http://www.w3.org/ns/prov#wasDerivedFrom	http://purl.org/dc/elements/1.1/format	name_CH1_1980	name_MSW1_1982	name_CH3_1991	name_MSW2_1993	name_Koopman_1994	name_MSW3_2005	name_HMW_2019	name_BatNames_2022	name_MDD_2022	name_IUCN_2022	name_BatNames_2023	name_MDD_2023	name_MDD_2025_2.0	name_batnames_2025_1.7	name_MDD_2025_2.2	column151	taxonomic_notes_concatenated	column171	synonyms_CH1	subspecies__MSW2	synonyms__MSW1	synonyms_CH3	synonyms_MSW2	subspecies_Koopman94_interpreted	subspecies_MSW3_interpreted	synonym_MSW3_interpreted	subspecies_HMW_interpreted	synonym_HMW_interpreted	subspecies_batnames_interpreted	synonym_batnames_interpreted	synonym_MDD_interpreted	synonym_IUCN_interpreted	subspecies_batnames2023_interpreted	synonym_batnames2023_interpreted	synonym_MDD2023_interpreted	synonym_MDD2025_interpreted	subspecies_batnames2025_interpreted	synonyms_batnames2025_interpreted	nominalNames	column391	docOrigin_CH1	commonName_CH1	distribution_CH1	docOrigin_MSW1	column451	typeLocality_MSW1	authority_MSW1	year_MSW1	citation_MSW1	distribution	comment_MSW1	docOrigin_CH3	commonName_CH3	distribution_CH3	docOrigin_MSW2	authority_MSW2	year_MSW2	citation_MSW2	comments_MSW2	distribution_MSW2	typeLocality_MSW2	docOrigin_Koopman94	authority_Koopman94	year_Koopman94	description_Koopman94	distribution_Koopman94	diversity_Koopman94	subspecies_Koopman94	page	rank	name	authority	year	parent	parent_rank	corrected_name	actual_species_count	claimed_species_count	dental_formula	description	diversity	full_subspecies_text	name_line	species_index	subspecies	synonym	text	docOrigin_MSW3	order_MSW3	family_MSW3	subfamily_MSW3	tribe_MSW3	name_MSW3	genus_MSW3	subgenus_MSW3	species_MSW3	authoritySpeciesAuthor_MSW3	(parentheses (1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common Name_batnames	synonyms_batnames	type_locality_batnames	Distribution_batnames	CITES_batnames	IUCN_batnames	comments_batnames	docOrigin_MDD	name_MDD	phylosort_MDD	mainCommonName_MDD	otherCommonNames_MDD	subclass_MDD	infraclass_MDD	magnorder_MDD	superorder_MDD	order_MDD	suborder_MDD	infraorder_MDD	parvorder_MDD	superfamily_MDD	family_MDD	subfamily_MDD	tribe_MDD	genus_MDD	subgenus_MDD	specificEpithet_MDD	authoritySpeciesAuthor_MDD	authoritySpeciesYear_MDD	authorityParentheses_MDD	originalNameCombination_MDD	authoritySpeciesCitation_MDD	authoritySpeciesLink_MDD	holotypeVoucher_MDD	holotypeVoucherURIs_MDD	typeLocality_MDD	typeLocalityLatitude_MDD	typeLocalityLongitude_MDD	nominalNames_MDD	taxonomyNotes_MDD	taxonomyNotesCitation_MDD	countryDistribution_MDD	continentDistribution_MDD	biogeographicRealm_MDD	iucnStatus_MDD	extinct_MDD	domestic_MDD	flagged_MDD	CMW_sciName_MDD	diffSinceCMW_MDD	MSW3_matchtype_MDD	MSW3_sciName_MDD	diffSinceMSW3_MDD	docOrigin_IUCN	internalTaxonId_IUCN	NAME_IUCN	kingdomName_IUCN	phylumName_IUCN	className_IUCN	orderName_IUCN	familyName_IUCN	genusName_IUCN	speciesName_IUCN	authoritySpeciesAuthorYear_IUCN	taxonomicNotes_IUCN	assessmentId_IUCN	scientificName_IUCN	redlistCategory_IUCN	redlistCriteria_IUCN	yearPublished_IUCN	assessmentDate_IUCN	criteriaVersion_IUCN	language_IUCN	rationale_IUCN	habitat_IUCN	threats_IUCN	population_IUCN	populationTrend_IUCN	range_IUCN	useTrade_IUCN	systems_IUCN	conservationActions_IUCN	realm_IUCN	yearLastSeen_IUCN	possiblyExtinct_IUCN	possiblyExtinctInTheWild_IUCN	scopes_IUCN	docOrigin_batnames2023	FAMILY_batnames2023	GENUS_batnames2023	SUBGENUS_batnames2023	SPECIES_batnames2023	authoritySpeciesAuthor_batnames2023	authoritySpeciesYearbatnames2023	PARENTHESES_batnames2023 (1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name MDD2023	id_MDD2023	phylosort_MDD2023	mainCommonName_MDD2023	otherCommonNames_MDD2023	subclass_MDD2023	infraclass_MDD2023	magnorder_MDD2023	superorder_MDD2023	order_MDD2023	suborder_MDD2023	infraorder_MDD2023	parvorder_MDD2023	superfamily_MDD2023	Family_mdd2023	subfamily_MDD2023	tribe_MDD2023	genus_MDD2023	subgenus_MDD2023	specificEpithet_MDD2023	authoritySpeciesAuthor_MDD2023	authoritySpeciesYear_MDD2023	authorityParentheses_MDD2023	originalNameCombination_MDD2023	authoritySpeciesCitation_MDD2023	authoritySpeciesLink_MDD2023	holotypeVoucher_MDD2023	holotypeVoucherURIs_MDD2023	typeLocality_MDD2023	typeLocalityLatitude_MDD2023	typeLocalityLongitude_MDD2023	nominalNames_MDD2023	taxonomyNotes_MDD2023	taxonomyNotesCitation_MDD2023	distributionNotes_MDD2023	distributionNotesCitation_MDD2023	subregionDistribution_MDD2023	countryDistribution_MDD2023	continentDistribution_MDD2023	biogeographicRealm_MDD2023	iucnStatus_MDD2023	extinct_MDD2023	domestic_MDD2023	flagged_MDD2023	CMW_sciName_MDD2023	diffSinceCMW_MDD2023	MSW3_matchtype_MDD2023	MSW3_sciName_MDD2023	diffSinceMSW3_MDD2023	docOrigin_MDD2025	sciName	id	phylosort	mainCommonName	otherCommonNames	subclass	infraclass	magnorder	superorder	order	suborder	infraorder	parvorder	superfamily	family	subfamily	tribe	genus	subgenus	specificEpithet	authoritySpeciesAuthor	authorityParentheses	originalNameCombination	authoritySpeciesCitation	authoritySpeciesLink	typeVoucher	typeKind	typeVoucherURIs	typeLocality	typeLocalityLatitude	typeLocalityLongitude	taxonomyNotes	taxonomyNotesCitation	distributionNotes	distributionNotesCitation	subregionDistribution	countryDistribution	continentDistribution	biogeographicRealm	iucnStatus	extinct	domestic	flagged	CMW_sciName	diffSinceCMW	MSW3_matchtype	MSW3_sciName	diffSinceMSW3	docOrigin_batnames2025	Family	Genus	Subgenus	Species	Author	Date	Parentheses (1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L928	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis nattereri	Myotis nattereri	Myotis nattereri	Myotis nattereri	Myotis nattereri	Myotis nattereri	Myotis nattereri	Myotis nattereri	Myotis nattereri	Myotis nattereri	Myotis nattereri	Myotis nattereri	Myotis nattereri	Myotis nattereri	Myotis nattereri		[MSW2] Subgenus Myotis. Does not include araxenus or bombinus; see Hor4iek and Hanak (1984).; [MSW3] Does not include araxenus or bombinus; see Horácek and Hanák (1984) and Kawai et al. (2003). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000). For discussion of correct spelling see Bogdanowicz and Kock (1998).; [HMW] Vespertilio natterer: Kuhl, 1817 , Hanau, Hessen , Germany . Subgenus Myotis ; myotis species group. The taxonomy of the nattereri species complex is currently very convoluted. Four cryptic species have currently been recognized as distinct ( M. nattereri , M. crypticus, M. escalerai , and M. zenatius) based on morphometric and genetic data, and a further two more species (hoveli and tschuliensis) may be recognizable from the Levant and Caucasus that appearto be related to M. schaubi . Anothercryptic species may also be found on Corsica based on limited genetic data; the populations on other large Mediterranean islandsare also currently uncertain due to the recent taxonomic changes. Myotis natterer and M. crypticus are sister species whereas M. escalerai and M. zenatiusare sister to one another with M. schaubi being related to either clade based on various genetic studies. The names hoveli and tschuliensis are here included as subspecies pending further studies that may prove their distinctiveness. Three subspecies recognized.; [batnames2022] Does not include araxenus or bombinus ; see Horácek and Hanák (1984), Kawai et al. (2003), and Çoraman et al. (2019). Does not include escalerai ; see Ibáñez et al. (2006). Does not include hoveli ; see Çoraman et al. (2019). Does not include tschuliensis ; see Çoraman et al. (2019), Smirnov et al. (2020), Kruskop and Solovyeva (2020), and Uvizl and Benda (2021). A cryptic species may occur in Austria and Northern Italy; see Mayer et al. (2007). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Puechmaille et al. (2012), Salicini et al. (2011), Çoraman et al. (2019), Juste et al. (2019), and Smirnov et al. (2020) for additional discussion of the M. nattereri complex.; [MDD2022] previously included M. escalarae, M. tschuliensis, and M. hoveli; [IUCN] Does not include ;Myotis schaubi ;(Armenia and western Iran), ;M. escalerai, ; M. crypticus, ; M. zenatius, ; M. hoveli, ; M. tschuliensis, ; and the not formally described Myotis sp. C. (endemic subpopulation of Corsica). It also doesn't include ;M. bombinus ; (southeastern Siberia, northeastern China, Korea, Japan).; [batnames2023] Does not include araxenus or bombinus ; see Horácek and Hanák (1984), Kawai et al. (2003), and Çoraman et al. (2019). Does not include escalerai ; see Ibáñez et al. (2006). Does not include hoveli ; see Çoraman et al. (2019). Does not include tschuliensis ; see Çoraman et al. (2019), Smirnov et al. (2020), Kruskop and Solovyeva (2020), and Uvizl and Benda (2021). A cryptic species may occur in Austria and Northern Italy; see Mayer et al. (2007). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Puechmaille et al. (2012), Salicini et al. (2011), Çoraman et al. (2019), Juste et al. (2019), and Smirnov et al. (2020) for additional discussion of the M. nattereri complex.; [MDD2023] previously included M. escalarae, M. tschuliensis, and M. hoveli; [MDD2025_2.0] previously included M. escalarae, M. tschuliensis, and M. hoveli; [batnames2025_1.7] Does not include araxenus or bombinus; see Horácek and Hanák (1984), Kawai et al. (2003), and Çoraman et al. (2019). Does not include escalerai; see Ibáñez et al. (2006). Does not include hoveli; see Çoraman et al. (2019). Does not include tschuliensis; see Çoraman et al. (2019), Smirnov et al. (2020), Kruskop and Solovyeva (2020), and Uvizl and Benda (2021). A cryptic species may occur in Austria and Northern Italy; see Mayer et al. (2007). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Puechmaille et al. (2012), Salicini et al. (2011), Çoraman et al. (2019), Juste et al. (2019), and Smirnov et al. (2020) for additional discussion of the M. nattereri complex.; [MDD2025_2.2] previously included M. escalarae, M. tschuliensis, and M. hoveli						escalerae, spelaeus, tschuliensis, typus.	tschuliensis, nattereri	nattereri, tschuliensis	escalerae, hoveli, spelaeus, typus	nattereri, hoveli, tschuliensis		nattereri, tschuliensis	nattereri - spelaeus; tschuliensis - typus	nattereri, spelaeus, typus	Does not include ;Myotis schaubi ;(Armenia and western Iran), ;M. escalerai, ; M. crypticus, ; M. zenatius, ; M. hoveli, ; M. tschuliensis, ; and the not formally described Myotis sp. C. (endemic subpopulation of Corsica). It also doesn't include ;M. bombinus ; (southeastern Siberia, northeastern China, Korea, Japan).	nattereri	nattereri - spelaeus, typus	nattereri, spelaeus, typus	nattereri, natererea, nattererii, spelaeus, typus	nattereri	nattereri - spelaeus, typus	nattereri (Kuhl, 1817)|natererea (S.D.W., 1836) [unjustified emendation]|nattererii (Loche, 1858) [incorrect subsequent spelling]|spelaeus (C. Koch, 1863)|typus (C. Koch, 1863)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Natterer's bat	Morocco, W Europe – SE Siberia, Japan	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis nattereri	Germany, Hessen, Hanau.	Kuhl	1818	Ann. Wetterau Ges. Naturk., 4(1):33.	Distribution: Ranging from Ireland, Portugal, and Morocco to the Urals, Israel and Turkmenia.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Natterer's bat	W Europe – Urals – Israel; N Africa	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Kuhl	1817	Die Deutschen Fledermause. Hanau, p. 14, 33.	Subgenus Myotis. Does not include araxenus or bombinus; see Hor4iek and Hanak (1984).	Europe (except Scandinavia); NW Africa; Turkey; Israel; Iraq; Crimea and Caucasus to Turkmenistan.	Germany, Hessen, Hanau.		KUHL	1817	Size medium (forearm length, 38-42 mm; condylobasal length, 13-16mm). Ear medium in length. Braincase relatively high but rostrum fairly long and narrow. Nasal emargination medium. Middle upper premolar in toothrow. Margin of uropatagium with a dense fringe of short hairs.	Distribution: Ranging from Ireland, Portugal, and Morocco to the Urals, Israel and Turkmenia.	Two subspecies are here recognized:	M. n. tschuliensis (Transcaucasia, Iraq, Turkmenia), M. n. nattereri (remainder or range).	101	species	M. nattereri	KUHL	1817	Myotis	subgenus	Myotis nattereri				Size medium (forearm length, 38-42 mm; condylobasal length, 13-16mm). Ear medium in length. Braincase relatively high but rostrum fairly long and narrow. Nasal emargination medium. Middle upper premolar in toothrow. Margin of uropatagium with a dense fringe of short hairs.	Two subspecies are here recognized:		8. M. nattereri (KUHL 1817) [nattereri group],	8	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis nattereri	Myotis		nattereri	Kuhl	y	1817		Die Deutschen Fledermäuse. Hanau			14, 33		Natterer's Myotis	Germany, Hessen, Hanau.	Ireland, Great Britain, Europe (except N Scandinavia), Morocco, N Algeria, Turkey, Israel, Jordan, Lebanon, Iraq, Iran, Bulgaria, Crimea and Caucasus to Turkmenistan.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	escalerae Cabrera, 1904; hoveli Harrison, 1964; spelaeus Koch, 1865; typus Koch, 1865; tschuliensis Kuzyakin, 1935.	Does not include araxenus or bombinus; see Horácek and Hanák (1984) and Kawai et al. (2003). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000). For discussion of correct spelling see Bogdanowicz and Kock (1998).	4C3D87E8FF226A9DFF7C97561941B82E	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	979	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF226A9DFF7C97561941B82E.xml	Myotis nattereri	Vespertilionidae	Myotis	nattereri		1817	Murin de Natterer @fr | Fransenfledermaus @de | Ratonero de Natterer @es | Natterer's Bat @en	Vespertilio natterer: Kuhl, 1817 , Hanau, Hessen , Germany . Subgenus Myotis ; myotis species group. The taxonomy of the nattereri species complex is currently very convoluted. Four cryptic species have currently been recognized as distinct ( M. nattereri , M. crypticus, M. escalerai , and M. zenatius) based on morphometric and genetic data, and a further two more species (hoveli and tschuliensis) may be recognizable from the Levant and Caucasus that appearto be related to M. schaubi . Anothercryptic species may also be found on Corsica based on limited genetic data; the populations on other large Mediterranean islandsare also currently uncertain due to the recent taxonomic changes. Myotis natterer and M. crypticus are sister species whereas M. escalerai and M. zenatiusare sister to one another with M. schaubi being related to either clade based on various genetic studies. The names hoveli and tschuliensis are here included as subspecies pending further studies that may prove their distinctiveness. Three subspecies recognized.	M. n. nattereriK uhl, 1817 — broadly extended in N, C & E Europe until 60° N and W Turkey . M.n. hoveli D.L.Harrison,1964—SCTurkey,WSyria,Lebanon,Israel,andWJordan. M. n. tschuliensis Kuzyakin, 1935 — Crimea S to Caucasus region. Isolated populations from W Russia , Crete, Cyprus , and Iraq , Iran , and Turkmenistan are not currently assigned to a subspecies.	Head-body 43-48 mm ,tail 38-47 mm , ear 15-4—17- 5 mm , hindfoot 7-1-9-3, forearm 38-5—41- 2 mm ; weight 7-10 g . Natterer’s Myotis has very distinct color between dorsum (generally brownish to grayish) and venter(pale grayish to whitish). Fur is remarkably long and shaggy. Young tend to be darker than adults, with skin and wings remarkably blackish. It has very broad wings and short wingspans ( 245-300 mm ) that allow very agile and maneuverable flight. Tail membrane has two rows of bristles onits edge, and it is supported by Sshaped calcar. This trait clearly separates Natterer’s Myotis from most other species of Myotis . This line of hairs looks like a brush and is one of the traits used to separate all cryptic species in the Myotis species group. Ears are relatively long and have several folds at outer edges, surpassing muzzle when flattened. Ears are distinctly curved due to multiple folds that allow backward movement. Natterer’s Myotis differs from other similar species such as Bechstein’s Myotis ( M. bechsteinii ) by the number of folds and total ear length. Tragus is long and sharp and surpasses one-half the ear length. Hairless face, especially around eyes,is noticeably pink, with narrow and long muzzle. Tibia is completely naked. Skull is small and delicate with high braincase and strongly concave forehead regions; there are no sagittal or occipital crests. P? is more than one-half height and about three-quarters crown area of P? and is within tooth row; lower molars are myotodont. Chromosomal complement has 2n = 44 and FNa = 52 ( Germany ).	Usually forested areas in a wide variety of deciduous and coniferous woodlands, orchards, gardens, parks, tree plantations, and pastures from sea level to up to ¢. 2000 m . Natterer’s Myotis hunts along rivers and in riparian forests and meadows.	Natterer’s Myotis foragesflying slowly 1-4 m above the ground and hunts prey on the ground and in vegetation. It can hover perfectly over prey before gleaning it from a surface. It mainly eats spiders (e.g. harvestmen), caterpillars, and flies. It eats beetles, moths, centipedes, and other insects in smaller proportions. It uses echolocation to detect prey far from a substrate and can hunt insects found in vegetation directly in flight or byfirst picking them off vegetation with their tail membrane. Natterer’s Myotis will move up to 4 km each night to foraging areas, covering up to 500 ha. It has been suggested thatits feeding preferences vary from Geoffroy’s Myotis ( M. emarginatus ) to avoid competition.	Maternity colonies are in tree holes (e.g. hollow trees, rotten branches, abandoned woodpecker nests, etc.), sometimes rock or wall crevices, bat boxes, hollow bricks and even buildings. These colonies usually have 20-50 adult females and young. Colonies in buildings can have more than 120 individuals. Births occur in early summer(from earlyJune to earlyJuly), and young start flying after 20 daysof life, being totally independent c.1 month after birth. Females only have one young per year. Single males often aggregate in colonies up to 25 individuals, but they are also found in maternity colonies. Natterer’s Myotis mate at swarming sites and in hibernacula.	Single Natterer’s Myotis can be found roosting in various roosts (e.g. tree holes, hollow trees, cliffs, small crevices and cracks, abandoned buildings, bat boxes, tunnels, caves, and mines). Natterer’s Myotis is very flexible relative to emerging from day roosts. They can be found foraging early at night, late at night, and even during the day. They generally emerge more than 30 minutes after the sunset. It has very slow and agile flight, capable of maneuvering in small and cluttered spaces. Echolocation calls are short and highly modulated pulses,starting at 100-150 kHz and ending at 20 kHz or even lower (one of the broadest frequency range of any Myotis ;.130 kHz). In general, its pulses are very short, with an average duration of only 3-5 milliseconds. Its echolocation is clearly distinct from the rest of small Myotis species.	Natterer’s Myotis is considered sedentary, usually not moving more than 40 km between summer, swarming, and hibernation roosts. In extreme cases, it reportedly moved 266-327 km . During the mating period, it swarms mostly in front of swarming sites, especially from Augustto late October. At swarming sites, it can occur with other species of Myotis such as Daubenton’s Myotis ( M. daubentonii ) or Bechstein’s Myotis . In winter, individuals form large hibernation coloniesin caves and mines, exceeding 8000 individuals in some cases, or small clusters isolated in mines, tunnels, and other underground roosts.	Classified as Least Concern on The IUCN Red List. Natterer’s Myotis is not very common, but populations in Central Europe seem to be stable. Itis sensitive to habitat loss and fragmentation. It is also one of the most affected bat species by sticky flypaper. Disturbance and loss of old buildings and refurbishing of houses used as roosts might reduce available roosts.	Arlettaz (1996b) | Coraman et al. (2019) | Halczok etal. (2017) | Hope & Jones (2012) | Hope etal. (2014) | Hutson, Aulagnier & Spitzenberger (2008) | Juste et al. (2019) | Melcén et al. (2007) | Mortimer (2006) | Parsons & Jones (2003) | Puechmaille, Allegrini et al. (2012) | Salicini et al. (2012, 2013) | Shiel et al. (1991) | Siemers & Schnitzler (2000) | Siemers & Swift (2006) | Smith & Racey (2005) | Swift (1997) | Swift & Racey (2002) | Volleth & Heller (2012) | Zeale etal. (2016)	https://zenodo.org/record/6399003/files/figure.png	489. Natterer’s Myotis Myotis nattereri French: Murin de Natterer / German: Fransenfledermaus / Spanish: Ratonero de Natterer Other common names: Natterer's Bat Taxonomy. Vespertilio natterer: Kuhl, 1817 , Hanau, Hessen , Germany . Subgenus Myotis ; myotis species group. The taxonomy of the nattereri species complex is currently very convoluted. Four cryptic species have currently been recognized as distinct ( M. nattereri , M. crypticus, M. escalerai , and M. zenatius) based on morphometric and genetic data, and a further two more species (hoveli and tschuliensis) may be recognizable from the Levant and Caucasus that appearto be related to M. schaubi . Anothercryptic species may also be found on Corsica based on limited genetic data; the populations on other large Mediterranean islandsare also currently uncertain due to the recent taxonomic changes. Myotis natterer and M. crypticus are sister species whereas M. escalerai and M. zenatiusare sister to one another with M. schaubi being related to either clade based on various genetic studies. The names hoveli and tschuliensis are here included as subspecies pending further studies that may prove their distinctiveness. Three subspecies recognized. Subspecies and Distribution. M. n. nattereriKuhl, 1817 — broadly extended in N, C & E Europe until 60° N and W Turkey . M.n.hoveliD.L.Harrison,1964—SCTurkey,WSyria,Lebanon,Israel,andWJordan. M. n. tschuliensis Kuzyakin, 1935 — Crimea S to Caucasus region. Isolated populations from W Russia , Crete, Cyprus , and Iraq , Iran , and Turkmenistan are not currently assigned to a subspecies. Descriptive notes. Head-body 43-48 mm ,tail 38-47 mm , ear 15-4—17- 5 mm , hindfoot 7-1-9-3, forearm 38-5—41- 2 mm ; weight 7-10 g . Natterer’s Myotis has very distinct color between dorsum (generally brownish to grayish) and venter(pale grayish to whitish). Fur is remarkably long and shaggy. Young tend to be darker than adults, with skin and wings remarkably blackish. It has very broad wings and short wingspans ( 245-300 mm ) that allow very agile and maneuverable flight. Tail membrane has two rows of bristles onits edge, and it is supported by Sshaped calcar. This trait clearly separates Natterer’s Myotis from most other species of Myotis . This line of hairs looks like a brush and is one of the traits used to separate all cryptic species in the Myotis species group. Ears are relatively long and have several folds at outer edges, surpassing muzzle when flattened. Ears are distinctly curved due to multiple folds that allow backward movement. Natterer’s Myotis differs from other similar species such as Bechstein’s Myotis ( M. bechsteinii ) by the number of folds and total ear length. Tragus is long and sharp and surpasses one-half the ear length. Hairless face, especially around eyes,is noticeably pink, with narrow and long muzzle. Tibia is completely naked. Skull is small and delicate with high braincase and strongly concave forehead regions; there are no sagittal or occipital crests. P? is more than one-half height and about three-quarters crown area of P? and is within tooth row; lower molars are myotodont. Chromosomal complement has 2n = 44 and FNa = 52 ( Germany ). Habitat. Usually forested areas in a wide variety of deciduous and coniferous woodlands, orchards, gardens, parks, tree plantations, and pastures from sea level to up to ¢. 2000 m . Natterer’s Myotis hunts along rivers and in riparian forests and meadows. Food and Feeding. Natterer’s Myotis foragesflying slowly 1-4 m above the ground and hunts prey on the ground and in vegetation. It can hover perfectly over prey before gleaning it from a surface. It mainly eats spiders (e.g. harvestmen), caterpillars, and flies. It eats beetles, moths, centipedes, and other insects in smaller proportions. It uses echolocation to detect prey far from a substrate and can hunt insects found in vegetation directly in flight or byfirst picking them off vegetation with their tail membrane. Natterer’s Myotis will move up to 4 km each night to foraging areas, covering up to 500 ha. It has been suggested thatits feeding preferences vary from Geoffroy’s Myotis ( M. emarginatus ) to avoid competition. Breeding. Maternity colonies are in tree holes (e.g. hollow trees, rotten branches, abandoned woodpecker nests, etc.), sometimes rock or wall crevices, bat boxes, hollow bricks and even buildings. These colonies usually have 20-50 adult females and young. Colonies in buildings can have more than 120 individuals. Births occur in early summer(from earlyJune to earlyJuly), and young start flying after 20 daysof life, being totally independent c.1 month after birth. Females only have one young per year. Single males often aggregate in colonies up to 25 individuals, but they are also found in maternity colonies. Natterer’s Myotis mate at swarming sites and in hibernacula. Activity patterns. Single Natterer’s Myotis can be found roosting in various roosts (e.g. tree holes, hollow trees, cliffs, small crevices and cracks, abandoned buildings, bat boxes, tunnels, caves, and mines). Natterer’s Myotis is very flexible relative to emerging from day roosts. They can be found foraging early at night, late at night, and even during the day. They generally emerge more than 30 minutes after the sunset. It has very slow and agile flight, capable of maneuvering in small and cluttered spaces. Echolocation calls are short and highly modulated pulses,starting at 100-150 kHz and ending at 20 kHz or even lower (one of the broadest frequency range of any Myotis ;.130 kHz). In general, its pulses are very short, with an average duration of only 3-5 milliseconds. Its echolocation is clearly distinct from the rest of small Myotis species. Movements, Home range and Social organization. Natterer’s Myotis is considered sedentary, usually not moving more than 40 km between summer, swarming, and hibernation roosts. In extreme cases, it reportedly moved 266-327 km . During the mating period, it swarms mostly in front of swarming sites, especially from Augustto late October. At swarming sites, it can occur with other species of Myotis such as Daubenton’s Myotis ( M. daubentonii ) or Bechstein’s Myotis . In winter, individuals form large hibernation coloniesin caves and mines, exceeding 8000 individuals in some cases, or small clusters isolated in mines, tunnels, and other underground roosts. Status and Conservation. Classified as Least Concern on The IUCN Red List. Natterer’s Myotis is not very common, but populations in Central Europe seem to be stable. Itis sensitive to habitat loss and fragmentation. It is also one of the most affected bat species by sticky flypaper. Disturbance and loss of old buildings and refurbishing of houses used as roosts might reduce available roosts. Bibliography. Arlettaz (1996b), Coraman et al. (2019), Halczok etal. (2017), Hope & Jones (2012), Hope etal. (2014), Hutson, Aulagnier & Spitzenberger (2008), Juste et al. (2019), Melcén et al. (2007), Mortimer (2006), Parsons & Jones (2003), Puechmaille, Allegrini et al. (2012), Salicini et al. (2012, 2013), Shiel et al. (1991), Siemers & Schnitzler (2000), Siemers & Swift (2006), Smith & Racey (2005), Swift (1997), Swift & Racey (2002), Volleth & Heller (2012), Zeale etal. (2016).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis nattereri	Myotis	Unassigned-Myotis	nattereri	Kuhl	1817	1	Die Deutschen Flederm&auml;use. Hanau	p. 14, 33	Natterer's Myotis	 spelaeus Koch, 1865; <b>tschuliensis</b> Kuzyakin, 1935;  typus Koch, 1865; ;	Germany, Hessen, Hanau.	Ireland, Great Britain, Europe (except N Scandinavia), Morocco, N Algeria, Turkey, Israel, Jordan, Lebanon, Iraq, Iran, Bulgaria, Crimea and Caucasus to Turkmenistan.	Not listed.	Least Concern	Does not include araxenus or bombinus ; see Horácek and Hanák (1984), Kawai et al. (2003), and Çoraman et al. (2019). Does not include escalerai ; see Ibáñez et al. (2006). Does not include hoveli ; see Çoraman et al. (2019). Does not include tschuliensis ; see Çoraman et al. (2019), Smirnov et al. (2020), Kruskop and Solovyeva (2020), and Uvizl and Benda (2021). A cryptic species may occur in Austria and Northern Italy; see Mayer et al. (2007). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Puechmaille et al. (2012), Salicini et al. (2011), Çoraman et al. (2019), Juste et al. (2019), and Smirnov et al. (2020) for additional discussion of the M. nattereri complex.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis nattereri	23	Natterer's Myotis	Natterer's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	nattereri	Kuhl	1817	1						Hanau, Hessen, Germany.			nattereri (Kuhl, 1817)|spelaeus (L. Koch, 1865)|typus (L. Koch, 1865)	previously included M. escalarae, M. tschuliensis, and M. hoveli	Ibáñez, C., García-Mudarra, J. L., Ruedi, M., Stadelmann, B., & Juste, J. (2006). The Iberian contribution to cryptic diversity in European bats. Acta Chiropterologica, 8(2), 277-297.|Kruskop, S. V. and Solovyeva, E. N. (2020). Validating the relationships: which species of Myotis "nattereri" group (Chiroptera: Vespertilionidae) actually inhabits the Caucasus. Mammali, 1-10.|Uvizl, M., & Benda, P. (2021). Diversity and distribution of the Myotis nattereri complex (Chiroptera: Vespertilionidae) in the Middle East: filling the gaps. Mammalian Biology, 1-15.	Ireland|United Kingdom|France|Luxembourg|Belgium|Netherlands|Germany|Denmark|Norway|Sweden|Finland|Switzerland|Liechtenstein|Austria|Czech Republic|Slovakia|Poland|Hungary|Slovenia|Croatia|Bosnia & Herzegovina|Serbia|Kosovo|Montenegro|Albania|North Macedonia|Greece|Bulgaria|Turkey|Romania|Ukraine|Belarus|Lithuania|Latvia|Estonia|Russia	Asia|Europe	Palearctic	LC	0	0	0	Myotis_nattereri	0	sciname match	Myotis_nattereri	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Myotis nattereri	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	nattereri	(Kuhl, 1817)	Does not include ;Myotis schaubi ;(Armenia and western Iran), ;M. escalerai, ; M. crypticus, ; M. zenatius, ; M. hoveli, ; M. tschuliensis, ; and the not formally described Myotis sp. C. (endemic subpopulation of Corsica). It also doesn't include ;M. bombinus ; (southeastern Siberia, northeastern China, Korea, Japan).	20000000	Myotis nattereri	Least Concern		2020	2019-05-14 00:00:00 UTC	3.1	English	The species is widespread and abundant, and there is no evidence of current significant population decline. Consequently it is assessed as Least Concern.	Maternity colonies roost in tree hollows, spaces under the loose bark and bat-boxes, crevices in buildings and bridges, frequently in cattle stables as well as in hollow blocks. The colony usually consists of 30-80 individuals. In buildings, however, they can also be more numerous (up to 200 females). As many other woodland bats, it is prone to frequent change of roosts (1-2 times a week). Male colonies of up to 30 animals also occupy tree cavities, bat and bird boxes and buildings. It hibernates in underground habitats (caves, cellars and mines), usually solitary or in small clusters of 2-5 individuals (Simon et al. 2012). The best known hibernation site is Nietoperek fortification system in Poland with more than 300 recorded hibernating bats (EUROBATS 2019). It hunts over a variety of habitats such as meadows, pastures, orchards, broadleaved woods, open coniferous forests and riparian areas. Intense foraging was observed inside cattle sheds. Feeding areas range between 128 and 580 ha. The core of a feeding area can be up to 4 km away from the roost and individuals are faithful to them, returning there regularly. Commuting routes between the roost and the core area are also utilised for foraging. In a German study, the average distance between maternity roosts and feeding areas was 1.5 km.M. nattereri can capture prey on the wing and by gleaning resting insects from the surface of vegetation using the tail membrane and/or feet. The chief components of its diet are Diptera and Coleoptera, especially Brachycera and Curculionidae. Diurnally active insects, insects which rarely fly and non-flying arthropods are also eaten (see the literature review in Kyheröinen et al. 2019). Faecal pellets collected at a hibernation site in southern England for two winters, indicated that M. nattereri forages throughout winter on non-volant Aranea, Isopoda and Lepidoptera (Hope et al. 2014). Movements between summer, autumn and winter roosts up to 185 km were recorded (Schober, Grimmberger 1998). The maximum known longevity for males and females was 21,5 and 17,5 years respectively (Ohlendorf 2002, Topal 2011).	Major threats in Europe are as follow (Simon et al. 2012): Habitat and roost loss due to abandonment of cattle husbandry (stable housing). Enlargement of crops leading to the destruction of small-scale, insect-rich cultivated landscapes. Insect loss due to the intensive agriculture, forestry and use of pesticides, also in livestock farming. Reduction of summer and winter roosts' availability due to forest operations (removal of deadwood and old trees with cavities), renovation of buildings, bridges, cattle stables etc. Disappearance of small waters in the forest and in the country. Loss of hibernacula due to collapse, closing or conversion of e.g. ice cellars, bunkers and other underground habitats.Disturbances (noise, vandalism) in the winter quarters.The species is sensitive to light pollution (Voigt et al. 2018). It is exposed to a high risk of collisions with vehicles (Fensome, Mathews 2016).	M. nattereri is widespread in Europe, but the population trend is uncertain. Within the European Union, it has ;Unfavourable-Inadequate status in all biogeographical regions but Boreal, where it was assessed as ;Unfavourable-Bad (EEA 2013). Among major range states outside the EU, Moldova and Belarus reported negative population trends (Moldova 2018, Shpak 2018), whereas it is rather stable in Ukraine (Ukraine 2018) and Russia (S. Gazaryan, pers. comm.). In Norway, it hasn't been found since 2010 despite special efforts (Henriksen, Hilmo 2015). According to the combined prototype European hibernating bat indicator (Haysom et al. 2013), a moderate increasing trend was observed.	Stable	Taxonomic position of M. nattereri ; is not completely resolved, because it was split off to a complex of segregate species. ;A series of genetic surveys focusing on Western European populations showed the existence of at least five major mitochondrial lineages which may represent separate species. Three of those lineages were located in continental Europe, one in Corsica and the another one in Northwestern Africa. The linage restricted to Iberia and the Balearic Islands was erected as a distinct species M. escalerai (Ibáñez et al ., 2006, Salicini et al. 2011). Lately, bats which inhabit mountain areas of provinces of central and northern Spain, southern France, the Italian Peninsula, adjacent southwestern parts of Austria and ;marginal areas to the north and west of the Alps ;were assigned to the new taxon Myotis crypticus . Another cryptic species ;Myotis zenatius ;is probably endemic and occurs in the Mediterranean region of Morocco and Algeria, and possibly Tunisia (Juste et al. 2019). Presumably, one more not yet described species of the complex (M. sp. C) occurs in Corsica (Puechmaille et al. 2012). In the light of differences in the nuclear DNA, Ñ€opulations from Israel and south‐eastern Anatolia were recently described as s separate species ;M. hoveli and the Caucasian and Crimean ones assigned to another segregate taxon ;M. tschuliensis ( Çoraman et al. 2019). If this taxonomic outlooks are accepted, at least 7 valid species may occur within the range of former M. nattereri s.l.: M. nattereri s.str., M. escalerai, Myotis crypticus, ; Myotis zenatius, ; M. hoveli, ; M. tschuliensis and ;not formally described M. sp. C. ;Accounting for aforementioned novelties, M. nattereri s.str occupies areas of Europe situated northwards and eastwards from the Alps, including the UK, France, Germany, Scandinavia, the Baltic states, the Balkans, Carpathians and entire Eastern Europe up to the Urals. However, distributional limits of the "mother" taxon in Central Europe are still uncertain after the resent splits.		Terrestrial	It is protected by national legislation in most of the main ; range states, except Russia. There are also international legal obligations for its protection through the Convention on Migratory Species (CMS) and EUROBATS Agreement. It is included in Appendix II ; of the Council of Europe’s Convention on the Conservation of European Wildlife and Natural Habitats (Bern Convention). Also included in Annex IV of EU Habitats and Species Directive. Important habitats partially protected by Natura 2000 sites and national networks of protected areas. The taxonomic position of ;M. nattereri ; is not completely resolved, because it was split off to a complex of segregate species.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	nattereri	Kuhl	1817	1	Die Deutschen Flederm&auml;use. Hanau	p. 14, 33	Natterer's Myotis	 spelaeus Koch, 1865; <b>tschuliensis</b> Kuzyakin, 1935;  typus Koch, 1865; ;	Germany, Hessen, Hanau.	Ireland, Great Britain, Europe (except N Scandinavia), Morocco, N Algeria, Turkey, Israel, Jordan, Lebanon, Iraq, Iran, Bulgaria, Crimea and Caucasus to Turkmenistan.	Not listed.	Least Concern	Does not include araxenus or bombinus ; see Horácek and Hanák (1984), Kawai et al. (2003), and Çoraman et al. (2019). Does not include escalerai ; see Ibáñez et al. (2006). Does not include hoveli ; see Çoraman et al. (2019). Does not include tschuliensis ; see Çoraman et al. (2019), Smirnov et al. (2020), Kruskop and Solovyeva (2020), and Uvizl and Benda (2021). A cryptic species may occur in Austria and Northern Italy; see Mayer et al. (2007). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Puechmaille et al. (2012), Salicini et al. (2011), Çoraman et al. (2019), Juste et al. (2019), and Smirnov et al. (2020) for additional discussion of the M. nattereri complex.	Myotis nattereri	1005445	23	Natterer's Myotis	Natterer's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	nattereri	Kuhl	1817	1						Hanau, Hessen, Germany.			nattereri (Kuhl, 1817)|spelaeus (L. Koch, 1865)|typus (L. Koch, 1865)	previously included M. escalarae, M. tschuliensis, and M. hoveli	Ibáñez, C., García-Mudarra, J. L., Ruedi, M., Stadelmann, B., & Juste, J. (2006). The Iberian contribution to cryptic diversity in European bats. Acta Chiropterologica, 8(2), 277-297.|Kruskop, S. V. and Solovyeva, E. N. (2020). Validating the relationships: which species of Myotis "nattereri" group (Chiroptera: Vespertilionidae) actually inhabits the Caucasus. Mammali, 1-10.|Uvizl, M., & Benda, P. (2021). Diversity and distribution of the Myotis nattereri complex (Chiroptera: Vespertilionidae) in the Middle East: filling the gaps. Mammalian Biology, 1-15.				Ireland|United Kingdom|France|Luxembourg|Belgium|Netherlands|Germany|Denmark|Norway|Sweden|Finland|Switzerland|Liechtenstein|Austria|Czech Republic|Slovakia|Poland|Hungary|Slovenia|Croatia|Bosnia & Herzegovina|Serbia|Kosovo|Montenegro|Albania|North Macedonia|Greece|Bulgaria|Turkey|Romania|Ukraine|Belarus|Lithuania|Latvia|Estonia|Russia	Asia|Europe	Palearctic	LC	0	0	0	Myotis_nattereri	0	sciname match	Myotis_nattereri	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_nattereri	1005445	23	Natterer's Myotis	Natterer's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	nattereri	Kuhl	1	Vespertilio Nattereri	Kuhl, H. 1817. Die deutschen Fledermäuse. None, Hanau, 67 pp.	https://nbn-resolving.org/urn:nbn:de:hebis:30-91692				Hanau, Hessen, Germany.			previously included M. escalarae, M. tschuliensis, and M. hoveli	Ibáñez, C., García-Mudarra, J. L., Ruedi, M., Stadelmann, B., & Juste, J. (2006). The Iberian contribution to cryptic diversity in European bats. Acta Chiropterologica, 8(2), 277-297.|Kruskop, S. V. and Solovyeva, E. N. (2020). Validating the relationships: which species of Myotis "nattereri" group (Chiroptera: Vespertilionidae) actually inhabits the Caucasus. Mammali, 1-10.|Uvizl, M., & Benda, P. (2021). Diversity and distribution of the Myotis nattereri complex (Chiroptera: Vespertilionidae) in the Middle East: filling the gaps. Mammalian Biology, 1-15.				Ireland|United Kingdom|France|Luxembourg|Belgium|Netherlands|Germany|Denmark|Norway|Sweden|Finland|Switzerland|Liechtenstein|Austria|Czech Republic|Slovakia|Poland|Hungary|Slovenia|Croatia|Bosnia and Herzegovina|Serbia|Kosovo|Montenegro|Albania|North Macedonia|Greece|Bulgaria|Turkey|Romania|Ukraine|Belarus|Lithuania|Latvia|Estonia|Russia	Asia|Europe	Palearctic	LC	0	0	0	Myotis_nattereri	0	sciname match	Myotis_nattereri	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	nattereri	Kuhl	1817	1	Die Deutschen Flederm&auml;use. Hanau	p. 6, 25	Natterer's Myotis	spelaeus Koch, 1865; tschuliensis Kuzyakin, 1935;  typus Koch, 1865;;	Germany, Hessen, Hanau.	Ireland, Great Britain, Europe (except N Scandinavia), Morocco, N Algeria, Turkey, Israel, Jordan, Lebanon, Iraq, Iran, Bulgaria, Crimea and Caucasus to Turkmenistan.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/215492021/211005466/' target='_blank'>Least Concern</a>	Does not include araxenus or bombinus; see Horácek and Hanák (1984), Kawai et al. (2003), and Çoraman et al. (2019). Does not include escalerai; see Ibáñez et al. (2006). Does not include hoveli; see Çoraman et al. (2019). Does not include tschuliensis; see Çoraman et al. (2019), Smirnov et al. (2020), Kruskop and Solovyeva (2020), and Uvizl and Benda (2021). A cryptic species may occur in Austria and Northern Italy; see Mayer et al. (2007). Reviewed in part by Harrison and Bates (1991) and Horácek et al. (2000). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Puechmaille et al. (2012), Salicini et al. (2011), Çoraman et al. (2019), Juste et al. (2019), and Smirnov et al. (2020) for additional discussion of the M. nattereri complex.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis nattereri; Myotis nattereri; Myotis nattereri; Myotis nattereri; Myotis nattereri; Myotis nattereri; nattereri; tschuliensis; escalerae; hoveli; spelaeus; typus; nattereri; hoveli; tschuliensis; tschuliensis; spelaeus; tschuliensis - typus; nattereri; spelaeus; typus; Murin de Natterer; Fransenfledermaus; Ratonero de Natterer; Natterer's Bat; Natterer's Myotis; Natterer's Bat; Natterer's Myotis; Natterer's Myotis; M. nattereri