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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L921	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis morrisi	Myotis morrisi	Myotis morrisi	Myotis morrisi	Myotis morrisi	Myotis morrisi	Myotis morrisi	Myotis morrisi	Myotis morrisi	Myotis morrisi	Myotis morrisi	Myotis morrisi	Myotis morrisi	Myotis morrisi	Myotis morrisi		[MSW2] Subgenus Myotis.; [HMW] Myotis morrist Hill in Hill & Morris, 1971 , “ ‘Forward Base Three’, mouth of Didessa River, Blue Nile Gorge [Walga, Ethiopia ]. 10° 05' N , 35° 38' E , alt. ¢. 1,000 m .” Subgenus Chrysopteron. This species has not yet been included in a phylogenetic study. Monotypic.; [batnames2022] None.; [batnames2023] None.; [batnames2025_1.7] None.														morrisi				morrisi 	morrisi 			morrisi J. Edwards Hill in J. Edwards Hill & P. Morris, 1971		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Ethiopia; ref. 4.80	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis morrisi	Ethiopia, Walaga, Didessa River mouth.	Hill	1971	Bull. Br. Mus. (Nat. Hist.) Zool., 21:43.	Distribution: Known only from northern Ethiopia.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Ethiopia, Nigeria; ref. 4.151	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Hill	1971	Bull. Brit. Mus. (Nat. Hist.) Zool., 21:43.	Subgenus Myotis.	Ethiopia, Nigeria.	Ethiopia, Walaga, Didessa River mouth.		HILL	1971	Size fairly large (forearm length, 45-46 mm; condylobasal length, 16-17 mm). Rostrum relatively narrow. Middle upper premolar somewhat displaced medially from the toothrow.	Distribution: Known only from northern Ethiopia.	No subspecies.		102	species	M. morrisi	HILL	1971	Myotis	subgenus	Myotis morrisi				Size fairly large (forearm length, 45-46 mm; condylobasal length, 16-17 mm). Rostrum relatively narrow. Middle upper premolar somewhat displaced medially from the toothrow.	No subspecies.		15. M. morrisi HILL 1971 [emarginatus group].	15	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis morrisi	Myotis		morrisi	Hill		1971		Bull. Brit. Mus. (Nat. Hist.) Zool.	21		43		Morris's Myotis	Ethiopia, Walaga, Didessa River mouth.	Ethiopia, Nigeria.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.			4C3D87E8FF456AFBFA5090C71B05B9DD	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	952	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF456AFBFA5090C71B05B9DD.xml	Myotis morrisi	Vespertilionidae	Myotis	morrisi	Hill	1971	Murin de Morris @fr | Morris-Mausohr @de | Ratonero de Morris @es | Morris's Bat @en | Morris's Hairy Bat @en	Myotis morrist Hill in Hill & Morris, 1971 , “ ‘Forward Base Three’, mouth of Didessa River, Blue Nile Gorge [Walga, Ethiopia ]. 10° 05' N , 35° 38' E , alt. ¢. 1,000 m .” Subgenus Chrysopteron. This species has not yet been included in a phylogenetic study. Monotypic.	Known only from two distant sites: type locality in W Ethiopia ; and NE Nigeria (Numan, Adamawa State ). May be widespread across Sudan Savanna Biome.	Head-body c. 48 mm , tail 45 mm , ear 16-17 mm , hindfoot 9- 5— 10 mm , forearm 45-2-45- 7 mm ; weight 8 g (all measurements from a single individual). Greatest length of skull 16:3-17- 5 mm , and maxillary tooth row 6:6- 6-8 mm . Pelage is dense and fairly woolly; dorsally bright orange-brown (hairs tricolored, creamy white, with basal quarter dark brown and tip orange-brown; dorsal hair length c. 7 mm ); ventrally dull creamy white to orange-beige, chin and flanks slightly darker (hairs unicolored, vs. bicolored or tricolored in all other African Myotis ). Muzzle is pink. Medium-sized ears are pinkish basally, shading to brown at tips; tragus is pink, bluntly pointed, and half ear-length,tip slightly directed backward. Wing membranes are dark brown, slightly pigmented yellow along leading edge and along a narrow band posterior to forearm, as well as between body and a line roughly joining elbow and ankle; wings are naked except for band on dorsal surface adjacent to body; wing membranes are attached to base of first toe; calcar extends half-way to tip of tail; no postcalcarial lobe; interfemoral membrane is yellow, with dorsal pelage extending over its distal half but no fringe or bristle-like hairs on posterior edge; terminal vertebra may or may not protrude beyond membrane. Two other African Myotis have dichromatic or contrasting wing pattern: in some populations of Temminck’s Myotis (M. trcolor), black and dark reddish-brown wing pattern might be present and ventral hairs bicolored; Welwitsch’s Myotis (M. welwitschii ) is larger (forearm length 52-60 mm ), with very conspicuous black-and-orange wing pattern and bicolored venter. Hindfoot is relatively short and tibiae hairy. Skull is elongated with uninflated, globular braincase and supraorbital region; palate long and narrow, anterior emargination rounded posteriorly, extending almost to a line joining the centers of the canines; no sagittal crests. Upper incisors bicuspid, I? is longer than wide, I* wider than long; P* of same height as I’, in contact with canine; P° reduced (reaching only slightly above cingulum of P? and slightly less than half crown area of P*) and slightly or fully displaced from tooth row, but P? and P* are not in contact; I, and I, with four cusps; lower premolars not particularly compressed; P, in contact with both others and slightly higher than P,. Anterior palatal emargination very narrow.	Morris's Myotis occupies dry scrub in lowlands. The holotype was found hunting over the Blue Nile in an area of riverine scrub and maize fields. The Nigerian specimen was found in a region of short grass and Acacia ( Fabaceae ) bush, with some silk-cotton trees.	Morris's Myotis forages at moderate height ( 1-3 m ).	No information.	No information.	No information.	Classified as Data Deficient on The IUCN Red List. Morris’s Myotis is known only from two specimens from widely separated sites.	ACR (2018) | Csorba, Chou Cheng-Han (2014) | Hill & Morris (1971) | Hill et al. (1988) | Jacobs et al. (2008h) | Koopman (1994) | Largen et al. (1974) | Simmons (2005) | Yalden (2013a) | Yalden & Largen (1992)	https://zenodo.org/record/6398843/files/figure.png	430. Morris's Myotis Myotis morrisi French: Murin de Morris / German: Morris-Mausohr / Spanish: Ratonero de Morris Other common names: Morris's Bat , Morris's Hairy Bat Taxonomy. Myotis morrist Hill in Hill & Morris, 1971 , “ ‘Forward Base Three’, mouth of Didessa River, Blue Nile Gorge [Walga, Ethiopia ]. 10° 05' N , 35° 38' E , alt. ¢. 1,000 m .” Subgenus Chrysopteron. This species has not yet been included in a phylogenetic study. Monotypic. Distribution. Known only from two distant sites: type locality in W Ethiopia ; and NE Nigeria (Numan, Adamawa State ). May be widespread across Sudan Savanna Biome. Descriptive notes. Head-body c. 48 mm , tail 45 mm , ear 16-17 mm , hindfoot 9- 5— 10 mm , forearm 45-2-45- 7 mm ; weight 8 g (all measurements from a single individual). Greatest length of skull 16:3-17- 5 mm , and maxillary tooth row 6:6- 6-8 mm . Pelage is dense and fairly woolly; dorsally bright orange-brown (hairs tricolored, creamy white, with basal quarter dark brown and tip orange-brown; dorsal hair length c. 7 mm ); ventrally dull creamy white to orange-beige, chin and flanks slightly darker (hairs unicolored, vs. bicolored or tricolored in all other African Myotis ). Muzzle is pink. Medium-sized ears are pinkish basally, shading to brown at tips; tragus is pink, bluntly pointed, and half ear-length,tip slightly directed backward. Wing membranes are dark brown, slightly pigmented yellow along leading edge and along a narrow band posterior to forearm, as well as between body and a line roughly joining elbow and ankle; wings are naked except for band on dorsal surface adjacent to body; wing membranes are attached to base of first toe; calcar extends half-way to tip of tail; no postcalcarial lobe; interfemoral membrane is yellow, with dorsal pelage extending over its distal half but no fringe or bristle-like hairs on posterior edge; terminal vertebra may or may not protrude beyond membrane. Two other African Myotis have dichromatic or contrasting wing pattern: in some populations of Temminck’s Myotis (M. trcolor), black and dark reddish-brown wing pattern might be present and ventral hairs bicolored; Welwitsch’s Myotis (M. welwitschii ) is larger (forearm length 52-60 mm ), with very conspicuous black-and-orange wing pattern and bicolored venter. Hindfoot is relatively short and tibiae hairy. Skull is elongated with uninflated, globular braincase and supraorbital region; palate long and narrow, anterior emargination rounded posteriorly, extending almost to a line joining the centers of the canines; no sagittal crests. Upper incisors bicuspid, I? is longer than wide, I* wider than long; P* of same height as I’, in contact with canine; P° reduced (reaching only slightly above cingulum of P? and slightly less than half crown area of P*) and slightly or fully displaced from tooth row, but P? and P* are not in contact; I, and I, with four cusps; lower premolars not particularly compressed; P, in contact with both others and slightly higher than P,. Anterior palatal emargination very narrow. Habitat. Morris's Myotis occupies dry scrub in lowlands. The holotype was found hunting over the Blue Nile in an area of riverine scrub and maize fields. The Nigerian specimen was found in a region of short grass and Acacia ( Fabaceae ) bush, with some silk-cotton trees. Food and Feeding. Morris's Myotis forages at moderate height ( 1-3 m ). Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. Morris’s Myotis is known only from two specimens from widely separated sites. Bibliography. ACR (2018), Csorba, Chou Cheng-Han (2014), Hill & Morris (1971), Hill et al. (1988), Jacobs et al. (2008h), Koopman (1994), Largen et al. (1974), Simmons (2005), Yalden (2013a), Yalden & Largen (1992).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis morrisi	Myotis	Chrysopteron	morrisi	Hill	1971	0	Bull. Br. Mus. Nat. Hist. Zool.	21:43	Morris' Myotis	None.	Ethiopia, Walaga, Didessa River mouth.	Ethiopia, Nigeria.	Not listed.	Data Deficient	None.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis morrisi	23	Morris's Myotis	Morris's Bat|Morris's Hairy Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Chrysopteron	morrisi	J. Edwards Hill in J. Edwards Hill & Morris	1971	0	Myotis_morrisi	Hill, J. E. (1971). In Hill, J. E. & Morris, P. Bats from Ethiopia collected by the Great Abbai Expedition, 1968. Bulletin of the British Museum (Natural History). Zoology, 21, 43.	https://www.biodiversitylibrary.org/item/19475#page/57/mode/1up	BM 1970.488		"‘Forward Base Three', mouth of Didessa River, Blue Nile Gorge [Walga, Ethiopia]. 10° 05' N, 35° 38' E, alt. c. 1,000 m."	10.083	35.633	morrisi J. Edwards Hill in J. Edwards Hill & Morris, 1971	NA	NA	Nigeria|Ethiopia	Africa	Afrotropic	DD	0	0	0	Myotis_morrisi	0	sciname match	Myotis_morrisi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14182	Myotis morrisi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	morrisi	Hill, 1971		20000000	Myotis morrisi	Data Deficient		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Data Deficient in view of the absence of sufficient information on its extent of occurrence, threats and conservation status.	Animals have been reported from caves and other subterranean locations in dry and moist savanna habitats. The specimen from Nigeria was 'mist-netted in the Sudan savanna, in a region of short grass and acacia bush, with some silk cotton trees, near the River Benue' (Hill et al. 1988).	The threats to this species are unclear. It may be threatened by cave disturbance and subsistence harvesting for food.	Little information is available on the population abundance or size of this species.	Unknown	This species has only been recorded from two very disjunct sites, the type locality of 'Didessa River mouth, Walaga' in Ethiopia (Hill and Morris 1971) and from Numan, Adamawa Province, in northeastern Nigeria (Hill et al. 1988).		Terrestrial	It is not known if the species is present in any protected areas. Further studies are needed into the distribution, abundance, natural history, and threats to this poorly known species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Chrysopteron	morrisi	Hill	1971	0	Bull. Br. Mus. Nat. Hist. Zool.	21:43	Morris' Myotis	None.	Ethiopia, Walaga, Didessa River mouth.	Ethiopia, Nigeria.	Not listed.	Data Deficient	None.	Myotis morrisi	1005441	23	Morris's Myotis	Morris's Bat|Morris's Hairy Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Chrysopteron	morrisi	J. Edwards Hill in J. Edwards Hill & Morris	1971	0	Myotis_morrisi	Hill, J. E. (1971). In Hill, J. E. & Morris, P. Bats from Ethiopia collected by the Great Abbai Expedition, 1968. Bulletin of the British Museum (Natural History). Zoology, 21, 43.	https://www.biodiversitylibrary.org/item/19475#page/57/mode/1up	BM 1970.488		"‘Forward Base Three', mouth of Didessa River, Blue Nile Gorge [Walga, Ethiopia]. 10° 05' N, 35° 38' E, alt. c. 1,000 m."	10.08333	35.63333	morrisi J. Edwards Hill in J. Edwards Hill & Morris, 1971	NA	NA				Nigeria|Ethiopia	Africa	Afrotropic	DD	0	0	0	Myotis_morrisi	0	sciname match	Myotis_morrisi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_morrisi	1005441	23	Morris's Myotis	Morris's Bat|Morris's Hairy Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Chrysopteron	morrisi	J. Edwards Hill in J. Edwards Hill & P. Morris	0	Myotis morrisi	Hill, J.E. and Morris, P. 1971. Bats from Ethiopia collected by the Great Abbai Expedition, 1968. Bulletin of the British Museum (Natural History) (Zoology)21(2):27-49.	https://www.biodiversitylibrary.org/page/2283109	BMNH:Mamm:1970.488	holotype	https://data.nhm.ac.uk/object/0f2f790f-2b7f-4222-b5ca-58c7e2d4f66f	"‘Forward Base Three', mouth of Didessa River, Blue Nile Gorge [Walga, Ethiopia]. 10° 05' N, 35° 38' E, alt. c. 1,000 m."	10.08333	35.63333	NA	NA				Nigeria|Ethiopia	Africa	Afrotropic	DD	0	0	0	Myotis_morrisi	0	sciname match	Myotis_morrisi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Chrysopteron	morrisi	Hill	1971	0	Bull. Br. Mus. Nat. Hist. Zool.	21:43	Morris' Myotis	None.	Ethiopia, Walaga, Didessa River mouth.	Ethiopia, Nigeria.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14182/22065314/' target='_blank'>Data Deficient</a>	None.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis morrisi; Myotis morrisi; Myotis morrisi; Myotis morrisi; Myotis morrisi; Myotis morrisi; morrisi; Murin de Morris; Morris-Mausohr; Ratonero de Morris; Morris's Bat; Morris's Hairy Bat; Morris's Myotis; Morris's Bat; Morris's Hairy Bat; Morris's Myotis; Morris' Myotis; M. morrisi