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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L946	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis planiceps	Myotis planiceps	Myotis planiceps	Myotis planiceps	Myotis planiceps	Myotis planiceps	Myotis planiceps	Myotis planiceps	Myotis planiceps	Myotis planiceps	Myotis planiceps	Myotis planiceps	Myotis planiceps	Myotis planiceps	Myotis planiceps		[MSW2] Subgenus Selysius. See Matson (1975, Mammalian Species, 60).; [MSW3] See Matson (1975).; [HMW] Myotis planiceps R. H. Baker, 1955 , “ 7 mi. [= 11 km ] Sand 4 mi. [= 6 km ] E Bella Unién, 7200 fi. [= 2195 m ], Coahuila ,” Mexico . Subgenus Pyzonix; lucifugus species group. Monotypic.; [batnames2022] See Matson (1975).; [batnames2023] See Matson (1975).; [batnames2025_1.7] See Matson (1975).														planiceps				planiceps	planiceps, planicpes			planiceps R. H. Baker, 1955|planicpes Mammal Diversity Database, 2018 [incorrect subsequent spelling | unpublished electronic]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Flat-headed myotis	N Mexico	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis planiceps	Mexico, Coahuila, 7 mi. (11 km) S. and 4 mi. (6 km) E. Bella Union, 7200 ft. (2195 m).	Baker	1955	Proc. Biol. Soc. Wash., 68:165.	Distribution: Confined to a small area in northeastern Mexico.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Flat-headed myotis	N Mexico	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Baker	1955	Proc. Biol. Soc. Washington, 68:165.	Subgenus Selysius. See Matson (1975, Mammalian Species, 60).	Coahuila, Nuevo Leon, and Zacatecas (Mexico).	Mexico, Coahuila, 7 mi. (11 km) S and 4 mi. (6 km) E Bella Union, 7,200 ft. (2,195 M).		BAKER	1955	Size small (forearm length, 25 28 mm; condylobasal length, 13-15 mm). Braincase greatly flattened and rostrum relatively slender. Middle upper premolar in toothrow.	Distribution: Confined to a small area in northeastern Mexico.	No subspecies.		104	species	M. planiceps	BAKER	1955	Selysius	subgenus	Myotis planiceps				Size small (forearm length, 25 28 mm; condylobasal length, 13-15 mm). Braincase greatly flattened and rostrum relatively slender. Middle upper premolar in toothrow.	No subspecies.		37. M. planiceps BAKER 1955 [leibii group].	37	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis planiceps	Myotis		planiceps	Baker		1955		Proc. Biol. Soc. Wash.	68		165		Flat-headed Myotis	Mexico, Coahuila, 7 mi. (11 km) S and 4 mi. (6 km) E Bella Union, 7,200 ft. (2,195 m).	Coahuila, Nuevo León, and Zacatecas (Mexico).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Critically Endangered.		See Matson (1975).	4C3D87E8FF536AEDFA4B91961C1EB82A	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	930	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF536AEDFA4B91961C1EB82A.xml	Myotis planiceps	Vespertilionidae	Myotis	planiceps	R. H. Baker	1955	Murin a téte plate @fr | Flachkopf-Mausohr @de | Ratonero de cabeza plana @es	Myotis planiceps R. H. Baker, 1955 , “ 7 mi. [= 11 km ] Sand 4 mi. [= 6 km ] E Bella Unién, 7200 fi. [= 2195 m ], Coahuila ,” Mexico . Subgenus Pyzonix; lucifugus species group. Monotypic.	Endemic to Mexico , restricted to a very small area in Coahuila , Nuevo Ledn, and Zacatecas .	Head-body ¢.30 51 mm , tail 21-25 mm , ear ¢. 10 mm , hindfoot 7-8 mm , forearm 26-5-27- 5 mm ; weight 5-7 g . The Flat-headed Myotis is morphologically distinct from its New World congeners. Fur is long and silky. Dorsal hairs are 8-10 mm , with blackish bases and cinnamon-brown tips. Ventral hairs are bicolored, with dark brown bases and buff tips and strong contrast between bases and tips. Ears are proportionally long, extending forward halfway from eye to nostril. Membranes are dark brown; plagiopatagium is broadly attached to foot at bases of toes. Upper and lower surfaces of uropatagium are barely covered with hairs. Skull is moderate in size (greatest skull lengths 13-5-14- 5 mm ) and flattened, more than in any other New World Myotis , sagittal crest is absent, and lambdoidal crests are present but very low; and rostrum narrows anteriorly. Teeth are small relative to palatal surface; occlusal surfaces of premolars are subequal in size, not crowded, and well-spaced; and P* and P* are very similar.	Coniferous and mixed forests in the northern parts of Sierra Madre Oriental and surrounding habitats, at elevations of 1000-3200 m .	Limited captures suggest that the Flat-headed Myotis is an aerial insectivore that forages in forested areas and near water bodies. Stomach contents of one specimen had dipterans ( Tachinidae and Therevidae ) and fragments of some lepidopteran.	A male Flat-headed Myotis with scrotal testes was captured in mid-September. Seven females appeared to be lactating with no embryos when captured in June.	The flattened skull of the Flat-headed Myotis is possibly an adaptation for roosting in crevices. Echolocation calls of indoor flying individuals were short (2-3 milliseconds), broadband (40-45 kHz), and steep FM. They swept from 80-90 kHz at the beginning of the call to 40-42 kHz at the end. Pulse interval was ¢.60-70 milliseconds. In open space, terminal frequency tended to decrease slightly (c.1 kHz); call duration and pulse interval increased from 2-3 milliseconds to 4 milliseconds and from ¢.60-70 milliseconds to 90 milliseconds, respectively, compared with indoor recordings. Bandwidth was similar (40-45 kHz).	No information.	Classified as Endangered on The IUCN Red List. The Flatheaded Myotis has an extremely restricted distribution, there are less than 250 mature individuals, and the population is probably declining because of wood exploitation and other disturbances in land use. Discovery of a new population near the known distribution in 2005 indicates it is not extinct, as it was once supposed to be.	Arroyo-Cabrales & Ospina-Garces (2016d) | Arroyo-Cabrales etal. (2005) | Baker (1955) | Jimenez-Guzman (1968, 2014) | Matson (1975)	https://zenodo.org/record/6398715/files/figure.png	382. Flat-headed Myotis Myotis planiceps French: Murin a téte plate / German: Flachkopf-Mausohr / Spanish: Ratonero de cabeza plana Taxonomy. Myotis planiceps R. H. Baker, 1955 , “ 7 mi. [= 11 km ] Sand 4 mi. [= 6 km ] E Bella Unién, 7200 fi. [= 2195 m ], Coahuila ,” Mexico . Subgenus Pyzonix; lucifugus species group. Monotypic. Distribution. Endemic to Mexico , restricted to a very small area in Coahuila , Nuevo Ledn, and Zacatecas . Descriptive notes. Head-body ¢.30 51 mm , tail 21-25 mm , ear ¢. 10 mm , hindfoot 7-8 mm , forearm 26-5-27- 5 mm ; weight 5-7 g . The Flat-headed Myotis is morphologically distinct from its New World congeners. Fur is long and silky. Dorsal hairs are 8-10 mm , with blackish bases and cinnamon-brown tips. Ventral hairs are bicolored, with dark brown bases and buff tips and strong contrast between bases and tips. Ears are proportionally long, extending forward halfway from eye to nostril. Membranes are dark brown; plagiopatagium is broadly attached to foot at bases of toes. Upper and lower surfaces of uropatagium are barely covered with hairs. Skull is moderate in size (greatest skull lengths 13-5-14- 5 mm ) and flattened, more than in any other New World Myotis , sagittal crest is absent, and lambdoidal crests are present but very low; and rostrum narrows anteriorly. Teeth are small relative to palatal surface; occlusal surfaces of premolars are subequal in size, not crowded, and well-spaced; and P* and P* are very similar. Habitat. Coniferous and mixed forests in the northern parts of Sierra Madre Oriental and surrounding habitats, at elevations of 1000-3200 m . Food and Feeding. Limited captures suggest that the Flat-headed Myotis is an aerial insectivore that forages in forested areas and near water bodies. Stomach contents of one specimen had dipterans ( Tachinidae and Therevidae ) and fragments of some lepidopteran. Breeding. A male Flat-headed Myotis with scrotal testes was captured in mid-September. Seven females appeared to be lactating with no embryos when captured in June. Activity patterns. The flattened skull of the Flat-headed Myotis is possibly an adaptation for roosting in crevices. Echolocation calls of indoor flying individuals were short (2-3 milliseconds), broadband (40-45 kHz), and steep FM. They swept from 80-90 kHz at the beginning of the call to 40-42 kHz at the end. Pulse interval was ¢.60-70 milliseconds. In open space, terminal frequency tended to decrease slightly (c.1 kHz); call duration and pulse interval increased from 2-3 milliseconds to 4 milliseconds and from ¢.60-70 milliseconds to 90 milliseconds, respectively, compared with indoor recordings. Bandwidth was similar (40-45 kHz). Movements, Home range and Social organization. No information. Status and Conservation. Classified as Endangered on The IUCN Red List. The Flatheaded Myotis has an extremely restricted distribution, there are less than 250 mature individuals, and the population is probably declining because of wood exploitation and other disturbances in land use. Discovery of a new population near the known distribution in 2005 indicates it is not extinct, as it was once supposed to be. Bibliography. Arroyo-Cabrales & Ospina-Garces (2016d), Arroyo-Cabrales etal. (2005), Baker (1955), Jimenez-Guzman (1968, 2014), Matson (1975).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis planiceps	Myotis	Unassigned-Myotis	planiceps	Baker	1955	0	Proc. Biol. Soc. Wash.	70:45:00	Flat-headed Myotis	None.	Mexico, Coahuila, 7 mi. (11 km) S and 4 mi. (6 km) E Bella Union, 7,200 ft. (2,195 m).	Coahuila, Nuevo León, and Zacatecas (Mexico).	Not listed.	Endangered	See Matson (1975).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis planiceps	23	Flat-headed Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	planiceps	R. H. Baker	1955	0	Myotis_planiceps	Baker, R. H. (1955). A new species of bat (genus Myotis) from Coahuila, Mexico. Proceedings of the Biological Society of Washington, 68, 165.	https://www.biodiversitylibrary.org/item/107489#page/179/mode/1up	KU 48242		"7 mi. [= 11 km] S and 4 mi. [= 6 km] E Bella Unión, 7200 ft. [= 2195 m], Coahuila," Mexico.			planiceps R. H. Baker, 1955	NA	NA	Mexico	North America	Nearctic	EN	0	0	0	Myotis_planiceps	0	sciname match	Myotis_planiceps	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14191	Myotis planiceps	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	planiceps	Baker, 1955		20000000	Myotis planiceps	Endangered	D	2016	2015-07-20 00:00:00 UTC	3.1	English	Flat-headed Myotis is listed as Endangered because there are less than 250 mature individuals, and the population is probably declining because of wood exploitation and changes in land use (Arriaga et al. 2000). This qualifies the species for Endangered under criterion D. Currently it is known to exist within an extent of occurrence of less than 20,000 km², which also qualifies the species as Vulnerable under criterion B1. Rediscovery of a new population near the known area of distribution indicates the species is not extinct, but is still subject to serious threats.	Little is known about the ecology of Myotis planiceps . Apparently, this species is confined to montane forests. Possibly it roosts in crevices. It is insectivorous; some stomach contents contained insects. No reproductive data are known (Matson 1975). Habitat loss has been estimated at between 16-21% over the last ten years (A.D. Cuaron and P.C. de Grammont pers. comm.).	The main threat is habitat loss due to its very restricted preferences for yucca trees accompanied by pinyon pines in the boreal montane forests of the Sierra Madre Oriental, northeastern Mexico (J. Arroyo-Cabrales pers. comm.). These forests are very limited in extent and highly disturbed by human activities (logging).	It is thought that there are less than 250 mature individuals and that the population is declining (J. Arroyo-Cabrales pers. comm.).	Decreasing	The insectivorous Flat-headed Myotis is an endemic species restricted to a very small area. It occurs in Coahuila, Nuevo León, and Zacatecas (Mexico) (Simmons 2005); it was thought extinct in 1996, but then it was rediscovered in three localities, including two new ones in 2004 (Arroyo-Cabrales et al . 2005).		Terrestrial	The species is included in the Mexican regulation for species conservation NOM-59-SEMARNAT-2001 (under P category which means Endangered with extinction). The protection of this species is being undertaken by the Program for Mexican Bat Conservation (PCMM) (J. Arroyo-Cabrales pers. comm.).	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	planiceps	Baker	1955	0	Proc. Biol. Soc. Wash.	70:45:00	Flat-headed Myotis	None.	Mexico, Coahuila, 7 mi. (11 km) S and 4 mi. (6 km) E Bella Union, 7,200 ft. (2,195 m).	Coahuila, Nuevo León, and Zacatecas (Mexico).	Not listed.	Endangered	See Matson (1975).	Myotis planiceps	1005458	23	Flat-headed Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	planiceps	R. H. Baker	1955	0	Myotis_planiceps	Baker, R. H. (1955). A new species of bat (genus Myotis) from Coahuila, Mexico. Proceedings of the Biological Society of Washington, 68, 165.	https://www.biodiversitylibrary.org/item/107489#page/179/mode/1up	KU 48242		"7 mi. [= 11 km] S and 4 mi. [= 6 km] E Bella Unión, 7200 ft. [= 2195 m], Coahuila," Mexico.			planiceps R. H. Baker, 1955	NA	NA				Mexico	North America	Nearctic	EN	0	0	0	Myotis_planiceps	0	sciname match	Myotis_planiceps	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_planiceps	1005458	23	Flat-headed Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	planiceps	R. H. Baker	0	Myotis planiceps	Baker, R.H. 1955-12-31. A new species of bat (genus _Myotis_) from Coahuila, Mexico. Proceedings of the Biological Society of Washington 68:165-166.	https://www.biodiversitylibrary.org/page/34549649	KU:M:48242	holotype	http://portal.vertnet.org/o/ku/kum?id=921256e2-1ed8-11e3-bfac-90b11c41863e	"7 mi. [= 11 km] S and 4 mi. [= 6 km] E Bella Unión, 7200 ft. [= 2195 m], Coahuila," Mexico.			NA	NA				Mexico	North America	Nearctic	EN	0	0	0	Myotis_planiceps	0	sciname match	Myotis_planiceps	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	planiceps	Baker	1955	0	Proc. Biol. Soc. Wash.	70:45:00	Flat-headed Myotis	None.	Mexico, Coahuila, 7 mi. (11 km) S and 4 mi. (6 km) E Bella Union, 7,200 ft. (2,195 m).	Coahuila, Nuevo León, and Zacatecas (Mexico).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14191/22066742/' target='_blank'>Endangered</a>	See Matson (1975).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis planiceps; Myotis planiceps; Myotis planiceps; Myotis planiceps; Myotis planiceps; Myotis planiceps; planiceps; Murin a téte plate; Flachkopf-Mausohr; Ratonero de cabeza plana; Flat-headed Myotis; Flat-headed Myotis; Flat-headed Myotis; M. planiceps