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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L932	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Myotis mystacinus [synonym of]	Myotis mystacinus nipalensis	Myotis nipalensis	Myotis davidii?	Myotis nipalensis	Myotis nipalensis	Myotis nipalensis	Myotis nipalensis	Myotis nipalensis	Myotis nipalensis	Myotis nipalensis	Myotis nipalensis		[MSW3] Often included in mystacinus, but see Benda and Tsytsulina (2000).; [batnames2022] Formerly included in davidii but distinct; see Ruedi et al. (2021). Likely includes both annatessae and ancricola , but types of these species have not been compared with nipalensis and we have retained them as valid species pending this additional research. Specimens identified as mystacinus from the Indian subcontinent (Bates and Harrison, 1997) and from Myanmar (Bates et al., 2005) have been assigned to this species by Ruedi et al. (2021). Note that the definition of this species by Srinivasulu and Srinivasulu (2019) is very different from that of Ruedi et al. (2021), which we follow here. Consequently, we do not include transcaspicus as a synonym.; [MDD2022] has been included under M. davidii; may include under M. ancricola and M. annatessae based on molecular data, but this remains to be fully tested; [IUCN] Formerly included in Myotis mystacinus (see Benda and Tsytsulina 2000) Myotis nipalensis possibly represents a complex of several similar species. Further studies are needed to clarify the taxonomic status of populations currently allocated to this species (K. Tsytsulina pers. comm. 2008).; [batnames2023] Formerly included in davidii but distinct; see Ruedi et al. (2021). Likely includes both annatessae and ancricola , but types of these species have not been compared with nipalensis and we have retained them as valid species pending this additional research. Specimens identified as mystacinus from the Indian subcontinent (Bates and Harrison, 1997) and from Myanmar (Bates et al., 2005) have been assigned to this species by Ruedi et al. (2021). Note that the definition of this species by Srinivasulu and Srinivasulu (2019) is very different from that of Ruedi et al. (2021), which we follow here. Consequently, we do not include transcaspicus as a synonym.; [MDD2023] has been included under M. davidii; may include under M. ancricola and M. annatessae based on molecular data, but this remains to be fully tested; [MDD2025_2.0] has been included under M. davidii; may include under M. ancricola and M. annatessae based on molecular data, but this remains to be fully tested; [batnames2025_1.7] Formerly included in davidii but distinct; see Ruedi et al. (2021). Likely includes both annatessae and ancricola, but types of these species have not been compared with nipalensis and we have retained them as valid species pending this additional research. Specimens identified as mystacinus from the Indian subcontinent (Bates and Harrison, 1997) and from Myanmar (Bates et al., 2005) have been assigned to this species by Ruedi et al. (2021). Note that the definition of this species by Srinivasulu and Srinivasulu (2019) is very different from that of Ruedi et al. (2021), which we follow here. Consequently, we do not include transcaspicus as a synonym.; [MDD2025_2.2] has been included under M. davidii; may include under M. ancricola and M. annatessae based on molecular data, but this remains to be fully tested								nipalensis, przewalskii, transcaspicus	kukunoriensis, meinertzhageni, pallidiventris; przewalskii - mongolicus; transcaspicus - pamirensis, sogdianus					pallidiventris, nipalensis, meinertzhageni	Formerly included in Myotis mystacinus (see Benda and Tsytsulina 2000) Myotis nipalensis possibly represents a complex of several similar species. Further studies are needed to clarify the taxonomic status of populations currently allocated to this species (K. Tsytsulina pers. comm. 2008).	nipalensis	nipalensis - meinertzhageni, pallidiventris	pallidiventris, nipalensis, meinertzhageni 	pallidiventris, nipalensis, meinertzhageni, kukunorensis, kukunoriensis	nipalensis	nipalensis - meinertzhageni, pallidiventris	pallidiventris (B. H. Hodgson, 1844) [nomen nudum]|nipalensis (Dobson, 1871)|meinertzhageni O. Thomas, 1926|kukunorensis Bobrinski, 1929|kukunoriensis Ellerman & Morrison-Scott, 1951 [incorrect subsequent spelling]						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis nipalensis	Myotis		nipalensis	Dobson		1871		Proc. Asiat. Soc. Bengal	1871		214		Nepalese Myotis	Nepal, Katmandu.	Iran, Turkey, and Uzbekistan to Nepal, Mongolia, Tibet and NW China, Siberia.	IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001).	kukunoriensis Bobrinskii, 1929; meinertzhageni Thomas, 1926; pallidiventris Hodgson, 1844 [nomen nudum]; przewalskii Bobrinski, 1926; mongolicus Kruskop and Borissenko, 1996; transcaspicus Ognev and Heptner, 1928; pamirensis Kuzyakin, 1935; sogdianus Kuzyakin, 1934.	Often included in mystacinus, but see Benda and Tsytsulina (2000).								synonym of Myotis davidii?					1871														Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis nipalensis	Myotis	Unassigned-Myotis	nipalensis	Dobson	1871	1	Proc. Asiat. Soc. Bengal	1874:34:00	Nepal Myotis	Yes	Nepal, Kathmandu	India, Nepal, and Myanmar	Not listed.	Least Concern	Formerly included in davidii but distinct; see Ruedi et al. (2021). Likely includes both annatessae and ancricola , but types of these species have not been compared with nipalensis and we have retained them as valid species pending this additional research. Specimens identified as mystacinus from the Indian subcontinent (Bates and Harrison, 1997) and from Myanmar (Bates et al., 2005) have been assigned to this species by Ruedi et al. (2021). Note that the definition of this species by Srinivasulu and Srinivasulu (2019) is very different from that of Ruedi et al. (2021), which we follow here. Consequently, we do not include transcaspicus as a synonym.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2031	Myotis nipalensis	23	Nepal Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	nipalensis	Dobson	1871	1	Vespertilio_nipalensis	Dobson, G. E. (1871). Notes on nine new species of Indian and Indo-Chinese Vespertilionidae, with remarks on the synonymy and classification of some other species of the same family. Proceedings of the Asiatic Society of Bengal, 1871, 214.	https://www.biodiversitylibrary.org/item/44923#page/260/mode/1up	ZSI 15573		"Kathmandu, Nepal."			pallidiventris (Hodgson, 1844) [nomen nudum]|nipalensis (Dobson, 1871)|meinertzhageni O. Thomas, 1926	has been included under M. davidii; may include under M. ancricola and M. annatessae based on molecular data, but this remains to be fully tested	Mayer, F., Dietz, C., & Kiefer, A. (2007). Molecular species identification boosts bat diversity. Frontiers in zoology, 4(1), 4.|Dietz, C., Gazaryan, A., Papov, G., Dundarova, H., & Mayer, F. (2016). Myotis hajastanicus is a local vicariant of a widespread species rather than a critically endangered endemic of the Sevan lake basin (Armenia). Mammalian Biology, 81(5), 518-522.|Benda, P., Abi Said, M. R., Jaoude, I. B., Karanouh, R., Lučan, R. K., Sadek, R., ... & Horáček, I. (2016). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 13. Review of distribution and ectoparasites of bats in Lebanon. Acta Societatis Zoologicae Bohemicae, 80, 207-316.|Ruedi, M., Saikia, U., Thabah, A., Görföl, T., Thapa, S., & Csorba, G. (2021). Molecular and morphological revision of small Myotinae from the Himalayas shed new light on the poorly known genus Submyotodon (Chiroptera: Vespertilionidae). Mammalian Biology.	India|Nepal|Bhutan|Myanmar	Asia	Palearctic|Indomalaya	LC	0	0	0	NA	1	sciname match	Myotis_nipalensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136495	Myotis nipalensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	nipalensis	(Dobson, 1871)	Formerly included in Myotis mystacinus (see Benda and Tsytsulina 2000) Myotis nipalensis possibly represents a complex of several similar species. Further studies are needed to clarify the taxonomic status of populations currently allocated to this species (K. Tsytsulina pers. comm. 2008).	20000000	Myotis nipalensis	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species is found in a wide variety of often arid or mountainous habitats, including forest, shrubland, grassland areas and desert. It has also been recorded from rural garden and urban areas in South Asia. Roosts in cracks in in buildings, rocks, caves and old mines (Smith and Xie 2008, Katerina Tsytsulina pers. comm. 8 May 2008). In Central Asia, this species has been recorded from desert lowlands and fertile mountains and from a wide variety of vegetation types (Benda et al. 2011). Animals usually begin feeding after dusk, hunting flying insects low above ground. The flight is fast and manoeuvring. It is most probably non-migratory, but populations can relocate to different roosting sites if they are disturbed. Winters in underground shelters. The species reproduces once a year, giving birth to a single young.	This is a widely distributed, generally common species. In South Asia, the abundance, population size and trends for this species are not known (S. Thapa pers. comm. 2017).	This is a widely distributed, generally common species. In South Asia, the abundance, population size and trends for this species are not known (S. Thapa pers. comm. 2017).	Unknown	Thedistribution range of this species is clouded by ongoing taxonomic problems, however, it has presently been widely recorded from the Caucasus in the west of its range, through much of Central Asia and western and central China, being found as far south as northern India and Nepal. In the Caucasus, it is likely present in northeastern Turkey, Georgia, Armenia, Azerbaijan and the Caspian region of northern Iran. In Central Asia, the species appears to be distributed in Turkmenistan, Kazakhstan, Uzbekistan, Tajikistan and Kyrgyzstan. In China, it has been recorded from Qinghai, Gansu and Xinjiang (Smith and Xie 2008), and is possible present in Xizang. In South Asia, this species has been recorded from India (Himachal Pradesh, Jammu and Kashmir, Meghalaya and West Bengal) and Nepal (Central Nepal) in South Asia (Srinivasulu et al . in press). Elevation details are not known.		Terrestrial	In view of the species wide range, it seems probable that it is present in many protected areas. There is a need for additional taxonomic research into this probable species complex.	Indomalayan|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	nipalensis	Dobson	1871	1	Proc. Asiat. Soc. Bengal	1874:34:00	Nepal Myotis	Yes	Nepal, Kathmandu	India, Nepal, and Myanmar	Not listed.	Least Concern	Formerly included in davidii but distinct; see Ruedi et al. (2021). Likely includes both annatessae and ancricola , but types of these species have not been compared with nipalensis and we have retained them as valid species pending this additional research. Specimens identified as mystacinus from the Indian subcontinent (Bates and Harrison, 1997) and from Myanmar (Bates et al., 2005) have been assigned to this species by Ruedi et al. (2021). Note that the definition of this species by Srinivasulu and Srinivasulu (2019) is very different from that of Ruedi et al. (2021), which we follow here. Consequently, we do not include transcaspicus as a synonym.	Myotis nipalensis	1006524	23	Nepal Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	nipalensis	Dobson	1871	1	Vespertilio_nipalensis	Dobson, G. E. (1871). Notes on nine new species of Indian and Indo-Chinese Vespertilionidae, with remarks on the synonymy and classification of some other species of the same family. Proceedings of the Asiatic Society of Bengal, 1871, 214.	https://www.biodiversitylibrary.org/item/44923#page/260/mode/1up	ZSI 15573		"Kathmandu, Nepal."			pallidiventris (Hodgson, 1844) [nomen nudum]|nipalensis (Dobson, 1871)|meinertzhageni O. Thomas, 1926	has been included under M. davidii; may include under M. ancricola and M. annatessae based on molecular data, but this remains to be fully tested	Mayer, F., Dietz, C., & Kiefer, A. (2007). Molecular species identification boosts bat diversity. Frontiers in zoology, 4(1), 4.|Dietz, C., Gazaryan, A., Papov, G., Dundarova, H., & Mayer, F. (2016). Myotis hajastanicus is a local vicariant of a widespread species rather than a critically endangered endemic of the Sevan lake basin (Armenia). Mammalian Biology, 81(5), 518-522.|Benda, P., Abi Said, M. R., Jaoude, I. B., Karanouh, R., Lučan, R. K., Sadek, R., ... & Horáček, I. (2016). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 13. Review of distribution and ectoparasites of bats in Lebanon. Acta Societatis Zoologicae Bohemicae, 80, 207-316.|Ruedi, M., Saikia, U., Thabah, A., Görföl, T., Thapa, S., & Csorba, G. (2021). Molecular and morphological revision of small Myotinae from the Himalayas shed new light on the poorly known genus Submyotodon (Chiroptera: Vespertilionidae). Mammalian Biology.				India|Nepal|Bhutan|Myanmar	Asia	Palearctic|Indomalaya	LC	0	0	0	NA	1	sciname match	Myotis_nipalensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_nipalensis	1006524	23	Nepal Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	nipalensis	Dobson	1	Vespertilio Nipalensis	Dobson, G.E. 1871. Notes on nine new species of Indian and Indo-Chinese Vespertilionidae, with remarks on the synonymy and classification of some other species of the same family. Proceedings of the Asiatic Society of Bengal 1871:210-215.	https://www.biodiversitylibrary.org/page/12393596	ZSI 155573	holotype		"Kathmandu, Nepal."			has been included under M. davidii; may include under M. ancricola and M. annatessae based on molecular data, but this remains to be fully tested	Mayer, F., Dietz, C., & Kiefer, A. (2007). Molecular species identification boosts bat diversity. Frontiers in zoology, 4(1), 4.|Dietz, C., Gazaryan, A., Papov, G., Dundarova, H., & Mayer, F. (2016). Myotis hajastanicus is a local vicariant of a widespread species rather than a critically endangered endemic of the Sevan lake basin (Armenia). Mammalian Biology, 81(5), 518-522.|Benda, P., Abi Said, M. R., Jaoude, I. B., Karanouh, R., Lučan, R. K., Sadek, R., ... & Horáček, I. (2016). Bats (Mammalia: Chiroptera) of the Eastern Mediterranean and Middle East. Part 13. Review of distribution and ectoparasites of bats in Lebanon. Acta Societatis Zoologicae Bohemicae, 80, 207-316.|Ruedi, M., Saikia, U., Thabah, A., Görföl, T., Thapa, S., & Csorba, G. (2021). Molecular and morphological revision of small Myotinae from the Himalayas shed new light on the poorly known genus Submyotodon (Chiroptera: Vespertilionidae). Mammalian Biology.				India|Nepal|Bhutan|Myanmar	Asia	Palearctic|Indomalaya	LC	0	0	0	NA	1	sciname match	Myotis_nipalensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	nipalensis	Dobson	1871	1	Proc. Asiat. Soc. Bengal	1874:34:00	Nepal Myotis	Yes	Nepal, Kathmandu	India, Nepal, and Myanmar	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136495/21976309/' target='_blank'>Least Concern</a>	Formerly included in davidii but distinct; see Ruedi et al. (2021). Likely includes both annatessae and ancricola, but types of these species have not been compared with nipalensis and we have retained them as valid species pending this additional research. Specimens identified as mystacinus from the Indian subcontinent (Bates and Harrison, 1997) and from Myanmar (Bates et al., 2005) have been assigned to this species by Ruedi et al. (2021). Note that the definition of this species by Srinivasulu and Srinivasulu (2019) is very different from that of Ruedi et al. (2021), which we follow here. Consequently, we do not include transcaspicus as a synonym.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis nipalensis; Myotis davidii?; Myotis nipalensis; Myotis nipalensis; Myotis nipalensis; Myotis nipalensis; nipalensis; przewalskii; transcaspicus; kukunoriensis; meinertzhageni; pallidiventris; przewalskii - mongolicus; transcaspicus - pamirensis; sogdianus; pallidiventris; nipalensis; meinertzhageni; Nepal Myotis; Nepalese Myotis; Nepal Myotis; M. nipalensis