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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L949	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi	Myotis ridleyi		[MSW2] Subgenus Selysius. Transferred from Pipistrellus; see Medway (1978:35).; [MSW3] Transferred from Pipistrellus; see Medway (1978); also see Hill and Topál (1973).; [HMW] Pipistrellus ridleyi Thomas, 1898 , “ Selangor , Malay Peninsula [= Peninsular Malaysia ].” Subgenus Myotis ; horsfieldii species group (9 species). In a 2013 phylogenetic study, M. ndleyi was related to M. rosseti and M. cf. browni and more distantly to an Oriental region clade of large-footed, “trawling” Myotis (hasselti, macropus , macrotarsus , and horsfieldii ). In a 2016 study, M. ridley and M. rosseti appeared as sister species (diverging c.4 million years ago) to a clade of “large-footed” Myotis ( horsfieldii , macrotarsus , macropus , adversus , and hasselti). Monotypic.; [batnames2022] Transferred from Pipistrellus ; see Medway (1978); also see Hill and Topál (1973).; [batnames2023] Transferred from Pipistrellus ; see Medway (1978); also see Hill and Topál (1973).; [batnames2025_1.7] Transferred from Pipistrellus; see Medway (1978); also see Hill and Topál (1973).														ridleyi				ridleyi	ridleyi			ridleyi (O. Thomas, 1898)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Ridley's bat	Malaya, ? Sumatra	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis ridleyi	Malaysia, Selangor.	Thomas	1898	Ann. Mag. Nat. Hist., ser. 7, 1:361.	Distribution: Known only from Malaya, Borneo and probably Sumatra.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Ridley's bat	Malaya, Borneo, ? Sumatra	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Thomas	1898	Ann. Mag. Nat. Hist., ser. 7, 1:361.	Subgenus Selysius. Transferred from Pipistrellus; see Medway (1978:35).	W Malaysia, Sumatra, Borneo.	Malaysia, Selangor.		THOMAS	1898	Size small (forearm length, 28-32 mm; condylobasal length, 11-12 mm). Braincase relatively high; rostrum relatively short and of medium width. Middle upper and lower premolars absent. Canines relatively short. No thumb or foot pads.	Distribution: Known only from Malaya, Borneo and probably Sumatra.	No subspecies.		104	species	M. ridleyi	THOMAS	1898	Selysius	subgenus	Myotis ridleyi				Size small (forearm length, 28-32 mm; condylobasal length, 11-12 mm). Braincase relatively high; rostrum relatively short and of medium width. Middle upper and lower premolars absent. Canines relatively short. No thumb or foot pads.	No subspecies.		32. M. ridleyi (THOMAS 1898) [muricola group].	32	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis ridleyi	Myotis		ridleyi	Thomas		1898		Ann. Mag. Nat. Hist., ser. 7	1		361		Ridley's Myotis	Malaysia, Selangor (= Kepong).	W Malaysia, Sumatra, Borneo.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		Transferred from Pipistrellus; see Medway (1978); also see Hill and Topál (1973).	4C3D87E8FF376A89FA509ACB1A87B133	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	966	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF376A89FA509ACB1A87B133.xml	Myotis ridleyi	Vespertilionidae	Myotis	ridleyi		1898	Murin de Ridley @fr | Ridley-Langful3fledermaus @de | Ratonero de Ridley @es | Ridley's Bat @en	Pipistrellus ridleyi Thomas, 1898 , “ Selangor , Malay Peninsula [= Peninsular Malaysia ].” Subgenus Myotis ; horsfieldii species group (9 species). In a 2013 phylogenetic study, M. ndleyi was related to M. rosseti and M. cf. browni and more distantly to an Oriental region clade of large-footed, “trawling” Myotis (hasselti, macropus , macrotarsus , and horsfieldii ). In a 2016 study, M. ridley and M. rosseti appeared as sister species (diverging c.4 million years ago) to a clade of “large-footed” Myotis ( horsfieldii , macrotarsus , macropus , adversus , and hasselti). Monotypic.	Malay Peninsula and N Borneo ( Sabah , N Kalimantan, and Gunung Gading National Park in SW Sarawak ). Data from Sumatra might be erroneous; specimens previously reported from there were identified as the Hairy-faced Myotis ( M. annectans ), but both species might not occur there. Data from Sulawesi identified as M. cf. ridleyi requires more detailed studies, and on zoogeographical grounds, they probably represent a different taxon.	Head-body 36-41 mm , tail 23-33 mm , ear 8-11 mm , hindfoot 5-1- 7: 5 mm , forearm 27-37 mm ; weight 4-6- 5 g . Ridley’s Myotis has short wings butrelatively heavy body and typical Myotis ears and tragus shape albeit the latter is relatively short and wide at its base. Upperparts are uniformly dark brown, with short velvety fur. Underparts are slightly paler, gray brown to dark brown. Face and muzzle are brown. Feet are small and naked, but toes are covered with stiff hairs. Wings are brownish black and attach at bases of toes. Ears are medium-sized and brownish black, with fairly long tragus, pointed tips, and distinct rounded basal lobe, curved forward. Baculum is less than 1 mm long and saddle-shaped. Skull has elevated frontal region. Braincase is relatively high and inflated. Rostrum is short and deep, with medium width. Dentition is reduced with only 34 teeth. P® and P, are lacking. Incisors are short and broad; canines are relatively short, C' barely exceeds P* in height, and C,is equal in height to P,. There are only two upper and lower premolars. P? is in tooth row, in contact with canine, but separated from P* by short diastema. Condylo-canine lengths are 11-12 mm ; maxillary tooth row lengths are 4-4— 4-8 mm . Chromosomal complement has 2n = 44 and FN = 50 (Ulu Gombak, Peninsular Malaysia ).	Dipterocarp and lowland primary forests, near lowland forest streams, and pristine lowland evergreen forest, near peat swamps, at upper elevations of ¢. 1000 m .	Ridley’s Myotis forages over small streams in forest understories and might be dependent on water to forage.	A pregnant Ridley’s Myotis was caught in April. At Krau Wildlife Reserve (Peninsular Malaysia ), lactating females were found in May and August and pregnant females in March and October.	Ridley’s Myotis is nocturnal and naturally roosts in caves, rock crevices and hollows, fallen logs, and small crevices in road culverts and under human houses/cabins. Some individuals were caught using hand nets near Krau Wildlife Reserve headquarters inside a culvert.	At Tabin Wildlife Reserve ( Sabah ), a colony of ¢.30 Ridley’s Myotis has roosted under a visitor cabin on a hill for many years.	Classified as Near Threatened on The IUCN Red List. Ridley’s Myotis has probably declined by close to 30% over the last 15 years due to significant loss ofits forest habitat. This rate of decline is predicted to continue for the next 15 years. Habitat loss from logging, agriculture, plantations, and forest fires is a major threat. It occurs in Kayan Mentarang National Park (Borneo) and other protected areas. It is rarely collected and requires additional research.	Amador et al. (2018) | Boitani et al. (2006) | Bumrungsri et al. (2006) | Chasen (1940) | Corbet & Hill (1992) | Francis (2008a) | Heller & Volleth (1989) | Hill (1969a) | Hill & Harrison (1987) | Hill & Topéal (1973) | Hutson, Francis, Kingston & Bumrungsri (2008) | Khan et al. (2008) | Kingston et al. (2006) | Koopman (1994) | Lim et al. (2017) | Medway (1978, 1983) | Payne et al. (1985) | Phillipps & Phillipps (2016) | Puri (1997) | Ruedi et al. (2013) | Simmons (2005) | Tate (1941d) | Thomas (1898b) | Volleth & Heller (2012) | Wiantoro et al. (2017)	https://zenodo.org/record/6398926/files/figure.png	466. Ridley’s Myotis Myotis ridleyi French: Murin de Ridley / German: Ridley-Langful3fledermaus / Spanish: Ratonero de Ridley Other common names: Ridley's Bat Taxonomy. Pipistrellus ridleyi Thomas, 1898 , “ Selangor , Malay Peninsula [= Peninsular Malaysia ].” Subgenus Myotis ; horsfieldii species group (9 species). In a 2013 phylogenetic study, M. ndleyi was related to M. rosseti and M. cf. browni and more distantly to an Oriental region clade of large-footed, “trawling” Myotis (hasselti, macropus , macrotarsus , and horsfieldii ). In a 2016 study, M. ridley and M. rosseti appeared as sister species (diverging c.4 million years ago) to a clade of “large-footed” Myotis ( horsfieldii , macrotarsus , macropus , adversus , and hasselti). Monotypic. Distribution. Malay Peninsula and N Borneo ( Sabah , N Kalimantan, and Gunung Gading National Park in SW Sarawak ). Data from Sumatra might be erroneous; specimens previously reported from there were identified as the Hairy-faced Myotis ( M. annectans ), but both species might not occur there. Data from Sulawesi identified as M. cf. ridleyi requires more detailed studies, and on zoogeographical grounds, they probably represent a different taxon. Descriptive notes. Head-body 36-41 mm , tail 23-33 mm , ear 8-11 mm , hindfoot 5-1- 7: 5 mm , forearm 27-37 mm ; weight 4-6- 5 g . Ridley’s Myotis has short wings butrelatively heavy body and typical Myotis ears and tragus shape albeit the latter is relatively short and wide at its base. Upperparts are uniformly dark brown, with short velvety fur. Underparts are slightly paler, gray brown to dark brown. Face and muzzle are brown. Feet are small and naked, but toes are covered with stiff hairs. Wings are brownish black and attach at bases of toes. Ears are medium-sized and brownish black, with fairly long tragus, pointed tips, and distinct rounded basal lobe, curved forward. Baculum is less than 1 mm long and saddle-shaped. Skull has elevated frontal region. Braincase is relatively high and inflated. Rostrum is short and deep, with medium width. Dentition is reduced with only 34 teeth. P® and P, are lacking. Incisors are short and broad; canines are relatively short, C' barely exceeds P* in height, and C,is equal in height to P,. There are only two upper and lower premolars. P? is in tooth row, in contact with canine, but separated from P* by short diastema. Condylo-canine lengths are 11-12 mm ; maxillary tooth row lengths are 4-4— 4-8 mm . Chromosomal complement has 2n = 44 and FN = 50 (Ulu Gombak, Peninsular Malaysia ). Habitat. Dipterocarp and lowland primary forests, near lowland forest streams, and pristine lowland evergreen forest, near peat swamps, at upper elevations of ¢. 1000 m . Food and Feeding. Ridley’s Myotis forages over small streams in forest understories and might be dependent on water to forage. Breeding. A pregnant Ridley’s Myotis was caught in April. At Krau Wildlife Reserve (Peninsular Malaysia ), lactating females were found in May and August and pregnant females in March and October. Activity patterns. Ridley’s Myotis is nocturnal and naturally roosts in caves, rock crevices and hollows, fallen logs, and small crevices in road culverts and under human houses/cabins. Some individuals were caught using hand nets near Krau Wildlife Reserve headquarters inside a culvert. Movements, Home range and Social organization. At Tabin Wildlife Reserve ( Sabah ), a colony of ¢.30 Ridley’s Myotis has roosted under a visitor cabin on a hill for many years. Status and Conservation. Classified as Near Threatened on The IUCN Red List. Ridley’s Myotis has probably declined by close to 30% over the last 15 years due to significant loss ofits forest habitat. This rate of decline is predicted to continue for the next 15 years. Habitat loss from logging, agriculture, plantations, and forest fires is a major threat. It occurs in Kayan Mentarang National Park (Borneo) and other protected areas. It is rarely collected and requires additional research. Bibliography. Amador et al. (2018), Boitani et al. (2006), Bumrungsri et al. (2006), Chasen (1940), Corbet & Hill (1992), Francis (2008a), Heller & Volleth (1989), Hill (1969a), Hill & Harrison (1987), Hill & Topéal (1973), Hutson, Francis, Kingston & Bumrungsri (2008), Khan et al. (2008), Kingston et al. (2006), Koopman (1994), Lim et al. (2017), Medway (1978, 1983), Payne et al. (1985), Phillipps & Phillipps (2016), Puri (1997), Ruedi et al. (2013), Simmons (2005), Tate (1941d), Thomas (1898b), Volleth & Heller (2012), Wiantoro et al. (2017).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis ridleyi	Myotis	Unassigned-Myotis	ridleyi	Thomas	1898	0	Ann. Mag. Nat. Hist.	ser. 7, 1: 361	Ridley's Myotis	None.	Malaysia, Selangor (= Kepong).	W Malaysia, Sumatra, Borneo.	Not listed.	Near Threatened	Transferred from Pipistrellus ; see Medway (1978); also see Hill and Topál (1973).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis ridleyi	23	Ridley's Myotis	Ridley's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	ridleyi	O. Thomas	1898	1	Pipistrellus_Ridleyi	Thomas, O. (1898). Description of a new bat from Selangore. Annals and Magazine of Natural History, Ser. 7, 1, 361.	https://www.biodiversitylibrary.org/item/53361#page/376/mode/1up	BM 1898.3.13.5		"Selangor, Malay Peninsula [= Peninsular Malaysia]."			ridleyi (O. Thomas, 1898)	NA	NA	Thailand|Malaysia|Indonesia|Brunei	Asia	Indomalaya	NT	0	0	0	Myotis_ridleyi	0	sciname match	Myotis_ridleyi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14194	Myotis ridleyi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	ridleyi	Thomas, 1898		20000000	Myotis ridleyi	Near Threatened	A2c+3c	2020	2018-09-10 00:00:00 UTC	3.1	English	This bat is listed as Near Threatened as it is dependent on forest habitat, and, having been recorded mainly from low altitudes, the population has probably undergone a decline of 25-30% over the last 15 years (three generations; generation length = 5 years, Pacifici et al. 2013) due to significant loss of its forest habitat due to logging, agriculture, plantations and forest fires. This rate of decline is predicted to continue for the next fifteen years. The species is close to qualifying for Vulnerable under criteria A2 and A3.	It is probably a understorey, forest dependent species. It has been recorded from caves, and found roosting in a small group under a house in forest on Borneo (Payne et al. 1985). They often forage over small streams in forest, and so perhaps it is water dependent. There were about 15-20 individuals were observed foraging over small streams in the forest trail of Gunung Mulu National Park, Sarawak (I. Azhar pers. comm). In Kawag Rainforest Lodge, Sabah, this species have been found roosting under a boulder near a waterfall (I. Azhar pers. comm). They have also been found roosting under rock near streams and also in fallen logs (S. Bumrungsri pers. comm). It was collected from pristine lowland evergreen forest in Thailand near a peat swamp, and was also found in peat swamp in Peninsular Malaysia (Bumrungsri, et al. 2006). There are no caves in the vicinity of the locality in Thailand suggesting that it may roost elsewhere (Bumrungsri et al. 2006).	Habitat loss due to logging, agriculture, plantations and forest fires represents a major threat to this species.	This is a fairly uncommon species and the global population has probably undergone a significant decline over the last 15 years due to significant loss of its forest habitat; the decline is expected to continue for the next 15 years.	Decreasing	This species is known from Peninsular Malaysia and extreme southeastern Thailand (Bumrungsri et al. 2006), few localities in Malaysian Borneo (Kubah National Park, Gunung Mulu National Park (Sarawak), Kawag Forest Lodge, Imbak Canyon Conservation Area, Gua Madai (Sabah), unpublished data) and also from Kalimantan in Indonesian Borneo (Puri 1997). Specimens from Sumatra were identified as M. annectans by Hill and Topal (1973) and the species may not occur there. The one locality in Thailand is at 100 m asl.		Terrestrial	It has been recorded on Borneo from Kayan Mentarang National Park (Puri 1997) and is found also in other protected areas throughout its range. Most of the records from Malaysian Borneo were from totally protected areas. This species has only been rarely collected and appears to have an ecology that is different from other Myotis and therefore needs to be studied further.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	ridleyi	Thomas	1898	0	Ann. Mag. Nat. Hist.	ser. 7, 1: 361	Ridley's Myotis	None.	Malaysia, Selangor (= Kepong).	W Malaysia, Sumatra, Borneo.	Not listed.	Near Threatened	Transferred from Pipistrellus ; see Medway (1978); also see Hill and Topál (1973).	Myotis ridleyi	1005461	23	Ridley's Myotis	Ridley's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	ridleyi	O. Thomas	1898	1	Pipistrellus_Ridleyi	Thomas, O. (1898). Description of a new bat from Selangore. Annals and Magazine of Natural History, Ser. 7, 1, 361.	https://www.biodiversitylibrary.org/item/53361#page/376/mode/1up	BM 1898.3.13.5		"Selangor, Malay Peninsula [= Peninsular Malaysia]."			ridleyi (O. Thomas, 1898)	NA	NA				Thailand|Malaysia|Indonesia|Brunei	Asia	Indomalaya	NT	0	0	0	Myotis_ridleyi	0	sciname match	Myotis_ridleyi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_ridleyi	1005461	23	Ridley's Myotis	Ridley's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	ridleyi	O. Thomas	1	Pipistrellus Ridleyi	Thomas, O. 1898-05-01. Description of a new bat from Selangore. Annals and Magazine of Natural History (7)1(5):360-362.	https://www.biodiversitylibrary.org/page/19365574	BMNH:Mamm:1898.3.13.5	holotype	https://data.nhm.ac.uk/object/31329f56-420d-4545-a070-fc1a3aa86da7	"Selangor, Malay Peninsula [= Peninsular Malaysia]."			NA	NA				Thailand|Malaysia|Indonesia|Brunei	Asia	Indomalaya	NT	0	0	0	Myotis_ridleyi	0	sciname match	Myotis_ridleyi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	ridleyi	Thomas	1898	1	Ann. Mag. Nat. Hist.	ser. 7, 1: 361	Ridley's Myotis	None.	Malaysia, Selangor (= Kepong).	W Malaysia, Sumatra, Borneo.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14194/22062376/' target='_blank'>Near Threatened</a>	Transferred from Pipistrellus; see Medway (1978); also see Hill and Topál (1973).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis ridleyi; Myotis ridleyi; Myotis ridleyi; Myotis ridleyi; Myotis ridleyi; Myotis ridleyi; ridleyi; Murin de Ridley; Ridley-Langful3fledermaus; Ratonero de Ridley; Ridley's Bat; Ridley's Myotis; Ridley's Bat; Ridley's Myotis; Ridley's Myotis; M. ridleyi