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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L944	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Myotis ricketti [synonym of]	N/A	N/A	Myotis pilosus	Myotis ricketti [synonym of]	Myotis pilosus	Myotis pilosus	Myotis pilosus	Myotis pilosus	Myotis pilosus	Myotis pilosus	Myotis pilosus	Myotis pilosus	Myotis pilosus		[MSW3] Myotis pilosus Peters, 1869 (type locality unknown) may be the oldest name for this species; see Ellerman and Morrison-Scott (1951) and Corbet and Hill (1992). Sometimes placed in its own subgenus Rickettia, see discussion in Findley (1972) and Corbet and Hill (1992). Reviewed in part by Bates et al. (1999) and Hendrichsen et al. (2001); also see Horácek et al. (2000), who discussed this taxon under the name pilosus.; [HMW] Vespertilio (Leuconoe) pilosus Peters, 1869 , type locality not given. Subgenus Myotis ; macrodactylus species group. See M. petax . Due to confusion with the type locality (thought to be Montevideo, Uruguay , South America), most authors used the name M. ricketti proposed by O. Thomas in 1894 and ignored the original name M. pilosus , which is the oldest and most appropriate name available. Based on phylogeography of M. pilosusin China , three lineages that correspond to distinct geographical populations were identified. Monotypic.; [batnames2022] Often called Myotis  ricketti (Thomas 1894); see Allen (1936) and Smith and Xie (2008), Ellerman and Morrison-Scott (1951), and Corbet and Hill(1992). Sometimes placed in its own subgenus Rickettia, see discussion in Findley (1972) and Corbet and Hill (1992). Reviewed in part by Bates et al. (1999) and Hendrichsen et al. (2001); also see HorÃ¡cek et al. (2000).; [MDD2022] generally known under the name ricketti, but pilosus has been shown to be the oldest name available for the species; [IUCN] This species has often used the name Myotis ricketti (Thomas, 1894), but pilosus has priority (Smith and Xie 2008). The erroneous reporting of the type locality as being in Uruguay, by Peters, led to the dual naming (Smith and Xie 2008).; [batnames2023] Often called Myotis  ricketti (Thomas 1894); see Allen (1936) and Smith and Xie (2008), Ellerman and Morrison-Scott (1951), and Corbet and Hill(1992). Sometimes placed in its own subgenus Rickettia, see discussion in Findley (1972) and Corbet and Hill (1992). Reviewed in part by Bates et al. (1999) and Hendrichsen et al. (2001); also see HorÃ¡cek et al. (2000).; [MDD2023] generally known under the name ricketti, but pilosus has been shown to be the oldest name available for the species; [MDD2025_2.0] generally known under the name ricketti, but pilosus has been shown to be the oldest name available for the species; [batnames2025_1.7] Often called Myotis ricketti (Thomas 1894); see Allen (1936) and Smith and Xie (2008), Ellerman and Morrison-Scott (1951), and Corbet and Hill(1992). Sometimes placed in its own subgenus Rickettia, see discussion in Findley (1972) and Corbet and Hill (1992). Reviewed in part by Bates et al. (1999) and Hendrichsen et al. (2001); also see HorÃ¡cek et al. (2000).; [MDD2025_2.2] generally known under the name ricketti, but pilosus has been shown to be the oldest name available for the species												pilosus 	pilosus - ricketti	pilosus, ricketti	This species has often used the name Myotis ricketti (Thomas, 1894), but pilosus has priority (Smith and Xie 2008). The erroneous reporting of the type locality as being in Uruguay, by Peters, led to the dual naming (Smith and Xie 2008).	pilosus 	pilosus - ricketti	pilosus, ricketti	pilosus, ricketti	pilosus	pilosus - ricketti	pilosus (W. C. H. Peters, 1869)|ricketti (O. Thomas, 1894)						N/A					Distribution: Confined to southern and eastern China.													PETERS	1869	Size relatively large (forearm length, 55-58 mm). Foot unusually large. Margin of plagiopatagium attached near middle of tibia. No fringe of hair on margin of uropatagium. Braincase of medium height. Rostrum relatively broad. Middle upper premolar displaced medially from the toothrow.	Distribution: Confined to southern and eastern China.	No subspecies.		109	species	M. pilosus	PETERS	1869	Leuconoe	subgenus	Myotis pilosus				Size relatively large (forearm length, 55-58 mm). Foot unusually large. Margin of plagiopatagium attached near middle of tibia. No fringe of hair on margin of uropatagium. Braincase of medium height. Rostrum relatively broad. Middle upper premolar displaced medially from the toothrow.	No subspecies.		81. M. pilosus (PETERS 1869) (= ricketti THOMAS 1894) [pilosus group],	81	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis ricketti	Myotis		ricketti	Thomas	y	1894		Ann. Mag. Nat. Hist., ser. 6	14		300		Rickett's Big-footed Myotis	China, Fukien (= Fujian), Foochow.	Fukien, Anhwei, Kiangsu, Shantung, Yunnan (China); Hong Kong; Vietnam and Laos.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).		Myotis pilosus Peters, 1869 (type locality unknown) may be the oldest name for this species; see Ellerman and Morrison-Scott (1951) and Corbet and Hill (1992). Sometimes placed in its own subgenus Rickettia, see discussion in Findley (1972) and Corbet and Hill (1992). Reviewed in part by Bates et al. (1999) and Hendrichsen et al. (2001); also see Horácek et al. (2000), who discussed this taxon under the name pilosus.	4C3D87E8FF2F6A90FF509F02145BB6A2	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	974	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF2F6A90FF509F02145BB6A2.xml	Myotis pilosus	Vespertilionidae	Myotis	pilosus		1869	Murin de Rickett @fr | Rickett-Wasserfledermaus @de | Ratonero de Rickett @es | Rickett's Big-footed Bat @en	Vespertilio (Leuconoe) pilosus Peters, 1869 , type locality not given. Subgenus Myotis ; macrodactylus species group. See M. petax . Due to confusion with the type locality (thought to be Montevideo, Uruguay , South America), most authors used the name M. ricketti proposed by O. Thomas in 1894 and ignored the original name M. pilosus , which is the oldest and most appropriate name available. Based on phylogeography of M. pilosusin China , three lineages that correspond to distinct geographical populations were identified. Monotypic.	S & E China ( Beijing , Shaanxi , Shandong , Jiangsu , Anhui , Zhejiang , Jiangxi , Fujian , Hunan , Guizhou , Guangdong , Guangxi , Yunnan , Xizang , and Hainan I), C Laos ( Khammouane Province ), N Vietnam (from Lang Son , Nghe An , Quang Binh , and Bac Kan provinces), and recently reported from Meghalaya State in NE India . Given this recent record far away from its main distribution, this species might occur in suitable habitats in Myanmar and Thailand .	Head-body 51-65 mm , tail 35-55 mm , ear 11-22- 3 mm , hindfoot 13-3-17- 5 mm , forearm 52-1-63- 5 mm ; weight 11-7-32.5 g (ranges from different sources). Compared with mainland specimens, 27 specimens from Hainan Island were smaller (forearm 48- 5-51 mm , tail 35-44 mm , ear 11-16- 2 mm , hindfoot 13-3— 16- 3 mm ; weight 117-222 g ), perhaps representing a subspecies. Rickett’s Big-footed Myotis has extreme large feet and very strong claws, well adapted for catching fish. Short fur is closely appressed to skin and velvety. Upperparts are drab, buffy gray-brown, with darker roots, and slightly darker on sides of head. Underparts are nearly off-white, with pale gray to white tips, and dark roots. Ears are medium brown and moderately large; anterior borders are smoothly convex but with well-defined notch at bases, bluntly pointed tips and straight posterior borders. Tragus is one-half the ear length and narrowly pointed. Muzzle is well covered with bristly whiskers. Tibiae are well haired above and below. Hindfeet are unusually large (c.80% the length oftibiae), with long, sharply pointed, pale, curved claws. Calcar is 9-10 mm and more than four-fifths the length of posterior border of uropatagium from foot to tip of tail. Uropatagium has some hair nearest to the body but no fringe on its margin. Wing membranes are dark brown and attach to ankle on ventral side of distal part of tibiae. Skull is medium-sized, with relatively broad rostrum and smooth dorsal profile. Anterior braincase does notrise sharply above rostrum, and there is only a very shallow concavity above posterior rostrum. Sagittal crest is poorly developed. Post-dental extension of palate is short and wide. Upper dentition is relatively weak; average length of upper tooth row (C-M?) is 8- 3 mm . Upper incisors are large, each with conspicuous supplementary cusp. Canines lack secondary cusps. P? is small, with single crown;it is situated in tooth row and usually in contact with C'. Crown area of P? is about one-half to two-thirds that of P%. P* displaced internally in tooth row and in close contact with P* which is about two-thirds the height of C' but has greater crown area. Upper molarsare typically myotodont. I, and I, are three-lobed from the front. C is about twice the height of Pail is about equal to or exceeds P, in height and crown area. P, is about one-half to two-thirds the crown area of P,, which is one-third to half that of C'. Condylo-basal lengths are 17-8-18- 8 mm ; maxillary tooth row lengths are 8-2-8- 4 mm . Chromosomal complement has 2n = 44 and FN = 52.	Strictly dependent on large areas of water (e.g. lakes, reservoir, and large still rivers) in lowland secondary forests, karst areas, and open habitats or heavily disturbed vegetation ( Vietnam ) at known elevations of 1400-3000 m .	Stomach of one Rickett’s Big-footed Myotis from Pu Mat, Vietham contained insects, but 97% of droppings from two specimens in Khammouan Limestone Area in Laos contained fish scales and a small number of insects. In a study of ¢.43 individuals from an area near Beijing , China , feces containedfish scales of Cyprinidae ( Zacco , Carassius , Phoxinus ) and insects ( Coleoptera , Lepidoptera, Homoptera, Ephemeroptera , Hemiptera , Lepidoptera , and Hymenoptera ). Scales of Z. platypus were the most abundantfish remains, and Coleoptera made up 70% of identifiable insects. These individuals flew back and forth above the water surface of the reservoir near their roosting cave.	Mating of Rickett’s Big-footed Myotis took place in October—April. Sperm was stored until ovulation occurred after hibernation in May-June. Births ofsingle young probably occurred from mid-June to July, and lactating females were reported from a temple in Du-le Si inJune-September. Sperm was present in cauda epididymis from late September to early April but absent from testes from mid-November to late August.	Flight of Rickett’s Big-footed Myotisis rapid and agile. They roost in caves. In a cave near Beijing , China , they hibernated from November to April. Echolocation calls have maximum frequency of 70-8 kHz, minimum frequency of 27-7 kHz, peak frequency of 40-6 kHz, average duration of 4-1 milliseconds, and average interpulse interval of 108-8 milliseconds. Calls of individuals in Hainan (forearm length 50 mm ) have peak frequency of 50-4 kHz, those from Henan (forearm 58- 1 mm ) 45-2 kHz, and those from Beijing (forearm 55-3-63- 5 mm ) 41-9 kHz, which might reflect subspecific differences.	Two female Rickett’s Big-footed Myotis were found together with Large Myotis (M. chinensis ) at the entrance of Lan Cat Cave, Vietnam , which was small and cylindrical and surrounded by arable land. They were captured in August-mid October, near Beijing over a water reservoir in a warm temperate zone forest, dominated by oaks and pines, ¢. 8 km away from a cave with Greater Horseshoe Bats (Rhinolophus ferrumequinum), Rufous Tube-nosed Bat ( Murina leucogaster ), and Greater Myotis ( M. myotis ). Rickett’s Big-footed Myotis is more dependent on fish than smaller trawling species of Myotis and travels considerable distances to forage. It is present in the southern part of the Funiu Mountains ( Henan , China ) in summer only, suggesting they move to the area to access greater food resources. In the Beijing area, China , cave colonies have up to 1000-2000 individuals. Summer roosts (including maternity roosts) are found in May-June in buildings or caves, some of which are used in winter. Some individuals roost in clusters, but territorial males are scattered in the cave, and other individuals are chased away when they approach a territory. Rickett’s Big-footed Myotis often shares its roosts with other bats and swifts ( Apodidae ).	Classified as Near Threatened on The IUCN Red Last. Rickett’s Big-footed Myotis has low tolerance to human disturbance. It is highly dependent on water sources forits food, and water pollution in China is a major threat. Research is needed on population status, ecology, habitat status, and conservation threats. It occurs in Tianmu Mountain National Nature Reserve ( China ), Qomolangma National Nature Reserve ( China ), and Kim Hy Nature Reserve (northern Vietnam ).	Aizpurua & Alberdi (2018) | Allen (1936, 1938b) | Amadoretal. (2018) | Bates et al. (1999) | Bianchi (1917) | Borisenko & Kruskop (2003) | Carter & Dolan (1978) | Corbet (1978) | Corbet & Hill (1992) | Csorba & Bates (2008b) | Ellerman & Morrison-Scott (1951, 1966) | Francis (2008a) | Furey & Racey (2016b) | Hendrichsen, Bates & Hayes (2001) | Honacki et al. (1982) | Horagek et al. (2000) | Hu Yiming et al. (2014) | Kawai et al. (2003) | Koopman (1993, 1994) | Kruskop (2013a) | Li Yuchun et al. (2006) | Lu Guanjun et al. (2013) | Ma Jie, Jones et al. (2003) | Ma Jie, Zhang Jinshuo et al. (2006) | Miller & Allen (1928) | Niu Hongxing, Wang Ning et al. (2007) | Peters (1869c) | Rasweiler et al. (2011) | Robinson & Webber (1998, 2000) | Ruedi, Csorba et al. (2015) | Ruedi, Stadelmann et al. (2013) | Saikia (2018) | Simmons (2005) | Stadelmann, Jacobs et al. (2004) | Stadelmann, Lin Liangkong et al. (2007) | Tang Zhanhui et al. (2012) | Thomas (1894) | Wang Sung & Xie Yan (2004) | Wang Zhe, Liang Bing et al. (2008) | Wang Zhe, Shi Qixian etal. (2008) | Wilson (2008b) | Wu Yi & Harada (2006) | Zhang Zhenzhen etal. (2009)	https://zenodo.org/record/6398975/files/figure.png	478. Ricketts Big-footed Myotis Myotis pilosus French: Murin de Rickett / German: Rickett-Wasserfledermaus / Spanish: Ratonero de Rickett Other common names: Rickett's Big-footed Bat Taxonomy. Vespertilio (Leuconoe) pilosus Peters, 1869 , type locality not given. Subgenus Myotis ; macrodactylus species group. See M. petax . Due to confusion with the type locality (thought to be Montevideo, Uruguay , South America), most authors used the name M. ricketti proposed by O. Thomas in 1894 and ignored the original name M. pilosus , which is the oldest and most appropriate name available. Based on phylogeography of M. pilosusin China , three lineages that correspond to distinct geographical populations were identified. Monotypic. Distribution. S & E China ( Beijing , Shaanxi , Shandong , Jiangsu , Anhui , Zhejiang , Jiangxi , Fujian , Hunan , Guizhou , Guangdong , Guangxi , Yunnan , Xizang , and Hainan I), C Laos ( Khammouane Province ), N Vietnam (from Lang Son , Nghe An , Quang Binh , and Bac Kan provinces), and recently reported from Meghalaya State in NE India . Given this recent record far away from its main distribution, this species might occur in suitable habitats in Myanmar and Thailand . Descriptive notes. Head-body 51-65 mm , tail 35-55 mm , ear 11-22- 3 mm , hindfoot 13-3-17- 5 mm , forearm 52-1-63- 5 mm ; weight 11-7-32.5 g (ranges from different sources). Compared with mainland specimens, 27 specimens from Hainan Island were smaller (forearm 48- 5-51 mm , tail 35-44 mm , ear 11-16- 2 mm , hindfoot 13-3— 16- 3 mm ; weight 117-222 g ), perhaps representing a subspecies. Rickett’s Big-footed Myotis has extreme large feet and very strong claws, well adapted for catching fish. Short fur is closely appressed to skin and velvety. Upperparts are drab, buffy gray-brown, with darker roots, and slightly darker on sides of head. Underparts are nearly off-white, with pale gray to white tips, and dark roots. Ears are medium brown and moderately large; anterior borders are smoothly convex but with well-defined notch at bases, bluntly pointed tips and straight posterior borders. Tragus is one-half the ear length and narrowly pointed. Muzzle is well covered with bristly whiskers. Tibiae are well haired above and below. Hindfeet are unusually large (c.80% the length oftibiae), with long, sharply pointed, pale, curved claws. Calcar is 9-10 mm and more than four-fifths the length of posterior border of uropatagium from foot to tip of tail. Uropatagium has some hair nearest to the body but no fringe on its margin. Wing membranes are dark brown and attach to ankle on ventral side of distal part of tibiae. Skull is medium-sized, with relatively broad rostrum and smooth dorsal profile. Anterior braincase does notrise sharply above rostrum, and there is only a very shallow concavity above posterior rostrum. Sagittal crest is poorly developed. Post-dental extension of palate is short and wide. Upper dentition is relatively weak; average length of upper tooth row (C-M?) is 8- 3 mm . Upper incisors are large, each with conspicuous supplementary cusp. Canines lack secondary cusps. P? is small, with single crown;it is situated in tooth row and usually in contact with C'. Crown area of P? is about one-half to two-thirds that of P%. P* displaced internally in tooth row and in close contact with P* which is about two-thirds the height of C' but has greater crown area. Upper molarsare typically myotodont. I, and I, are three-lobed from the front. C is about twice the height of Pail is about equal to or exceeds P, in height and crown area. P, is about one-half to two-thirds the crown area of P,, which is one-third to half that of C'. Condylo-basal lengths are 17-8-18- 8 mm ; maxillary tooth row lengths are 8-2-8- 4 mm . Chromosomal complement has 2n = 44 and FN = 52. Habitat. Strictly dependent on large areas of water (e.g. lakes, reservoir, and large still rivers) in lowland secondary forests, karst areas, and open habitats or heavily disturbed vegetation ( Vietnam ) at known elevations of 1400-3000 m . Food and Feeding. Stomach of one Rickett’s Big-footed Myotis from Pu Mat, Vietham contained insects, but 97% of droppings from two specimens in Khammouan Limestone Area in Laos contained fish scales and a small number of insects. In a study of ¢.43 individuals from an area near Beijing , China , feces containedfish scales of Cyprinidae ( Zacco , Carassius , Phoxinus ) and insects ( Coleoptera , Lepidoptera, Homoptera, Ephemeroptera , Hemiptera , Lepidoptera , and Hymenoptera ). Scales of Z. platypus were the most abundantfish remains, and Coleoptera made up 70% of identifiable insects. These individuals flew back and forth above the water surface of the reservoir near their roosting cave. Breeding. Mating of Rickett’s Big-footed Myotis took place in October—April. Sperm was stored until ovulation occurred after hibernation in May-June. Births ofsingle young probably occurred from mid-June to July, and lactating females were reported from a temple in Du-le Si inJune-September. Sperm was present in cauda epididymis from late September to early April but absent from testes from mid-November to late August. Activity patterns. Flight of Rickett’s Big-footed Myotisis rapid and agile. They roost in caves. In a cave near Beijing , China , they hibernated from November to April. Echolocation calls have maximum frequency of 70-8 kHz, minimum frequency of 27-7 kHz, peak frequency of 40-6 kHz, average duration of 4-1 milliseconds, and average interpulse interval of 108-8 milliseconds. Calls of individuals in Hainan (forearm length 50 mm ) have peak frequency of 50-4 kHz, those from Henan (forearm 58- 1 mm ) 45-2 kHz, and those from Beijing (forearm 55-3-63- 5 mm ) 41-9 kHz, which might reflect subspecific differences. Movements, Home range and Social organization. Two female Rickett’s Big-footed Myotis were found together with Large Myotis (M. chinensis ) at the entrance of Lan Cat Cave, Vietnam , which was small and cylindrical and surrounded by arable land. They were captured in August-mid October, near Beijing over a water reservoir in a warm temperate zone forest, dominated by oaks and pines, ¢. 8 km away from a cave with Greater Horseshoe Bats (Rhinolophus ferrumequinum), Rufous Tube-nosed Bat ( Murina leucogaster ), and Greater Myotis ( M. myotis ). Rickett’s Big-footed Myotis is more dependent on fish than smaller trawling species of Myotis and travels considerable distances to forage. It is present in the southern part of the Funiu Mountains ( Henan , China ) in summer only, suggesting they move to the area to access greater food resources. In the Beijing area, China , cave colonies have up to 1000-2000 individuals. Summer roosts (including maternity roosts) are found in May-June in buildings or caves, some of which are used in winter. Some individuals roost in clusters, but territorial males are scattered in the cave, and other individuals are chased away when they approach a territory. Rickett’s Big-footed Myotis often shares its roosts with other bats and swifts ( Apodidae ). Status and Conservation. Classified as Near Threatened on The IUCN Red Last. Rickett’s Big-footed Myotis has low tolerance to human disturbance. It is highly dependent on water sources forits food, and water pollution in China is a major threat. Research is needed on population status, ecology, habitat status, and conservation threats. It occurs in Tianmu Mountain National Nature Reserve ( China ), Qomolangma National Nature Reserve ( China ), and Kim Hy Nature Reserve (northern Vietnam ). Bibliography. Aizpurua & Alberdi (2018), Allen (1936, 1938b), Amadoretal. (2018), Bates et al. (1999), Bianchi (1917), Borisenko & Kruskop (2003), Carter & Dolan (1978), Corbet (1978), Corbet & Hill (1992), Csorba & Bates (2008b), Ellerman & Morrison-Scott (1951, 1966), Francis (2008a), Furey & Racey (2016b), Hendrichsen, Bates & Hayes (2001), Honacki et al. (1982), Horagek et al. (2000), Hu Yiming et al. (2014), Kawai et al. (2003), Koopman (1993, 1994), Kruskop (2013a), Li Yuchun et al. (2006), Lu Guanjun et al. (2013), Ma Jie, Jones et al. (2003), Ma Jie, Zhang Jinshuo et al. (2006), Miller & Allen (1928), Niu Hongxing, Wang Ning et al. (2007), Peters (1869c), Rasweiler et al. (2011), Robinson & Webber (1998, 2000), Ruedi, Csorba et al. (2015), Ruedi, Stadelmann et al. (2013), Saikia (2018), Simmons (2005), Stadelmann, Jacobs et al. (2004), Stadelmann, Lin Liangkong et al. (2007), Tang Zhanhui et al. (2012), Thomas (1894), Wang Sung & Xie Yan (2004), Wang Zhe, Liang Bing et al. (2008), Wang Zhe, Shi Qixian etal. (2008), Wilson (2008b), Wu Yi & Harada (2006), Zhang Zhenzhen etal. (2009).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis pilosus	Myotis	Unassigned-Myotis	pilosus	Peters	1869	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	1875:43:00	Rickett's Big-footed Myotis	 ricketti Thomas, 1894.	China, Fukien (= Fujian), Foochow.	Fukien, Anhwei, Kiangsu, Shantung, Yunnan (China); Hong Kong; Vietnam and Laos.	Not listed.	Vulnerable	Often called Myotis  ricketti (Thomas 1894); see Allen (1936) and Smith and Xie (2008), Ellerman and Morrison-Scott (1951), and Corbet and Hill(1992). Sometimes placed in its own subgenus Rickettia, see discussion in Findley (1972) and Corbet and Hill (1992). Reviewed in part by Bates et al. (1999) and Hendrichsen et al. (2001); also see HorÃ¡cek et al. (2000).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis pilosus	23	Rickett's Big-footed Myotis	Rickett's Big-footed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	pilosus	W. Peters	1869	1	Vespertilio_(LeuconoÃ«)_pilosus	Peters, W. C. H. (1869). Ãœber neue oder weniger bekannte Flederthiere, besonders des Pariser Museums. Monatsberichte der KÃ¶niglich Preussischen Akademie der Wissenschaften zu Berlin, 1869, 403.	https://www.biodiversitylibrary.org/item/111869#page/437/mode/1up	MNHN 1869-14		type locality not given.			pilosus (W. Peters, 1869)|ricketti (O. Thomas, 1894)	generally known under the name ricketti, but pilosus has been shown to be the oldest name available for the species	Smith, A. T., & Xie, Y. (2008). A Guide to the Mammals of China. Princeton University Press, Princeton, New Jersey.|Lu, G., Lin, A., Luo, J., Blondel, D. V., Meiklejohn, K. A., Sun, K., & Feng, J. (2013). Phylogeography of the Rickett's big-footed bat, Myotis pilosus (Chiroptera: Vespertilionidae): a novel pattern of genetic structure of bats in China. BMC evolutionary biology, 13(1), 1-11.	China|Laos|Vietnam|India	Asia	Palearctic|Indomalaya	VU	0	0	0	Myotis_pilosus	0	manual	Myotis_ricketti	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14193	Myotis pilosus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	pilosus	(Peters, 1869)	This species has often used the name Myotis ricketti (Thomas, 1894), but pilosus has priority (Smith and Xie 2008). The erroneous reporting of the type locality as being in Uruguay, by Peters, led to the dual naming (Smith and Xie 2008).	20000000	Myotis pilosus	Vulnerable	A4c	2019	2018-08-22 00:00:00 UTC	3.1	English	This bat is assessed as Vulnerable as it is a widely distributed habitat specialist with a low tolerance to human disturbance. It is highly dependent on water sources for its food, and water pollution in China continues to be a major threat. Although there are no quantitative data, it is suspected that this species has declined more than 30% over the last 10 years due to water pollution and is expected to continue to decline by a similar rate for at least another 5 years (GL = 5 years; Pacifici et al. 2013).	This species is strictly dependent on water since its diet is comprised of fish. This is the most specialized, fish eating bat within the region. It has low tolerance to human disturbance (Csorba and Bates 2006).	This is a widely distributed freshwater habitat specialist, which has low tolerance to human disturbance. Highly dependent on water sources for its food, water pollution (domestic and commercial/industrial) in China (the bulk of its distribution) is a major threat. In Southeast Asia, water sources are not as degraded and therefore, it does not face a similar threat as it does in China. This species may be hunted for food locally.	This species is widespread (T. Jiang pers. comm. August 2018) and is thought to be declining across its range due to decreasing water quality.	Decreasing	This species occurs in China (including Hong Kong), Lao PDR, Hong Kong, and Viet Nam (Simmons 2005). In China, this species occurs in the provinces of Anhui, Beijing, Fujian, Guangdong, Guangxi, Shandong, Shanxi, Yunnan, Zhejiang (Smith and Xie 2008), Jiangxi (CSIS 2008), Guizhou (Jiang et al. 2008), and may be present in Hunan and Jiangsu. The distribution in Lao PDR and Viet Nam is very scattered (Csorba, G. pers. comm.). In 2004, 27 specimens of this species were collected on Hainan, which the authors noted were smaller than those of mainland specimens (Li et al. 2006).	This species may be targeted for subsistence as it is a relatively large species, but as it is a rare form impact on the population is likely to be low (G. Csorba pers. comm.).	Terrestrial	This species occurs in Tianmushan Nature Reserve (CSIS 2008), but it is not known if the species is present in any other protected areas. It has been regionally Red Listed in China as Least Concern (Wang and Xie 2004), but recently it has been considered as Near Threatened (Jiang et al. 2016). Research is needed in the areas of population status, biology and ecology, habitat status, and threats.	Indomalayan|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	pilosus	Peters	1869	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	1875:43:00	Rickett's Big-footed Myotis	 ricketti Thomas, 1894.	China, Fukien (= Fujian), Foochow.	Fukien, Anhwei, Kiangsu, Shantung, Yunnan (China); Hong Kong; Vietnam and Laos.	Not listed.	Vulnerable	Often called Myotis  ricketti (Thomas 1894); see Allen (1936) and Smith and Xie (2008), Ellerman and Morrison-Scott (1951), and Corbet and Hill(1992). Sometimes placed in its own subgenus Rickettia, see discussion in Findley (1972) and Corbet and Hill (1992). Reviewed in part by Bates et al. (1999) and Hendrichsen et al. (2001); also see HorÃ¡cek et al. (2000).	Myotis pilosus	1005457	23	Rickett's Big-footed Myotis	Rickett's Big-footed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	pilosus	W. Peters	1869	1	Vespertilio_(LeuconoÃ«)_pilosus	Peters, W. C. H. (1869). Ãœber neue oder weniger bekannte Flederthiere, besonders des Pariser Museums. Monatsberichte der KÃ¶niglich Preussischen Akademie der Wissenschaften zu Berlin, 1869, 403.	https://www.biodiversitylibrary.org/item/111869#page/437/mode/1up	MNHN 1869-14		type locality not given.			pilosus (W. Peters, 1869)|ricketti (O. Thomas, 1894)	generally known under the name ricketti, but pilosus has been shown to be the oldest name available for the species	Smith, A. T., & Xie, Y. (2008). A Guide to the Mammals of China. Princeton University Press, Princeton, New Jersey.|Lu, G., Lin, A., Luo, J., Blondel, D. V., Meiklejohn, K. A., Sun, K., & Feng, J. (2013). Phylogeography of the Rickett's big-footed bat, Myotis pilosus (Chiroptera: Vespertilionidae): a novel pattern of genetic structure of bats in China. BMC evolutionary biology, 13(1), 1-11.				China|Laos|Vietnam|India	Asia	Palearctic|Indomalaya	VU	0	0	0	Myotis_pilosus	0	manual	Myotis_ricketti	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_pilosus	1005457	23	Rickett's Big-footed Myotis	Rickett's Big-footed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	pilosus	W. C. H. Peters	1	Vespertilio (LeuconoÃ«) pilosus	Peters, W.C.H. 1869. Bemerkungen Ã¼ber neue oder weniger bekannte Flederthiere, besonders des Pariser Museums. Monatsberichte der KÃ¶niglichen Preussischen Akademie der Wissenschaften zu Berlin 1869:393-408.	https://www.biodiversitylibrary.org/page/36276616	MNHN-ZM-MO-1869-14	holotype	http://coldb.mnhn.fr/catalognumber/mnhn/zm/mo-1869-14	type locality not given.			generally known under the name ricketti, but pilosus has been shown to be the oldest name available for the species	Smith, A. T., & Xie, Y. (2008). A Guide to the Mammals of China. Princeton University Press, Princeton, New Jersey.|Lu, G., Lin, A., Luo, J., Blondel, D. V., Meiklejohn, K. A., Sun, K., & Feng, J. (2013). Phylogeography of the Rickett's big-footed bat, Myotis pilosus (Chiroptera: Vespertilionidae): a novel pattern of genetic structure of bats in China. BMC evolutionary biology, 13(1), 1-11.				China|Laos|Vietnam|India	Asia	Palearctic|Indomalaya	VU	0	0	0	Myotis_pilosus	0	manual	Myotis_ricketti	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	pilosus	Peters	1869	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	1875:43:00	Rickett's Big-footed Myotis	ricketti Thomas, 1894.	China, Fukien (= Fujian), Foochow.	Fukien, Anhwei, Kiangsu, Shantung, Yunnan (China); Hong Kong; Vietnam and Laos.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14193/22062554/' target='_blank'>Vulnerable</a>	Often called Myotis ricketti (Thomas 1894); see Allen (1936) and Smith and Xie (2008), Ellerman and Morrison-Scott (1951), and Corbet and Hill(1992). Sometimes placed in its own subgenus Rickettia, see discussion in Findley (1972) and Corbet and Hill (1992). Reviewed in part by Bates et al. (1999) and Hendrichsen et al. (2001); also see HorÃ¡cek et al. (2000).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis ricketti; Myotis pilosus; Myotis pilosus; Myotis pilosus; Myotis pilosus; Myotis pilosus; ricketti; pilosus; ricketti; Murin de Rickett; Rickett-Wasserfledermaus; Ratonero de Rickett; Rickett's Big-footed Bat; Rickett's Big-footed Myotis; Rickett's Big-footed Bat; Rickett's Big-footed Myotis; Rickett's Big-footed Myotis; M. pilosus
