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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L951	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis rosseti	Myotis rosseti	Myotis rosseti	Myotis rosseti	Myotis rosseti	Myotis rosseti	Myotis rosseti	Myotis rosseti	Myotis rosseti	Myotis rosseti	Myotis rosseti	Myotis rosseti	Myotis rosseti	Myotis rosseti	Myotis rosseti		[MSW2] Subgenus Selysius. Originally described as a species of Glischropus; see Hill and Topal (1973).; [MSW3] Originally described as a species of Glischropus; see Hill and Topál (1973). Vietnamese record is not well documented; see Bates et al. (1999).; [HMW] Glischropus rosseti Oey, 1951 , Cambodia . Subgenus Myotis ; horsfieldii species group. See M. ridleyi . Based on neighbor-joining tree analysis in 2008, Myotis rosseti (partly as ndleyr) was related to M. cf. browni (as muricola ) and some “long-footed” Myotis (e.g. hasseltir). Monotypic.; [batnames2022] Originally described as a species of Glischropus ; see Hill and Topál (1973). Vietnamese record is not well documented; see Bates etal. (1999).; [IUCN] This species was originally described within the genus Glischropus, but was moved to Myotis by Hill and Topal (1973).; [batnames2023] Originally described as a species of Glischropus ; see Hill and Topál (1973). Vietnamese record is not well documented; see Bates etal. (1999).; [batnames2025_1.7] Originally described as a species of Glischropus; see Hill and Topál (1973). Vietnamese record is not well documented; see Bates etal. (1999).														rosseti	This species was originally described within the genus Glischropus, but was moved to Myotis by Hill and Topal (1973).			rosseti 	rosseti 			rosseti (Oei Hong Peng, 1951)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Thailand, Cambodia; ref. 4.83	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis rosseti	Kampuchea.	Oey	1951	Beaufortia, 1(8):4.	Distribution: Known only from Thailand and Cambodia.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Thick-thumbed myotis	Thailand, Kampuchea	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Oey	1951	Beaufortia, 1(8):4.	Subgenus Selysius. Originally described as a species of Glischropus; see Hill and Topal (1973).	Cambodia, Thailand.	Cambodia.		OEY	1951	Size small (forearm length, 27-31 mm; condylobasal length, 11-12 mm). Braincase fairly high; rostrum relatively short and broad. Middle upper and lower premolars absent. Thumb and foot pads present.	Distribution: Known only from Thailand and Cambodia.	No subspecies.		104	species	M. rosseti	OEY	1951	Selysius	subgenus	Myotis rosseti				Size small (forearm length, 27-31 mm; condylobasal length, 11-12 mm). Braincase fairly high; rostrum relatively short and broad. Middle upper and lower premolars absent. Thumb and foot pads present.	No subspecies.		33. M. rosseti (OEY 1951) [muricola group],	33	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis rosseti	Myotis		rosseti	Oey	y	1951		Beaufortia	1	8	4		Thick-thumbed Myotis	Cambodia.	Cambodia, Thailand, possibly Vietnam.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		Originally described as a species of Glischropus; see Hill and Topál (1973). Vietnamese record is not well documented; see Bates et al. (1999).	4C3D87E8FF366A89FF759C1F195BB11F	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	967	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF366A89FF759C1F195BB11F.xml	Myotis rosseti	Vespertilionidae	Myotis	rosseti		1951	Murin de Rosset @fr | Kambodscha-Langfu Rfledermaus @de | Ratonero de Rosset @es	Glischropus rosseti Oey, 1951 , Cambodia . Subgenus Myotis ; horsfieldii species group. See M. ridleyi . Based on neighbor-joining tree analysis in 2008, Myotis rosseti (partly as ndleyr) was related to M. cf. browni (as muricola ) and some “long-footed” Myotis (e.g. hasseltir). Monotypic.	S & SE Thailand , Cambodia , S Laos , C & S Vietnam ( Ha Tinh Province and Cat Tien Nadonal Park); in S Vienam , it might occur in nearby Binh Phuoc Province where “bamboo” habitat occurs.	Head-body 36-47 mm , tail 32-42 mm , ear 11-14 mm , hindfoot 5-1-7- 5 mm , forearm 27- 5-31 mm ; weight 3-8- 6 g . The Thick-thumbed Myotis has thick pinkish pads on thumbs and small triangular pads on feet. Fur is relatively short, light gray to dark grayish brown on upperparts, with hairs having dark brown bases and pale tips. Underparts are lighter. Ears are gray, long, and funnelshaped, with distinctive emargination on outer margins. Tragus (6-2-6- 8 mm ) is ¢.50% of ear length, thickened at base, and angled forward, tapering to tip. Muzzle and limbs are pinkish and not especially pigmented. Wing membrane is dark gray, and uropatagium is attached just below ankle. Calcar lobe is more or less developed. The Thick-thumbed Myotis differs from other species of Myotis by thickened pads on thumbs and feet and absence of P’ and P,. Ridley’s Myotis ( M. ridleyi ) and the Thick-thumbed Myotis can be readily recognized among other Myotis by their expanded, globose braincase, shortened, massive rostrum, and reduced premolar number. Glischropus spp. have shorter and broader ears, smaller and more rounded tragus, smaller and pinker thumb and foot pads, and distinctive upperincisors. Ridley’s Myotis is more blackish and lacks thumb pads. Characteristic thumb and foot pads of the Thick-thumbed Myotis also are found in one or more species of Pipistrellus , Glischropus , Eudiscopus , Tylonycteris , and Hesperoptenus . These distinctive morphological features also suggest Thick-thumbed Myotis are confined to bamboo associations. Baculum of a Thick-thumbed Myotis from southern Vietnam was moderately large (1- 1 mm in length), relatively wide, and weakly converging forward, with wide rounded tip; border between main body and “wings” was not apparent, and thickened edges were absent; upper side was concaved and had large rounded protrusion in basal one-third hanging over wide and shallow basal notch; and urethral groove was wide and deep. Skull is similar to that of Ridley’s Myotis , but braincase is a little more swollen frontally; interorbital region is shorter; rostrum is slightly shorter and wider, median depression is shallower and less clearly defined; narial emargination is more nearly U-shaped rather than V-shaped as in Ridley’s Myotis , not extending as far posteriorly; basial depressions more pronounced. Dentition of the Thick-thumbed Myotis is reduced with only 34 teeth. P* and P, are lacking. I? is short and wide and has anterior and posterior cusp, the former cusp is prominent and twice the height of the latter. I is ¢.50% the size of I,. Canines have strong cingulum but no prominent cingulum cusps and are only slightly taller than posterior premolars. Condylo-canine lengths are 10-5-11- 3 mm ; maxillary tooth row lengths are 4-1-4- 5 mm .	Densely forested mountain slopes, narrow valleys, and turbulent boulder strewn rivers (northern Vietnam ), semi-open landscapes with large number of tall bamboo (southern Vietnam ), disturbed habitats, and urban areas up to elevations of ¢. 500 m . Recent captures suggest that the Thick-thumbed Myotis is confined to bamboo associations and particularly adapted to using bamboo stems. In Thailand ,it roosted in tree hollows, house roofs, and bamboo.	The Thick-thumbed Myotis is known to forage for insects over disturbed areas including fields.	In Cat Tien, male Thick-thumbed Myotis with enlarged testes were observed in April-May and a pregnant female in November. Females captured in Vietnam in November had conspicuous evidence of post-lactation, suggesting late summer or early autumn birth and thus polyestrous reproduction.	In Cat Loc (Cat Tien National Park), the Thick-thumbed Myotis foraged c. 1-2 m aboveground among thickets and over cornfields in semi-disturbed and agricultural landscapes. An interesting community of bats adapted to living in bamboo stems, including poorly known species such as the Thick-thumbed Myotis , the Indochinese Thick-thumbed Bat ( Glischropus bucephalus ), and the mysterious Disk-footed Bat ( Eudiscopus denticulus ), occupy extensive bamboo-dominated formations in Cat Loc.	Heavy infestations of ectoparasites (nycteribiid flies) on Thick-thumbed Myotis suggest gregarious habits.	Classified as Least Concern on The IUCN Red List. Extent of occurrence of the Thick-thumbed Myotis is greater than 20,000 km 2, and recent evidence shows thatit has a wider distribution than was previously known. It has been reported in disturbed habitat and urban areas in some parts ofits distribution suggesting that it is tolerant of disturbance. Its populations are not believed to be declining fast enough to qualify for listing in a more threatened category.	Amador et al. (2018) | Bates et al. (1999) | Borisenko & Ivanova (2003) | Borisenko & Kruskop (2003) | Borisenko, Kruskop & Ivanova (2008) | Corbet & Hill (1992) | Csorba, Bumrungsri & Bates (2008) | Francis (2008a) | Francis et al. (2010) | Hill (1969a) | Hill & Topal (1973) | Kock (2000) | Koopman (1994) | Kruskop (2013a, 2013b) | Lekagul & McNeely (1988) | Oey (1951) | Pearch & Writer (2009) | Ruedi et al. (2013) | Simmons (2005) | Thomas et al. (2013) | Thong Vu Dinh (2015)	https://zenodo.org/record/6398928/files/figure.png	467. Thick-thumbed Myotis Myotis rosseti French: Murin de Rosset / German: Kambodscha-LangfuRfledermaus / Spanish: Ratonero de Rosset Taxonomy. Glischropus rosseti Oey, 1951 , Cambodia . Subgenus Myotis ; horsfieldii species group. See M. ridleyi . Based on neighbor-joining tree analysis in 2008, Myotis rosseti (partly as ndleyr) was related to M. cf. browni (as muricola ) and some “long-footed” Myotis (e.g. hasseltir). Monotypic. Distribution. S & SE Thailand , Cambodia , S Laos , C & S Vietnam ( Ha Tinh Province and Cat Tien Nadonal Park); in S Vienam , it might occur in nearby Binh Phuoc Province where “bamboo” habitat occurs. Descriptive notes. Head-body 36-47 mm , tail 32-42 mm , ear 11-14 mm , hindfoot 5-1-7- 5 mm , forearm 27- 5-31 mm ; weight 3-8- 6 g . The Thick-thumbed Myotis has thick pinkish pads on thumbs and small triangular pads on feet. Fur is relatively short, light gray to dark grayish brown on upperparts, with hairs having dark brown bases and pale tips. Underparts are lighter. Ears are gray, long, and funnelshaped, with distinctive emargination on outer margins. Tragus (6-2-6- 8 mm ) is ¢.50% of ear length, thickened at base, and angled forward, tapering to tip. Muzzle and limbs are pinkish and not especially pigmented. Wing membrane is dark gray, and uropatagium is attached just below ankle. Calcar lobe is more or less developed. The Thick-thumbed Myotis differs from other species of Myotis by thickened pads on thumbs and feet and absence of P’ and P,. Ridley’s Myotis ( M. ridleyi ) and the Thick-thumbed Myotis can be readily recognized among other Myotis by their expanded, globose braincase, shortened, massive rostrum, and reduced premolar number. Glischropus spp. have shorter and broader ears, smaller and more rounded tragus, smaller and pinker thumb and foot pads, and distinctive upperincisors. Ridley’s Myotis is more blackish and lacks thumb pads. Characteristic thumb and foot pads of the Thick-thumbed Myotis also are found in one or more species of Pipistrellus , Glischropus , Eudiscopus , Tylonycteris , and Hesperoptenus . These distinctive morphological features also suggest Thick-thumbed Myotis are confined to bamboo associations. Baculum of a Thick-thumbed Myotis from southern Vietnam was moderately large (1- 1 mm in length), relatively wide, and weakly converging forward, with wide rounded tip; border between main body and “wings” was not apparent, and thickened edges were absent; upper side was concaved and had large rounded protrusion in basal one-third hanging over wide and shallow basal notch; and urethral groove was wide and deep. Skull is similar to that of Ridley’s Myotis , but braincase is a little more swollen frontally; interorbital region is shorter; rostrum is slightly shorter and wider, median depression is shallower and less clearly defined; narial emargination is more nearly U-shaped rather than V-shaped as in Ridley’s Myotis , not extending as far posteriorly; basial depressions more pronounced. Dentition of the Thick-thumbed Myotis is reduced with only 34 teeth. P* and P, are lacking. I? is short and wide and has anterior and posterior cusp, the former cusp is prominent and twice the height of the latter. I is ¢.50% the size of I,. Canines have strong cingulum but no prominent cingulum cusps and are only slightly taller than posterior premolars. Condylo-canine lengths are 10-5-11- 3 mm ; maxillary tooth row lengths are 4-1-4- 5 mm . Habitat. Densely forested mountain slopes, narrow valleys, and turbulent boulder strewn rivers (northern Vietnam ), semi-open landscapes with large number of tall bamboo (southern Vietnam ), disturbed habitats, and urban areas up to elevations of ¢. 500 m . Recent captures suggest that the Thick-thumbed Myotis is confined to bamboo associations and particularly adapted to using bamboo stems. In Thailand ,it roosted in tree hollows, house roofs, and bamboo. Food and Feeding. The Thick-thumbed Myotis is known to forage for insects over disturbed areas including fields. Breeding. In Cat Tien, male Thick-thumbed Myotis with enlarged testes were observed in April-May and a pregnant female in November. Females captured in Vietnam in November had conspicuous evidence of post-lactation, suggesting late summer or early autumn birth and thus polyestrous reproduction. Activity patterns. In Cat Loc (Cat Tien National Park), the Thick-thumbed Myotis foraged c. 1-2 m aboveground among thickets and over cornfields in semi-disturbed and agricultural landscapes. An interesting community of bats adapted to living in bamboo stems, including poorly known species such as the Thick-thumbed Myotis , the Indochinese Thick-thumbed Bat ( Glischropus bucephalus ), and the mysterious Disk-footed Bat ( Eudiscopus denticulus ), occupy extensive bamboo-dominated formations in Cat Loc. Movements, Home range and Social organization. Heavy infestations of ectoparasites (nycteribiid flies) on Thick-thumbed Myotis suggest gregarious habits. Status and Conservation. Classified as Least Concern on The IUCN Red List. Extent of occurrence of the Thick-thumbed Myotis is greater than 20,000 km 2, and recent evidence shows thatit has a wider distribution than was previously known. It has been reported in disturbed habitat and urban areas in some parts ofits distribution suggesting that it is tolerant of disturbance. Its populations are not believed to be declining fast enough to qualify for listing in a more threatened category. Bibliography. Amador et al. (2018), Bates et al. (1999), Borisenko & Ivanova (2003), Borisenko & Kruskop (2003), Borisenko, Kruskop & Ivanova (2008), Corbet & Hill (1992), Csorba, Bumrungsri & Bates (2008), Francis (2008a), Francis et al. (2010), Hill (1969a), Hill & Topal (1973), Kock (2000), Koopman (1994), Kruskop (2013a, 2013b), Lekagul & McNeely (1988), Oey (1951), Pearch & Writer (2009), Ruedi et al. (2013), Simmons (2005), Thomas et al. (2013), Thong Vu Dinh (2015).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis rosseti	Myotis	Unassigned-Myotis	rosseti	Oey	1951	1	Beaufortia	1(8): 4	Thick-thumbed Myotis	None.	Cambodia.	Cambodia, Thailand, possibly Vietnam.	Not listed.	Least Concern	Originally described as a species of Glischropus ; see Hill and Topál (1973). Vietnamese record is not well documented; see Bates etal. (1999).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis rosseti	23	Thick-thumbed Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	rosseti	Oei Hong Peng	1951	1						Cambodia.			rosseti (Oei Hong Peng, 1951)	NA	NA	Thailand|Cambodia|Vietnam|Laos	Asia	Indomalaya	LC	0	0	0	Myotis_rosseti	0	sciname match	Myotis_rosseti	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14196	Myotis rosseti	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	rosseti	(Oey, 1951)	This species was originally described within the genus Glischropus, but was moved to Myotis by Hill and Topal (1973).	20000000	Myotis rosseti	Least Concern		2020	2018-09-01 00:00:00 UTC	3.1	English	Listed as Least Concern as its extent of occurrence is greater than 20,000 km², recent evidence shows that it has a wider distribution than was previously known, and it has been reported from disturbed habitat and urban areas in some parts of its range suggesting that it is quite tolerant of disturbed habitats. Its populations are not believed to be declining fast enough to qualify for listing in a more threatened category.	This species uses disturbed habitats, in Thailand it is found in tree hollows and house roofs, and bamboo. In Viet Nam it was found in semi-open habitats ; with large number of tall bamboo (Kruskop 2013). The anatomical feature (thickened pads on thumb) also suggests an adaptation to bamboo groves. In Viet Nam, females captured in November showed signs of postlactation while one pregnant female was reported in the beginning of November (Kruskop 2013).	There are no major threats to this adaptable species.	This species is known from only a small number of individuals in any country and are not believed to be declining fast enough to qualify for listing in a more threatened category.	Unknown	This species occurs in Cambodia, Thailand, and Viet Nam (Kruskop 2013), and is known from one locality in Lao PDR near Ban Paam, Dong AMphan NBCA, Attapu Province (Thomas et al. 2013).		Terrestrial	This species occurs in some protected areas throughout its range i.e. from Nam Cat Tien and Cat Tien NPs in Vietnam and Dong Amphan NBCA in Lao. Further studies are needed into the distribution, abundance, and ecology of this species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	rosseti	Oey	1951	1	Beaufortia	1(8): 4	Thick-thumbed Myotis	None.	Cambodia.	Cambodia, Thailand, possibly Vietnam.	Not listed.	Least Concern	Originally described as a species of Glischropus ; see Hill and Topál (1973). Vietnamese record is not well documented; see Bates etal. (1999).	Myotis rosseti	1005463	23	Thick-thumbed Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	rosseti	Oei Hong Peng	1951	1						Cambodia.			rosseti (Oei Hong Peng, 1951)	NA	NA				Thailand|Cambodia|Vietnam|Laos	Asia	Indomalaya	LC	0	0	0	Myotis_rosseti	0	sciname match	Myotis_rosseti	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_rosseti	1005463	23	Thick-thumbed Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	rosseti	Oei Hong Peng	1	Glischropus rosseti	Oei, H.P. 1951. A new species of Bat from Cambodge, _Glischropus rosseti_ sp. nov. Beaufortia 8:1-6.	https://repository.naturalis.nl/pub/505036	RMNH.MAM.19629, RMNH.MAM.19630	syntypes	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.19629.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.19629.b | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.19630	Cambodia.			NA	NA				Thailand|Cambodia|Vietnam|Laos	Asia	Indomalaya	LC	0	0	0	Myotis_rosseti	0	sciname match	Myotis_rosseti	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	rosseti	Oey	1951	1	Beaufortia	1(8): 4	Thick-thumbed Myotis	None.	Cambodia.	Cambodia, Thailand, possibly Vietnam.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href=”https://www.iucnredlist.org/species/14196/22062800/” target=”_blank>Least Concern</a>	Originally described as a species of Glischropus; see Hill and Topál (1973). Vietnamese record is not well documented; see Bates etal. (1999).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis rosseti; Myotis rosseti; Myotis rosseti; Myotis rosseti; Myotis rosseti; Myotis rosseti; rosseti; Murin de Rosset; Kambodscha-Langfu Rfledermaus; Ratonero de Rosset; Thick-thumbed Myotis; Thick-thumbed Myotis; Thick-thumbed Myotis; M. rosseti