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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L943	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Myotis keaysi pilosotibialis	Myotis pilosatibialis	Myotis pilosatibialis	Myotis pilosatibialis	N/A	Myotis pilosatibialis	Myotis pilosatibialis	Myotis pilosatibialis	Myotis pilosatibialis	Myotis pilosatibialis		[HMW] Myotis keaysi pilosatibialis LaVal, 1973 , “ 1 km W Talanga, Francisco Morazan [= Morazrin], Honduras , elevation 750 m .” Subgenus Pyzonix; ruber species group (7 species). Myotis pilosatibialis became a distinct species after analysis of morphological data in 2014. Monotypic.; [batnames2022] Distinct from keaysi; see Mantilla-Meluk and Munoz-Garay (2014). May not be monophyletic; see Novaes et al. (2022).; [MDD2022] split from M. keaysi; [batnames2023] Distinct from keaysi; see Mantilla-Meluk and Munoz-Garay (2014). May not be monophyletic; see Novaes et al. (2022).; [MDD2023] split from M. keaysi; [MDD2025_2.0] split from M. keaysi; [batnames2025_1.7] Distinct from keaysi; see Mantilla-Meluk and Munoz-Garay (2014). May not be monophyletic; see Novaes et al. (2022).; [MDD2025_2.2] split from M. keaysi														pilosatibialis				pilosatibialis 	pilosatibialis, pilosotibialis			pilosatibialis LaVal, 1973|pilosotibialis Koopman, 1994 [incorrect subsequent spelling]						N/A																																								NA																											4C3D87E8FF516AEEFA5D95EA1652B7E6	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	932	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF516AEEFA5D95EA1652B7E6.xml	Myotis pilosatibialis	Vespertilionidae	Myotis	pilosatibialis	LaVal	1973	Murin a pattes velues @fr | Nordliches Haarfi Riges Mausohr @de | Ratonero de patas peludas @es	Myotis keaysi pilosatibialis LaVal, 1973 , “ 1 km W Talanga, Francisco Morazan [= Morazrin], Honduras , elevation 750 m .” Subgenus Pyzonix; ruber species group (7 species). Myotis pilosatibialis became a distinct species after analysis of morphological data in 2014. Monotypic.	From EC Mexico S through Central America into Colombia and N Venezuela ; also on Trinidad I.	Head-body c. 45-51 mm , tail 34-39 mm , ear 12-14 mm , hindfoot 7-9 mm ; forearm 32- 2-39 mm ; weight 4-7 g . The Northern Hairy-legged Myotis is morphologically similar to its Neotropical congeners. Fur is short to medium long (dorsal fur 3-5- 6 mm ; ventral fur 3-5 mm ) and woolly. Dorsal fur is unicolored and dark brown to cinnamon-brown, but a few individuals can have slightly darker bases. Ventral hairs are strongly bicolored, with dark brown bases and reddish or yellowish brown tips. Hair color can vary seasonally. Ears are comparatively short, extending forward halfway from eye to nostril. Antitragal notch is barely evident. Membranes are mummy brown; plagiopatagium is broadly attached to foot at bases of toes. Fringe of hairs along trailing edge of uropatagium is absent; dorsal surface of uropatagium is covered with dense hair, extending to middle of tail and exceeding to foot, including fur across tibia onto plagiopatagium. Skull is moderate in size (greatest skull lengths 12:5-14- 1 mm , generally less than 14 mm ); frontal bone is vertically sloped laterally; palatal angle is acute (angle formed between palatal plane and plane of basicranium is greater than 30°); acute nasal angle is noticeable laterally (angle formed between nasals and plane of frontals is less than 45); parietalis inclined forward; occipital region is comparatively flattened posteriorly; and sagittal and lambdoidal crests are present, ranging from low to high. P? is displaced to lingualside with respect to tooth row. Chromosomal complement has 2n = 44 and FN = 50, with large submetacentric X-chromosome and small acrocentric Y-chromosome.	Various habitats, including scrub-steppe, upland semideciduous forest, oakconifer forest, and upland and lowland rainforests from sea level up to elevations of c. 2500 m .	The Northern Hairy-legged Myotis forages in forested areas and over water, where captures mostly lepidopterans, coleopterans, and dipterans in flight.	In Costa Rica , the Northern Hairy-legged Myotis is seasonal polyestrous, without bimodal birth peaks. Births occur mostly in May-June, but breeding can extend through the wet season. In Mexico, one pregnant female captured had a 15-mm long embryo in April.	Activity of the Northern Hairy-legged Myotis begins at sunset and ends c.1 hour before sunrise. It seems to be cave roosting. In Mexico , calls had steep down sweeps that end in narrowband tail, with most energy at 59-63 kHz; initial frequency was ¢.110 kHz but could be lower (c.80 kHz).	In Costa Rica , a few recaptures at the same site and long-distance recaptures ( 200-1450 m ) suggested that Northern Hairy-legged Myotis have large home ranges. In Mexico, colonies of more than 500 individuals in caves are common. In Costa Rica , a colony of 200 individuals was recorded roosting under a bridge. Colonies also can have less than ten individuals.	Not assessed on The IUCN Red List. The Northern Hairylegged Myotis is widespread and ecological flexible and probably does not face any major threats.	Allen (1914) | Arita & Vargas (1995) | Barquez & Diaz ( 2016g) | Bickham et al. (1986) | Hernandez-Meza et al. (2005) | LaVal (1973b) | LaVal & Fitch (1977) | Mantilla-Meluk & Munoz-Garay (2014) | Moratelli et al. (2013) | Rydell et al. (2002) | Timm et al. (1989) | Wilson (2008b)	https://zenodo.org/record/6398739/files/figure.png	386. Northern Hairy-legged Myotis Myotis pilosatibialis French: Murin a pattes velues / German: Nordliches HaarfiRiges Mausohr / Spanish: Ratonero de patas peludas Taxonomy. Myotis keaysi pilosatibialis LaVal, 1973 , “ 1 km W Talanga, Francisco Morazan [= Morazrin], Honduras , elevation 750 m .” Subgenus Pyzonix; ruber species group (7 species). Myotis pilosatibialis became a distinct species after analysis of morphological data in 2014. Monotypic. Distribution. From EC Mexico S through Central America into Colombia and N Venezuela ; also on Trinidad I. Descriptive notes. Head-body c. 45-51 mm , tail 34-39 mm , ear 12-14 mm , hindfoot 7-9 mm ; forearm 32- 2-39 mm ; weight 4-7 g . The Northern Hairy-legged Myotis is morphologically similar to its Neotropical congeners. Fur is short to medium long (dorsal fur 3-5- 6 mm ; ventral fur 3-5 mm ) and woolly. Dorsal fur is unicolored and dark brown to cinnamon-brown, but a few individuals can have slightly darker bases. Ventral hairs are strongly bicolored, with dark brown bases and reddish or yellowish brown tips. Hair color can vary seasonally. Ears are comparatively short, extending forward halfway from eye to nostril. Antitragal notch is barely evident. Membranes are mummy brown; plagiopatagium is broadly attached to foot at bases of toes. Fringe of hairs along trailing edge of uropatagium is absent; dorsal surface of uropatagium is covered with dense hair, extending to middle of tail and exceeding to foot, including fur across tibia onto plagiopatagium. Skull is moderate in size (greatest skull lengths 12:5-14- 1 mm , generally less than 14 mm ); frontal bone is vertically sloped laterally; palatal angle is acute (angle formed between palatal plane and plane of basicranium is greater than 30°); acute nasal angle is noticeable laterally (angle formed between nasals and plane of frontals is less than 45); parietalis inclined forward; occipital region is comparatively flattened posteriorly; and sagittal and lambdoidal crests are present, ranging from low to high. P? is displaced to lingualside with respect to tooth row. Chromosomal complement has 2n = 44 and FN = 50, with large submetacentric X-chromosome and small acrocentric Y-chromosome. Habitat. Various habitats, including scrub-steppe, upland semideciduous forest, oakconifer forest, and upland and lowland rainforests from sea level up to elevations of c. 2500 m . Food and Feeding. The Northern Hairy-legged Myotis forages in forested areas and over water, where captures mostly lepidopterans, coleopterans, and dipterans in flight. Breeding. In Costa Rica , the Northern Hairy-legged Myotis is seasonal polyestrous, without bimodal birth peaks. Births occur mostly in May-June, but breeding can extend through the wet season. In Mexico, one pregnant female captured had a 15-mm long embryo in April. Activity patterns. Activity of the Northern Hairy-legged Myotis begins at sunset and ends c.1 hour before sunrise. It seems to be cave roosting. In Mexico , calls had steep down sweeps that end in narrowband tail, with most energy at 59-63 kHz; initial frequency was ¢.110 kHz but could be lower (c.80 kHz). Movements, Home range and Social organization. In Costa Rica , a few recaptures at the same site and long-distance recaptures ( 200-1450 m ) suggested that Northern Hairy-legged Myotis have large home ranges. In Mexico, colonies of more than 500 individuals in caves are common. In Costa Rica , a colony of 200 individuals was recorded roosting under a bridge. Colonies also can have less than ten individuals. Status and Conservation. Not assessed on The IUCN Red List. The Northern Hairylegged Myotis is widespread and ecological flexible and probably does not face any major threats. Bibliography. Allen (1914), Arita & Vargas (1995), Barquez & Diaz ( 2016g ), Bickham et al. (1986), Hernandez-Meza et al. (2005), LaVal (1973b), LaVal & Fitch (1977), Mantilla-Meluk & Munoz-Garay (2014), Moratelli et al. (2013), Rydell et al. (2002), Timm et al. (1989), Wilson (2008b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis pilosatibialis	Myotis	Unassigned-Myotis	pilosatibialis	La Val	1973	0	Bull. Nat. Hist. Mus. Los Angeles Co.	15:24	Hairy-legged Myotis	None.	Honduras, Francisco Moraz&aacute;n, 1 km W Talanga	S Mexico (San Luis Potosi, Tamaulipas) south through Panama, Ecuador, Venezuela and Trinidad	Not listed.	Least Concern under Myotis keaysi 	Distinct from keaysi; see Mantilla-Meluk and Munoz-Garay (2014). May not be monophyletic; see Novaes et al. (2022).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis pilosatibialis	23	Northern Hairy-legged Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	pilosatibialis	LaVal	1973	0						"1 km W Talanga, Francisco Morazán [= Morazrin], Honduras, elevation 750 m."			pilosatibialis LaVal, 1973	split from M. keaysi	Mantilla-Meluk, H., & Munoz-Garay, J. (2014). Biogeography and taxonomic status of Myotis keaysi pilosatibialis LaVal 1973 (Chiroptera: Vespertilionidae). Zootaxa, 3793(1), 60-70.	Mexico|Guatemala|Belize|El Salvador|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Trinidad & Tobago	North America|South America	Neotropic	NA	0	0	0	Myotis_pilosatibialis	0	unmatched	NA	1																																			Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	pilosatibialis	La Val	1973	0	Bull. Nat. Hist. Mus. Los Angeles Co.	15:24	Hairy-legged Myotis	None.	Honduras, Francisco Moraz&aacute;n, 1 km W Talanga	S Mexico (San Luis Potosi, Tamaulipas) south through Panama, Ecuador, Venezuela and Trinidad	Not listed.	Least Concern under Myotis keaysi 	Distinct from keaysi; see Mantilla-Meluk and Munoz-Garay (2014). May not be monophyletic; see Novaes et al. (2022).	Myotis pilosatibialis	1005456	23	Northern Hairy-legged Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	pilosatibialis	LaVal	1973	0						"1 km W Talanga, Francisco Morazán [= Morazrin], Honduras, elevation 750 m."			pilosatibialis LaVal, 1973	split from M. keaysi	Mantilla-Meluk, H., & Munoz-Garay, J. (2014). Biogeography and taxonomic status of Myotis keaysi pilosatibialis LaVal 1973 (Chiroptera: Vespertilionidae). Zootaxa, 3793(1), 60-70.				Mexico|Guatemala|Belize|El Salvador|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Trinidad & Tobago	North America|South America	Neotropic	NA	0	0	0	Myotis_pilosatibialis	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_pilosatibialis	1005456	23	Northern Hairy-legged Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	pilosatibialis	LaVal	0	Myotis keaysi pilosatibialis	LaVal, R.K. 1973-02-14. A revision of the Neotropical bats of the genus _Myotis_. Natural History Museum of Los Angeles County, Science Bulletin 15:1-54.		LACM:Mamm:36879	holotype	http://portal.vertnet.org/o/lacm/mammals?id=86f3e136-7971-44c1-b1e3-e07192d0ee77	"1 km W Talanga, Francisco Morazán [= Morazrin], Honduras, elevation 750 m."	14.4	-87.0833	split from M. keaysi	Mantilla-Meluk, H., & Munoz-Garay, J. (2014). Biogeography and taxonomic status of Myotis keaysi pilosatibialis LaVal 1973 (Chiroptera: Vespertilionidae). Zootaxa, 3793(1), 60-70.				Mexico|Guatemala|Belize|El Salvador|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Trinidad and Tobago	North America|South America	Neotropic	NE	0	0	0	Myotis_pilosatibialis	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	pilosatibialis	La Val	1973	0	Bull. Nat. Hist. Mus. Los Angeles Co.	15:24	Hairy-legged Myotis	None.	Honduras, Francisco Moraz&aacute;n, 1 km W Talanga	S Mexico (San Luis Potosi, Tamaulipas) south through Panama, Ecuador, Venezuela and Trinidad	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	Not Evaluated	Distinct from keaysi; see Mantilla-Meluk and Munoz-Garay (2014). May not be monophyletic; see Novaes et al. (2022).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis pilosatibialis; Myotis pilosatibialis; Myotis pilosatibialis; Myotis pilosatibialis; pilosatibialis; Murin a pattes velues; Nordliches Haarfi Riges Mausohr; Ratonero de patas peludas; Northern Hairy-legged Myotis; Hairy-legged Myotis; M. pilosatibialis