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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L954	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis rufopictus	Myotis formosus [synonym of]	Myotis formosus [synonym of]	Myotis formosus [synonym of]	Myotis formosus rufopictus	Myotis formosus rufopictus	Myotis rufopictus	Myotis rufopictus	Myotis rufopictus	Myotis rufopictus	Myotis rufopictus	Myotis rufopictus	Myotis rufopictus	Myotis rufopictus	Myotis rufopictus		[HMW] Vespertilio rufo-pictus Waterhouse, 1845 , “ Philippines .” Subgenus Chrysopteron. See M. formosus , from which it was split based on cranial, dental, and external morphology. Evidence suggests there may be another closely related species in the Philippines , as yet undescribed, which in coloration and size is close to M. weberi . Monotypic.; [batnames2022] Distinct from formosus; see Csorba et al. (2014).; [MDD2022] split from M. formosus; [IUCN] <p><span lang="HU">Previously Myotis rufopictus was considered to be a subspecies of M. formosus (Corbet and Hill 1992, Koopman 1993, Simmons 2005), but was elevated to full species on the basis of cranial, dental and external morphology (Heaney et al . 1998, Csorba et al . 2014). Large differences in skull measurements of Chrysopteron specimens within the Philippines and records of a reddish form in the islands (L. Heaney pers. comm. 2015) imply the presence of a second species beside M. rufopictus . However, published information were written before it became clear how to distinguish these species, so the present assessment lumps the two Philippine species together<span lang="HU">.</span></p>; [batnames2023] Distinct from formosus; see Csorba et al. (2014).; [MDD2023] split from M. formosus; [MDD2025_2.0] split from M. formosus; [batnames2025_1.7] Distinct from formosus; see Csorba et al. (2014).; [MDD2025_2.2] split from M. formosus														rufopictus	<p><span lang="HU">Previously Myotis rufopictus was considered to be a subspecies of M. formosus (Corbet and Hill 1992, Koopman 1993, Simmons 2005), but was elevated to full species on the basis of cranial, dental and external morphology (Heaney et al . 1998, Csorba et al . 2014). Large differences in skull measurements of Chrysopteron specimens within the Philippines and records of a reddish form in the islands (L. Heaney pers. comm. 2015) imply the presence of a second species beside M. rufopictus . However, published information were written before it became clear how to distinguish these species, so the present assessment lumps the two Philippine species together<span lang="HU">.</span></p>			rufopictus	rufopictus			rufopictus (G. R. Waterhouse, 1845)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Philippines		N/A																																								NA																											4C3D87E8FF406AF8FA829C9D1F42BD63	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	949	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF406AF8FA829C9D1F42BD63.xml	Myotis rufopictus	Vespertilionidae	Myotis	rufopictus		1845	Murin a doigts roux @fr | Orangefinger Mausohr @de | Ratonero de dedos narajnas @es | Red-painted Myotis @en	Vespertilio rufo-pictus Waterhouse, 1845 , “ Philippines .” Subgenus Chrysopteron. See M. formosus , from which it was split based on cranial, dental, and external morphology. Evidence suggests there may be another closely related species in the Philippines , as yet undescribed, which in coloration and size is close to M. weberi . Monotypic.	Philippines , occurring on Luzon ( Camarines Sur , Isabela, Nueva Viscaya, and Rizal provinces), Mindoro, Sibuyan, Negros, and Palawan (including Cuyo Is). On following pages: 424. Black-and-orange Myotis ( Myotis formosus ); 425. Geoffroy's Myotis ( Myotis emarginatus ); 426. Temminck's Myotis ( Myotis tricolon; 427. Welwitsch’'s Myotis ( Myotis welwitschii ); 428. Reddish-black Myotis ( Myotis rufonigen; 429. Scott's Myotis ( Myotis scott); 430. Morris's Myotis ( Myotis morrisi ); 431. Bocage's Myotis ( Myotis bocagil); 432. Malagasy Myotis ( Myotis goudoti ); 433. Anjouan Myotis ( Myotis anjouanensis ); 434. Kock's Myotis ( Myotis dieteri ); 435. Alcathoe Whiskered Myotis ( Myotis alcathoe ); 436. Hyrcanian Myotis ( Myotis hyrcanicus); 437. Pond Myotis ( Myotis dasycneme ); 438. lkonnikov's Myotis ( Myotis ikonnikovi ); 439. Sichuan Myotis ( Myotis altarium ); 440. Common Whiskered Myotis ( Myotis mystacinus ); 441. David's Myotis ( Myotis davidii ); 442. Valley Myotis ( Myotis ancricola ); 443. Anna Tess's Myotis ( Myotis annatessae );, 444. Gomantong Myotis ( Myotis gomantongensis ); 445. Peters's Myotis ( Myotis ater ); 446. Nepalese Whiskered Myotis ( Myotis muricola ); 447. Brown's Whiskered Myotis ( Myotis brown); 448. Insular Myotis ( Myotis insularum ).	Head-body 63- 4 mm (type), tail 43-59 mm , ear 19-22 mm , hindfoot 12-14 mm , forearm 47-58 mm ; weight 9- 5-17 g . The Orange-fingered Myotis is one of the most strikingly colored bats of Asia. It has fairly long pelage; upperparts usually bright yellow-orange, with pale gray roots; ventrally paler, yellowish white, scarcely tinted with gray at roots. Wing membranes are black, except wing bones, which are orange or orange brown, and a large bright red basal area roughly behind a line from thumb to heel; small strip of membrane above arm is red, clouded with black; limbs and interfemoral membrane are also red; naked part of muzzle and ears are pale flesh-colored; wing membranes attached to side of foot, at base oftoes. Ears are long; tragus long and slender, coming to blunt point. Nostrils simple (not tubular); muzzle not swollen. Skull profile is rather flat with no frontal depression;sagittal and lambdoid crests are only moderately developed, whereas skull is globose posteriorly. Canines are moderately strong; upper molars relatively robust with developed talons; P, very small and intruded lingually half-way out of line of tooth row. Condylo-canine length 16- 1 mm ; maxillary tooth row 7-2-7- 5 mm .	Primary lowland and montane forest, sometimes over limestone; also secondgrowth forest and fields near forest. Elevational range 50-1450 m .	No information.	No information.	No information.	[Location of roosts unknown, but the species has not been found in caves.	Classified as Data Deficient on The IUCN Red List. Probably threatened by deforestation in lowlands. It occurs in several protected areas.	Allen (1922) | Boitani et al. (2006) | Corbet & Hill (1992) | Csorba, Chou Cheng-Han et al. (2014) | Csorba, Rosell-Ambal et al. (2016) | Esselstyn, Widmann & Heaney (2004) | Heaney, Balete, Dolar et al. (1998) | Heaney, Balete & Rickart (2016) | Ingle & Heaney (1992) | Koopman (1993, 1994) | MBCFI (2018) | Mudar & Allen (1986) | Sedlock, Ingle & Balete (2011) | Sedlock, Weyandt et al. (2008) | Simmons (2005) | Tate (1941d) | Water house (1845)	https://zenodo.org/record/6398825/files/figure.png	423. Orange-fingered Myotis Myotis rufopictus French: Murin a doigts roux / German: OrangefingerMausohr / Spanish: Ratonero de dedos narajnas Other common names: Red-painted Myotis Taxonomy. Vespertilio rufo-pictus Waterhouse, 1845 , “ Philippines .” Subgenus Chrysopteron. See M. formosus , from which it was split based on cranial, dental, and external morphology. Evidence suggests there may be another closely related species in the Philippines , as yet undescribed, which in coloration and size is close to M. weberi . Monotypic. Distribution. Philippines , occurring on Luzon ( Camarines Sur , Isabela, Nueva Viscaya, and Rizal provinces), Mindoro, Sibuyan, Negros, and Palawan (including Cuyo Is). On following pages: 424. Black-and-orange Myotis ( Myotis formosus ); 425. Geoffroy's Myotis ( Myotis emarginatus ); 426. Temminck's Myotis ( Myotis tricolon; 427. Welwitsch’'s Myotis ( Myotis welwitschii ); 428. Reddish-black Myotis ( Myotis rufonigen; 429. Scott's Myotis ( Myotis scott); 430. Morris's Myotis ( Myotis morrisi ); 431. Bocage's Myotis ( Myotis bocagil); 432. Malagasy Myotis ( Myotis goudoti ); 433. Anjouan Myotis ( Myotis anjouanensis ); 434. Kock's Myotis ( Myotis dieteri ); 435. Alcathoe Whiskered Myotis ( Myotis alcathoe ); 436. Hyrcanian Myotis ( Myotis hyrcanicus); 437. Pond Myotis ( Myotis dasycneme ); 438. lkonnikov's Myotis ( Myotis ikonnikovi ); 439. Sichuan Myotis ( Myotis altarium ); 440. Common Whiskered Myotis ( Myotis mystacinus ); 441. David's Myotis ( Myotis davidii ); 442. Valley Myotis ( Myotis ancricola ); 443. Anna Tess's Myotis ( Myotis annatessae );, 444. Gomantong Myotis ( Myotis gomantongensis ); 445. Peters's Myotis ( Myotis ater ); 446. Nepalese Whiskered Myotis ( Myotis muricola ); 447. Brown's Whiskered Myotis ( Myotis brown); 448. Insular Myotis ( Myotis insularum ). Descriptive notes. Head-body 63- 4 mm (type), tail 43-59 mm , ear 19-22 mm , hindfoot 12-14 mm , forearm 47-58 mm ; weight 9- 5-17 g . The Orange-fingered Myotis is one of the most strikingly colored bats of Asia. It has fairly long pelage; upperparts usually bright yellow-orange, with pale gray roots; ventrally paler, yellowish white, scarcely tinted with gray at roots. Wing membranes are black, except wing bones, which are orange or orange brown, and a large bright red basal area roughly behind a line from thumb to heel; small strip of membrane above arm is red, clouded with black; limbs and interfemoral membrane are also red; naked part of muzzle and ears are pale flesh-colored; wing membranes attached to side of foot, at base oftoes. Ears are long; tragus long and slender, coming to blunt point. Nostrils simple (not tubular); muzzle not swollen. Skull profile is rather flat with no frontal depression;sagittal and lambdoid crests are only moderately developed, whereas skull is globose posteriorly. Canines are moderately strong; upper molars relatively robust with developed talons; P, very small and intruded lingually half-way out of line of tooth row. Condylo-canine length 16- 1 mm ; maxillary tooth row 7-2-7- 5 mm . Habitat. Primary lowland and montane forest, sometimes over limestone; also secondgrowth forest and fields near forest. Elevational range 50-1450 m . Food and Feeding. No information. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. [Location of roosts unknown, but the species has not been found in caves. Status and Conservation. Classified as Data Deficient on The IUCN Red List. Probably threatened by deforestation in lowlands. It occurs in several protected areas. Bibliography. Allen (1922), Boitani et al. (2006), Corbet & Hill (1992), Csorba, Chou Cheng-Han et al. (2014), Csorba, Rosell-Ambal et al. (2016), Esselstyn, Widmann & Heaney (2004), Heaney, Balete, Dolar et al. (1998), Heaney, Balete & Rickart (2016), Ingle & Heaney (1992), Koopman (1993, 1994), MBCFI (2018), Mudar & Allen (1986), Sedlock, Ingle & Balete (2011), Sedlock, Weyandt et al. (2008), Simmons (2005), Tate (1941d), Water house (1845).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis rufopictus	Myotis	Chrysopteron	rufopictus	Waterhouse	1845	1	Proc. Zool. Soc. Lond.	######	Red-painted Myotis	None.	Phillippines	Phillippines	Not listed.	Data Deficient	Distinct from formosus; see Csorba et al. (2014).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis rufopictus	23	Orange-fingered Myotis	Red-painted Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Chrysopteron	rufopictus	Waterhouse	1845	1						"Philippines."			rufopictus (Waterhouse, 1845)	split from M. formosus	Csorba, G., Chou, C. H., Ruedi, M., Görföl, T., Motokawa, M., Wiantoro, S., ... & Furey, N. (2014). The reds and the yellows: a review of Asian Chrysopteron Jentink, 1910 (Chiroptera: Vespertilionidae: Myotis). Journal of Mammalogy, 95(4), 663-678.	Philippines	Asia	Indomalaya	DD	0	0	0	Myotis_rufopictus	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136411	Myotis rufopictus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	rufopictus	(Waterhouse, 1845)	<p><span lang="HU">Previously Myotis rufopictus was considered to be a subspecies of M. formosus (Corbet and Hill 1992, Koopman 1993, Simmons 2005), but was elevated to full species on the basis of cranial, dental and external morphology (Heaney et al . 1998, Csorba et al . 2014). Large differences in skull measurements of Chrysopteron specimens within the Philippines and records of a reddish form in the islands (L. Heaney pers. comm. 2015) imply the presence of a second species beside M. rufopictus . However, published information were written before it became clear how to distinguish these species, so the present assessment lumps the two Philippine species together<span lang="HU">.</span></p>	20000000	Myotis rufopictus	Data Deficient		2016	2015-12-30 00:00:00 UTC	3.1	English	This species is assessed as Data Deficient in view of continuing doubts as to its, extent of occurrence, status and ecological requirements.	<p><span lang="HU">The habitat preferences of this species are poorly known. It has been found in primary lowland and montane forest and in an agricultural area near sea level (Mudar and Allen 1986, Heaney et al. 1998). It has been recorded on Luzon in secondary and primary lowland and montane forest, from near sea level to about 1,450 m elevation, including forest over limestone and agricultural fields near forest (Heaney et al. 2016).</p>	The species is probably threatened by deforestation in the lowland parts of its distribution.	This is a widespread but probably uncommon species (Heaney et al . 1998, 2016).	Unknown	This species is endemic to the Philippines.		Terrestrial	It occurs in a number of protected areas.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Chrysopteron	rufopictus	Waterhouse	1845	1	Proc. Zool. Soc. Lond.	########	Red-painted Myotis	None.	Phillippines	Phillippines	Not listed.	Data Deficient	Distinct from formosus; see Csorba et al. (2014).	Myotis rufopictus	1005466	23	Orange-fingered Myotis	Red-painted Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Chrysopteron	rufopictus	Waterhouse	1845	1						"Philippines."			rufopictus (Waterhouse, 1845)	split from M. formosus	Csorba, G., Chou, C. H., Ruedi, M., Görföl, T., Motokawa, M., Wiantoro, S., ... & Furey, N. (2014). The reds and the yellows: a review of Asian Chrysopteron Jentink, 1910 (Chiroptera: Vespertilionidae: Myotis). Journal of Mammalogy, 95(4), 663-678.				Philippines	Asia	Indomalaya	DD	0	0	0	Myotis_rufopictus	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_rufopictus	1005466	23	Orange-fingered Myotis	Red-painted Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Chrysopteron	rufopictus	G. R. Waterhouse	1	Vespertilio rufo-pictus	Waterhouse, G.R. 1845-04. Descriptions of species of Bats collectd in the Philippine Islands, and presented to the Society by H. Cuming, Esq. Proceedings of the Zoological Society of London 1845:3-10.	https://www.biodiversitylibrary.org/page/12862504	BMNH:Mamm:1907.1.1.533	holotype	https://data.nhm.ac.uk/object/8146120b-9cc1-4855-9603-27d4803b6be8	"Philippines."			split from M. formosus	Csorba, G., Chou, C. H., Ruedi, M., Görföl, T., Motokawa, M., Wiantoro, S., ... & Furey, N. (2014). The reds and the yellows: a review of Asian Chrysopteron Jentink, 1910 (Chiroptera: Vespertilionidae: Myotis). Journal of Mammalogy, 95(4), 663-678.				Philippines	Asia	Indomalaya	DD	0	0	0	Myotis_rufopictus	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Chrysopteron	rufopictus	Waterhouse	1845	1	Proc. Zool. Soc. Lond.	########	Red-painted Myotis	None.	Phillippines	Phillippines	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136411/22017446/' target='_blank'>Data Deficient</a>	Distinct from formosus; see Csorba et al. (2014).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis rufopictus; Myotis rufopictus; Myotis rufopictus; Myotis rufopictus; Myotis rufopictus; rufopictus; Murin a doigts roux; Orangefinger Mausohr; Ratonero de dedos narajnas; Red-painted Myotis; Orange-fingered Myotis; Red-painted Myotis; Red-painted Myotis; M. rufopictus