b. gracilis , which is now considered a synonym of M. sibiricus . Myotis gracilis has been variously recognized as a distinct species, but M. sibiricus 1s the oldest available name and thus has nomenclatural priority. Myotis sibiricus appears to be sister species to M. brandtii , but they are very tae Both species are very peculiarly embedded in the New World clade of Myotis , although the exact position of M. brandtii and M. sibiricus with respect to the New World Myotis is still not completely clear. Monotypic.	E Asia from NE Mongolia to Russian Far East, NE China , Korea (including Jeju I), N Japan ( Hokkaido ), and Kuril Is.	Head-body 38-51 mm ,tail 32-45 mm , ear 12-15- 5 mm , hindfoot 8-10 mm , forearm 32-38 mm ; weight 4-9- 5 g . The Siberian Whiskered Myotisis relatively small. Fur is shaggy and long. Dorsal pelage is dark brown to yellowish brown (hairs with metallic sheen on tips); venter is paler and grayer than dorsum. Bare parts of ears, muzzle, and membranes are dark brown. Ears are not truncated but are rounded at tips, and anterior margins are turned outward above centers; tragusis straight or slightly turned outward, being under one-half the ear length. Wing attaches to distal part of metatarsus on first toe, and calcar is long, lacking postcalcarial lobe. Uropatagium has straight vein running to ankle from just above base oftail. Anteorbital ridge is distinct, and lacrimal foramen is not exposed; braincase is low; basicochlear fissure 1s long; P? and P, are in tooth rows; and upper molars have protoconules. Chromosomal complement Bos 2n = 44 and FN = 50 ( Korea ).	Generally forested habitats at higher elevations and agricultural areas at lower elevations.	Siberian Whiskered Myotis fly low and forage along streams between mountains and agriculturalfields.	Births of Siberian Whiskered Myotis probably occur in early July in Korea and Japan , and lactating females have been observed from late July to early August in Hokkaido . Females have one young per pregnancy.	Siberian Whiskered Myotis emerge in early evening and forage through the night. They use hollowsin trees for roots and will use caves as hibernacula. Calls are steep FM sweep, apparently with peak frequencies of 32-45 kHz.	Siberian Whiskered Myotis are known to roost alone, with one individual using a hole or crevice in a cave for hibernation. Maternity colonies can have 20-60 females and their young.	Not assessed on The [UCN Red List. The Siberian Whiskered Myotis was previously included in Brandt's Myotis ( M. brandtii ), which is classified as Least Concern. The Siberian Whiskered Myotis is widespread but rare throughout much of its distribution.	Jo Yeong-Seok et al. (2018) | Kawai et al. (2003) | Kruskop et al. (2012) | Ohdachi et al. (2009) | Stadelmann et al. (2007) | Yoo & Yoon Myung-Hee (1992)	https://zenodo.org/record/6398815/files/figure.png	419. Siberian Whiskered Myots Myotis sibiricus French: Murin de Sibérie / German: Sibirien-Mausohr / Spanish: Ratonero de Siberia Other common names: Siberian Myotis Taxonomy. Vespertilio mystacinus sibiricus Kastschenko, 1905 , Tomsk , Russia . Subgenus Pizonyx; brandtii species group. Myotis sibiricus is traditionally considered a subspecies of M. brandi under the name M. b. gracilis , which is now considered a synonym of M. sibiricus . Myotis gracilis has been variously recognized as a distinct species, but M. sibiricus 1s the oldest available name and thus has nomenclatural priority. Myotis sibiricus appears to be sister species to M. brandtii , but they are very tae Both species are very peculiarly embedded in the New World clade of Myotis , although the exact position of M. brandtii and M. sibiricus with respect to the New World Myotis is still not completely clear. Monotypic. Distribution. E Asia from NE Mongolia to Russian Far East, NE China , Korea (including Jeju I), N Japan ( Hokkaido ), and Kuril Is. Descriptive notes. Head-body 38-51 mm ,tail 32-45 mm , ear 12-15- 5 mm , hindfoot 8-10 mm , forearm 32-38 mm ; weight 4-9- 5 g . The Siberian Whiskered Myotisis relatively small. Fur is shaggy and long. Dorsal pelage is dark brown to yellowish brown (hairs with metallic sheen on tips); venter is paler and grayer than dorsum. Bare parts of ears, muzzle, and membranes are dark brown. Ears are not truncated but are rounded at tips, and anterior margins are turned outward above centers; tragusis straight or slightly turned outward, being under one-half the ear length. Wing attaches to distal part of metatarsus on first toe, and calcar is long, lacking postcalcarial lobe. Uropatagium has straight vein running to ankle from just above base oftail. Anteorbital ridge is distinct, and lacrimal foramen is not exposed; braincase is low; basicochlear fissure 1s long; P? and P, are in tooth rows; and upper molars have protoconules. Chromosomal complement Bos 2n = 44 and FN = 50 ( Korea ). Habitat. Generally forested habitats at higher elevations and agricultural areas at lower elevations. Food and Feeding. Siberian Whiskered Myotis fly low and forage along streams between mountains and agriculturalfields. Breeding. Births of Siberian Whiskered Myotis probably occur in early July in Korea and Japan , and lactating females have been observed from late July to early August in Hokkaido . Females have one young per pregnancy. Activity patterns. Siberian Whiskered Myotis emerge in early evening and forage through the night. They use hollowsin trees for roots and will use caves as hibernacula. Calls are steep FM sweep, apparently with peak frequencies of 32-45 kHz. Movements, Home range and Social organization. Siberian Whiskered Myotis are known to roost alone, with one individual using a hole or crevice in a cave for hibernation. Maternity colonies can have 20-60 females and their young. Status and Conservation. Not assessed on The [UCN Red List. The Siberian Whiskered Myotis was previously included in Brandt's Myotis ( M. brandtii ), which is classified as Least Concern. The Siberian Whiskered Myotis is widespread but rare throughout much of its distribution. Bibliography. Jo Yeong-Seok et al. (2018), Kawai et al. (2003), Kruskop et al. (2012), Ohdachi et al. (2009), Stadelmann et al. (2007), Yoo & Yoon Myung-Hee (1992).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis sibiricus	Myotis	Unassigned-Myotis	sibiricus	Kastschenko	1905	0	A Review of Mammals of Western Siberia. Part I. Chiroptera and Insectivora		Siberian Myotis	 sibiricus Katschenko, 1905.	Russia, Tomsk	E Russia, Lake Baikal Region to Ussuri Region, Sakhalin Isl., Kamchatka Peninsula, Kurile Isls., Korea and North Japan in Hokkaido	Not listed.	Least Concern	Distinct from brandtii; see Kruskop et al. 2012, who suggested that sibiricus is the correct name for this species, rather than gracilis , a suggestion we have adopted here although additional work is required to clarify the relationships among sibiricus , gracilis , and specimens from Japan. See review in Kruskop et al. (2012).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis sibiricus	23	Siberian Whiskered Myotis	Siberian Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	sibiricus	Kastschenko	1905	1						Tomsk, Russia.			sibiricus (Kastschenko, 1905)|gracilis Ognev, 1927	split from M. brandtii; includes gracilis	Ohdachi, S. D. I, Ishibashi, Y., Iwasa, M. A., & Saitoh, Takashi (2009). The Wild Mammals of Japan, Shoukadoh, Kyoto.|Kruskop, S. V., Borisenko, A. V., Ivanova, N. V., Lim, B. K., & Eger, J. L. (2012). Genetic diversity of northeastern Palaearctic bats as revealed by DNA barcodes. Acta Chiropterologica, 14(1), 1-14.	Russia|Mongolia|China|North Korea|South Korea|Japan	Asia	Indomalaya|Palearctic	LC	0	0	0	Myotis_sibiricus	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Myotis sibiricus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	sibiricus	Kastschenko, 1905	This species belongs to the subgenus Aeorestes Fitzinger, 1870 (Stadelmann et al. 2007). Most close to Myotis brandtii and treated formerly within it (Corbet 1978, Pavlinov et al. 1995, Tiunov 1997, Simmons 2005). Known geographic hiatus between western and eastern populations of M. brandtii s. lato runs somewhat W of the Ob’ River. Animals captured east of the Ob’ are genetically identical to Far East M. gracilis and differ from M. brandtii s. str. (Kruskop et al. 2007, 2008, 2012; Zhigalin et al. 2016, 2019).	90000000	Myotis sibiricus	Least Concern		2020	2019-07-12 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.ssessed as Near Threatened.	The species is widely distributed in flat and mountain forests and forest-steppe. In the summer lives in hollows of trees, buildings, and caves. It winters in caves near the summer habitat. The female gives birth to 1, rarely 2 pups. It has been documented to live to 41 years and has a forearm length of 4.1 cm.	Urban development is the main threat for the the species as it is both destroys and degrades forest and cave habitats upon which the species depends for foraging and roosting. Deforestation and fire reduce the extent and quality of forest habitat for the species whereas cavers cause winter mortality in roosts.	The species global population is currently suspected to be in decline, due to forest loss, degradation, and fragmentation, and loss of key roost sites in caves, but not at a rate to qualify as a threatened species.	Decreasing	This species is widely distributed in north Asia east of the Ob’ River from the Altai and Siberia to Primorye and Korea, Kamchatka, Kurile and Sakhalin Isls, Japan (Yoshiyuki 1989, Tiunov 1997, Botvinkin 2002, Kawai 2009, Zhigalin et al. 2019). The species is found in Kazakhstan and Mongolia (Bekenov et al. 1985, Datzman et al. 2012).		Terrestrial	Given the species wide distribution it is assumed to be present in some protected areas within its range. Known roosts should be protected and diverse educational efforts are needed to strengthen knowledge, appreciation and protection of the species and its habitat. Additional research is needed into its taxonomy, population status, life history and threats. The species habitat and known colonies need to be monitored.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	sibiricus	Kastschenko	1905	0	A Review of Mammals of Western Siberia. Part I. Chiroptera and Insectivora		Siberian Myotis	 sibiricus Katschenko, 1905.	Russia, Tomsk	E Russia, Lake Baikal Region to Ussuri Region, Sakhalin Isl., Kamchatka Peninsula, Kurile Isls., Korea and North Japan in Hokkaido	Not listed.	Least Concern	Distinct from brandtii; see Kruskop et al. 2012, who suggested that sibiricus is the correct name for this species, rather than gracilis , a suggestion we have adopted here although additional work is required to clarify the relationships among sibiricus , gracilis , and specimens from Japan. See review in Kruskop et al. (2012).	Myotis sibiricus	1005471	23	Siberian Whiskered Myotis	Siberian Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	sibiricus	Kastschenko	1905	1						Tomsk, Russia.			sibiricus (Kastschenko, 1905)|gracilis Ognev, 1927	split from M. brandtii; includes gracilis	Ohdachi, S. D. I, Ishibashi, Y., Iwasa, M. A., & Saitoh, Takashi (2009). The Wild Mammals of Japan, Shoukadoh, Kyoto.|Kruskop, S. V., Borisenko, A. V., Ivanova, N. V., Lim, B. K., & Eger, J. L. (2012). Genetic diversity of northeastern Palaearctic bats as revealed by DNA barcodes. Acta Chiropterologica, 14(1), 1-14.				Russia|Mongolia|China|North Korea|South Korea|Japan	Asia	Indomalaya|Palearctic	LC	0	0	0	Myotis_sibiricus	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_sibiricus	1005471	23	Siberian Whiskered Myotis	Siberian Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	sibiricus	Kastschenko	1	Vespertilio mystacinus subsp. sibiricus	Kastschenko, N.F. 1905. Обзоръ млекопитающихъ Западной Сибири и Туркестана. Выпускъ 1-й. Chiroptera, рукокрылыя.—Insectivora, насикомоядныя. Ðœ. Н. Кононов, Tomsk, 102 pp.					Tomsk, Russia.			split from M. brandtii; includes gracilis	Ohdachi, S. D. I, Ishibashi, Y., Iwasa, M. A., & Saitoh, Takashi (2009). The Wild Mammals of Japan, Shoukadoh, Kyoto.|Kruskop, S. V., Borisenko, A. V., Ivanova, N. V., Lim, B. K., & Eger, J. L. (2012). Genetic diversity of northeastern Palaearctic bats as revealed by DNA barcodes. Acta Chiropterologica, 14(1), 1-14.				Russia|Mongolia|China|North Korea|South Korea|Japan	Asia	Indomalaya|Palearctic	LC	0	0	0	Myotis_sibiricus	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	sibiricus	Kastschenko	1905	0	A Review of Mammals of Western Siberia. Part I. Chiroptera and Insectivora		Siberian Myotis	sibiricus Katschenko, 1905.	Russia, Tomsk	E Russia, Lake Baikal Region to Ussuri Region, Sakhalin Isl., Kamchatka Peninsula, Kurile Isls., Korea and North Japan in Hokkaido	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/85567062/85567065/' target='_blank'>Least Concern</a>	Distinct from brandtii; see Kruskop et al. 2012, who suggested that sibiricus is the correct name for this species, rather than gracilis, a suggestion we have adopted here although additional work is required to clarify the relationships among sibiricus, gracilis, and specimens from Japan. See review in Kruskop et al. (2012).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis sibiricus; Myotis sibiricus; Myotis sibiricus; Myotis sibiricus; Myotis sibiricus; sibiricus; sibiricus; gracilis; Murin de Sibérie; Sibirien-Mausohr; Ratonero de Siberia; Siberian Myotis; Siberian Whiskered Myotis; Siberian Myotis; Siberian Myotis; M. sibiricus