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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L936	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus	Myotis oxyotus		[MSW2] Subgenus Leuconoe. Revised by LaVal (1973fl).; [MSW3] Revised by LaVal (1973a). Subspecies allocation of populations from coastal Peru is uncertain. Apparently closely related to nigricans and levis; see Ruedi and Mayer (2001).; [HMW] Vespertilio oxyotus Peters, 1866 , “ Chimborazo , in einer Hohe von 9 bis 10,000 Fuss,” Chimborazo , Ecuador . Base on neotype selection, restricted by R. K. LaVal in 1973 to “Gruta Rumichaca, 2 mi E La Paz, Carchi , Ecuador , elevation 2600 m .” Subgenus Pizonyx; albescens species group. Myotis oxyotus was considered a subspecies of M. chiloensis , but studies based on morphology and molecular data indicate clear distinction between the two. Studies of molecular data indicated that subspecies gardneri might be a distinct species. Two subspecies recognized.; [batnames2022] Revised by LaVal (1973 a ). Subspecies allocation of populations from coastal Peru is uncertain. Apparently closely related to nigricans <u> </u>and<u> </u> levis ; see Ruedi and Mayer (2001).; [IUCN] Woodman (1993) notes that many mammalian generic names ending in -otis use the wrong gender for specific names. If this advice is to be followed, the species name should be M. oxyota . The Chiroptera SG advise keeping the names as they are for now while a decision is still to be made on these.; [batnames2023] Revised by LaVal (1973 a ). Subspecies allocation of populations from coastal Peru is uncertain. Apparently closely related to nigricans <u> </u>and<u> </u> levis ; see Ruedi and Mayer (2001).; [batnames2025_1.7] Revised by LaVal (1973a). Subspecies allocation of populations from coastal Peru is uncertain. Apparently closely related to nigricans<u> </u>and<u> </u>levis; see Ruedi and Mayer (2001).						gardneri, thomasi.	gardneri, oxyotus	oxyotus, gardneri	thomasi	oxyotus, gardneri		oxyotus, gardneri	oxyotus - thomasi	oxyotus, thomasi, gardneri	Woodman (1993) notes that many mammalian generic names ending in -otis use the wrong gender for specific names. If this advice is to be followed, the species name should be M. oxyota . The Chiroptera SG advise keeping the names as they are for now while a decision is still to be made on these.	oxyotus, gardneri	oxyotus - thomasi	oxyotus, thomasi, gardneri 	oxyotus, thomasi, gardneri, oxyotus	gardneri, oxyotus	oxyotus - thomasi	oxyotus (W. C. H. Peters, 1866)|thomasi Cabrera, 1901|gardneri LaVal, 1973|oxyotus Woodman, 1993 [unjustified emendation]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Costa Rica – Peru, N Bolivia	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis oxyotus	Ecuador, Mount Chimborazo, between 2743 and 3048 m.	Peters	1867	Monatsb. Preuss. Akad. Wiss. Berlin, p. 19.	Distribution: Known from Costa Rica and western Panama and in South America from Venezuela to Bolivia, mostly in mountain and Pacific coastal areas.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Montane myotis	Costa Rica – Peru, N Bolivia	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1867	Monatsb. K. Preuss. Akad. Wiss. Berlin, 1867:19.	Subgenus Leuconoe. Revised by LaVal (1973fl).	Venezuela to Bolivia; Panama; Costa Rica.	Ecuador, Mount Chimborazo, between 2,743 and 3,048 m.		PETERS	1867	Size fairly small (forearm length, 36-44 mm; condylobasal length, 13-15 mm). Margin of plagiopatagium attached to side of foot. Sagittal crest of braincase absent or poorly developed. Rostrum of medium width or fairly slender.	Distribution: Known from Costa Rica and western Panama and in South America from Venezuela to Bolivia, mostly in mountain and Pacific coastal areas.	Two subspecies are recognized:	M. o. gardneri (Costa Rica, Panama), M. o. oxyotus (remainder of range except for coastal Peru, where subspecific allocation is uncertain).	107	species	M. oxyotus	PETERS	1867	Leuconoe	subgenus	Myotis oxyotus				Size fairly small (forearm length, 36-44 mm; condylobasal length, 13-15 mm). Margin of plagiopatagium attached to side of foot. Sagittal crest of braincase absent or poorly developed. Rostrum of medium width or fairly slender.	Two subspecies are recognized:		63. M. oxyotus (PETERS 1867) [levis group].	63	_M. o. gardneri_ LaVal, 1973; _M. o. oxyotus_ (Peters, 1866) (synonyms: _thomasi_ Cabrera, 1901)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis oxyotus	Myotis		oxyotus	Peters	y	1867		Monatsb. K. Preuss. Akad. Wiss. Berlin	1867		19		Montane Myotis	Ecuador, Mount Chimborazo, between 2,743 and 3,048 m.	Venezuela to Bolivia; Panama; Costa Rica.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	thomasi Cabrera, 1901; gardneri LaVal, 1973.	Revised by LaVal (1973a). Subspecies allocation of populations from coastal Peru is uncertain. Apparently closely related to nigricans and levis; see Ruedi and Mayer (2001).	4C3D87E8FF4F6AF0FF449DA01612BDAE	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	942	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF4F6AF0FF449DA01612BDAE.xml	Myotis oxyotus	Vespertilionidae	Myotis	oxyotus		1866	Murin a oreilles effilées @fr | Gebirgsmausohr @de | Ratonero montano @es	Vespertilio oxyotus Peters, 1866 , “ Chimborazo , in einer Hohe von 9 bis 10,000 Fuss,” Chimborazo , Ecuador . Base on neotype selection, restricted by R. K. LaVal in 1973 to “Gruta Rumichaca, 2 mi E La Paz, Carchi , Ecuador , elevation 2600 m .” Subgenus Pizonyx; albescens species group. Myotis oxyotus was considered a subspecies of M. chiloensis , but studies based on morphology and molecular data indicate clear distinction between the two. Studies of molecular data indicated that subspecies gardneri might be a distinct species. Two subspecies recognized.	M.o.oxyotusPeters,1866—fromColombiaandVenezuelaStoEcuador,Peru,BoliviaandNArgentina. M. o. gardneri LaVal, 1973 — Cordillera Central of Costa Rica and Cordillera de Talamanca of Costa Rica and Panama .	Head-body c¢.40 53 mm , tail 41-46 mm , ear 12-19 mm , hindfoot 6-9 mm , forearm 36-9-43- 4 mm ; weight 4-7 g . Fur is long (dorsal fur 7-10 mm ; ventral fur 5-7 mm ) and woolly. Dorsal hairs are mummy brown, with bases sometimes darker than tips, but dorsal fur generally appears unicolored. Ventral hairs are strongly bicolored, with blackish bases and pale pinkish buff to Dresden brown tips. Ears are medium-sized, extending forward halfway from eye to nostril. Antitragal notch is barely evident. Membranes are mummy brown orslightly darker; plagiopatagium 1s broadly attached to foot at base of toes; uropatagium lacks fringe of hairs. Skull is large (greatest length of skull 14-1-16- 1 mm ). Frontals are steeply sloping; sagittal crest is usually absent, but when present,it is always very low; lambdoidal crests are low and weakly developed; parietals are inclined forward; and occipital region is always rounded and projects behind posterior surfaces of occipital condyles; and P? is generally aligned in tooth row and visible in labial view. Bacula are 0-64- 0-82 mm long, 0-22- 0-35 mm deep, and 0-38- 0-46 mm wide, and specimens from South America are substantially larger. Chromosomal complement has 2n = 44 and FN = 50. X-chromosome is large submetacentric, and Y-chromosome is small acrocentric.	Highland forests, including wet montane forests, at elevations of 1500-3450 m (rarely below 1000 m ).	The Montane Myotis is insectivorous and forages within forested areas and over water.	No information.	No information.	No information.	Classified as Least Concern on The IUCN Red List. The Montane Myotis is widespread and tolerates habitat alterations by humans.	Chaverri et al. (2016) | LaVal (1973b) | Miller & Allen (1928) | Moratelli et al. (2013) | Solari ( 2018m) | Trouessart (1904) | Urquizo et al. (2017) | Wilson (2008b)	https://zenodo.org/record/6398785/files/figure.png	406. Montane Myotis Myotis oxyotus French: Murin a oreilles effilées / German: Gebirgsmausohr / Spanish: Ratonero montano Taxonomy. Vespertilio oxyotus Peters, 1866 , “ Chimborazo , in einer Hohe von 9 bis 10,000 Fuss,” Chimborazo , Ecuador . Base on neotype selection, restricted by R. K. LaVal in 1973 to “Gruta Rumichaca, 2 mi E La Paz, Carchi , Ecuador , elevation 2600 m .” Subgenus Pizonyx; albescens species group. Myotis oxyotus was considered a subspecies of M. chiloensis , but studies based on morphology and molecular data indicate clear distinction between the two. Studies of molecular data indicated that subspecies gardneri might be a distinct species. Two subspecies recognized. Subspecies and Distribution. M.o.oxyotusPeters,1866—fromColombiaandVenezuelaStoEcuador,Peru,BoliviaandNArgentina. M. o. gardneri LaVal, 1973 — Cordillera Central of Costa Rica and Cordillera de Talamanca of Costa Rica and Panama . Descriptive notes. Head-body c¢.40 53 mm , tail 41-46 mm , ear 12-19 mm , hindfoot 6-9 mm , forearm 36-9-43- 4 mm ; weight 4-7 g . Fur is long (dorsal fur 7-10 mm ; ventral fur 5-7 mm ) and woolly. Dorsal hairs are mummy brown, with bases sometimes darker than tips, but dorsal fur generally appears unicolored. Ventral hairs are strongly bicolored, with blackish bases and pale pinkish buff to Dresden brown tips. Ears are medium-sized, extending forward halfway from eye to nostril. Antitragal notch is barely evident. Membranes are mummy brown orslightly darker; plagiopatagium 1s broadly attached to foot at base of toes; uropatagium lacks fringe of hairs. Skull is large (greatest length of skull 14-1-16- 1 mm ). Frontals are steeply sloping; sagittal crest is usually absent, but when present,it is always very low; lambdoidal crests are low and weakly developed; parietals are inclined forward; and occipital region is always rounded and projects behind posterior surfaces of occipital condyles; and P? is generally aligned in tooth row and visible in labial view. Bacula are 0-64- 0-82 mm long, 0-22- 0-35 mm deep, and 0-38- 0-46 mm wide, and specimens from South America are substantially larger. Chromosomal complement has 2n = 44 and FN = 50. X-chromosome is large submetacentric, and Y-chromosome is small acrocentric. Habitat. Highland forests, including wet montane forests, at elevations of 1500-3450 m (rarely below 1000 m ). Food and Feeding. The Montane Myotis is insectivorous and forages within forested areas and over water. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Montane Myotis is widespread and tolerates habitat alterations by humans. Bibliography. Chaverri et al. (2016), LaVal (1973b), Miller & Allen (1928), Moratelli et al. (2013), Solari ( 2018m ), Trouessart (1904), Urquizo et al. (2017), Wilson (2008b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis oxyotus	Myotis	Unassigned-Myotis	oxyotus	Peters	1867	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	######	Montane Myotis	 thomasi Cabrera, 1901; <b> gardneri </b> LaVal, 1973.	Ecuador, Mount Chimborazo, between 2,743 and 3,048 m.	Venezuela to Bolivia; Panama; Costa Rica.	Not listed.	Least Concern	Revised by LaVal (1973 a ). Subspecies allocation of populations from coastal Peru is uncertain. Apparently closely related to nigricans <u> </u>and<u> </u> levis ; see Ruedi and Mayer (2001).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis oxyotus	23	Montane Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	oxyotus	W. Peters	1866	1						"Chimborazo, in einer Höhe von 9 bis 10,000 Fuss," Chimborazo, Ecuador. Base on neotype selection, restricted by R. K. LaVal in 1973 to "Gruta Rumichaca, 2 mi E La Paz, Carchi, Ecuador, elevation 2600 m."			oxyotus (W. Peters, 1866)|thomasi Cabrera, 1901|gardneri LaVal, 1973	NA	NA	Costa Rica|Panama|Colombia|Venezuela|Ecuador|Peru|Bolivia|Argentina	North America|South America	Neotropic	LC	0	0	0	Myotis_oxyotus	0	sciname match	Myotis_oxyotus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14187	Myotis oxyotus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	oxyotus	(Peters, 1867)	Woodman (1993) notes that many mammalian generic names ending in -otis use the wrong gender for specific names. If this advice is to be followed, the species name should be M. oxyota . The Chiroptera SG advise keeping the names as they are for now while a decision is still to be made on these.	20000000	Myotis oxyotus	Least Concern		2018	2018-03-05 00:00:00 UTC	3.1	English	Although restricted to montane and highland areas, this species is listed as Least Concern in view of its wide distribution, tolerance of human habitat conversion and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	The natural history of this bat is poorly known. It is found in evergreen highland forest and forest edges. Found in montane forests and strongly associated with highland habitats (Wilson 2008). It apparently prefers forested areas in these cooler montane areas, where day roosts are known to include hollow trees. They have been mistnetted over roads and in clearings in the forest, as well as in more open areas adjacent to the ;forest (Linares 1998). Pregnant females have been recorded in February and July (Gardner ;et al.  ;1970, Reid 2009). ;It seems to tolerate some range of human disturbance.	There are no known major threats throughout the species' range. Change in land use are not frequent or extreme at the higher parts of the Andes but, when present could affect some specific populations over part of its geographic distribution.	It is uncommon to fairly common (Eisenberg and Redford 1999), with great variation in abundance among localities.	Unknown	This species is known from the Andes of Venezuela to Bolivia, plus the mountains of Panama and Costa Rica (Simmons 2005, Wilson 2008). In Bolivia it is found up to 3,800 m, in Venezuela between 1,800-3,100 m, in Colombia between 1,000-2,880 m, and in Peru between 2,600-3,900 m. It seems to be restricted to Andean Montane formations, especially to forested areas.		Terrestrial	This species occurs in protected areas through its geographic distribution.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	oxyotus	Peters	1867	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	########	Montane Myotis	 thomasi Cabrera, 1901; <b> gardneri </b> LaVal, 1973.	Ecuador, Mount Chimborazo, between 2,743 and 3,048 m.	Venezuela to Bolivia; Panama; Costa Rica.	Not listed.	Least Concern	Revised by LaVal (1973 a ). Subspecies allocation of populations from coastal Peru is uncertain. Apparently closely related to nigricans <u> </u>and<u> </u> levis ; see Ruedi and Mayer (2001).	Myotis oxyotus	1005450	23	Montane Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	oxyotus	W. Peters	1866	1						"Chimborazo, in einer Höhe von 9 bis 10,000 Fuss," Chimborazo, Ecuador. Base on neotype selection, restricted by R. K. LaVal in 1973 to "Gruta Rumichaca, 2 mi E La Paz, Carchi, Ecuador, elevation 2600 m."			oxyotus (W. Peters, 1866)|thomasi Cabrera, 1901|gardneri LaVal, 1973	NA	NA				Costa Rica|Panama|Colombia|Venezuela|Ecuador|Peru|Bolivia|Argentina	North America|South America	Neotropic	LC	0	0	0	Myotis_oxyotus	0	sciname match	Myotis_oxyotus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_oxyotus	1005450	23	Montane Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	oxyotus	W. C. H. Peters	1	Vespertilio oxyotus	Peters, W.C.H. 1866. Ãœber einige neue oder weniger bekannte Flederthiere. Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin 1866:16-25.	https://www.biodiversitylibrary.org/page/39052569	LACM:Mamm:36878	neotype	http://portal.vertnet.org/o/lacm/mammals?id=e80df7e9-bf5e-4dee-93a6-3c8ecfe663fb	"Chimborazo, in einer Höhe von 9 bis 10,000 Fuss," Chimborazo, Ecuador. Base on neotype selection, restricted by R. K. LaVal in 1973 to "Gruta Rumichaca, 2 mi E La Paz, Carchi, Ecuador, elevation 2600 m."	0.483333	-77.8333	NA	NA				Costa Rica|Panama|Colombia|Venezuela|Ecuador|Peru|Bolivia|Argentina	North America|South America	Neotropic	LC	0	0	0	Myotis_oxyotus	0	sciname match	Myotis_oxyotus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	oxyotus	Peters	1867	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	########	Montane Myotis	thomasi Cabrera, 1901; gardneri LaVal, 1973.	Ecuador, Mount Chimborazo, between 2,743 and 3,048 m.	Venezuela to Bolivia; Panama; Costa Rica.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14187/22067211/' target='_blank'>Least Concern</a>	Revised by LaVal (1973a). Subspecies allocation of populations from coastal Peru is uncertain. Apparently closely related to nigricans<u> </u>and<u> </u>levis; see Ruedi and Mayer (2001).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis oxyotus; Myotis oxyotus; Myotis oxyotus; Myotis oxyotus; Myotis oxyotus; Myotis oxyotus; oxyotus; gardneri; thomasi; oxyotus; gardneri; gardneri; thomasi; oxyotus; thomasi; gardneri; Murin a oreilles effilées; Gebirgsmausohr; Ratonero montano; Montane Myotis; Montane Myotis; Montane Myotis; M. oxyotus