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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L917	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Myotis midastactus?	Myotis midastactus	Myotis midastactus	Myotis midastactus	Myotis macropus	Myotis midastactus	Myotis midastactus	Myotis midastactus	Myotis midastactus	Myotis midastactus		[MSW3] Distinct from adversus and moluccarum; see Kitchener et al. (1995b), who revised this complex.; [HMW] Myotis midastactus Moratelli & Wilson, 2014 , “Cercado, Rio Mamoré, Beni , Bolivia , about 23 km W of San Javier (1434S, 64°B5"W).” Subgenus Pyzonix; ruber species group. Myotis midastactus was recently described from 28 specimens originally identified as Myotis simus . Monotypic.; [batnames2022] Specimens of midastactus were previously identified as Myotis simus ; see Moratelli and Wilson (2014).; [MDD2022] recently described; [batnames2023] Specimens of midastactus were previously identified as Myotis simus ; see Moratelli and Wilson (2014).; [MDD2023] recently described; [MDD2025_2.0] recently described; [batnames2025_1.7] Specimens of midastactus were previously identified as Myotis simus; see Moratelli and Wilson (2014).; [MDD2025_2.2] recently described														midastactus				midastactus	midastactus			midastactus Moratelli & D. E. Wilson, 2014						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis macropus	Myotis		macropus	Gould	y	1854		Mammals of Australia			unnumbered page of text		Gould's Large-footed Myotis	South Australia.	S Australia, Victoria (Australia).	IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001).		Distinct from adversus and moluccarum; see Kitchener et al. (1995b), who revised this complex.	4C3D87E8FF576AE8FA5E95AF16EAB77D	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	934	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF576AE8FA5E95AF16EAB77D.xml	Myotis midastactus	Vespertilionidae	Myotis	midastactus	Moratelli & Wilson	2014	Murin midas @fr | Goldgelbes Mausohr @de | Ratonero dorado @es | Midas Myotis @en	Myotis midastactus Moratelli & Wilson, 2014 , “Cercado, Rio Mamoré, Beni , Bolivia , about 23 km W of San Javier (1434S, 64°B5"W).” Subgenus Pyzonix; ruber species group. Myotis midastactus was recently described from 28 specimens originally identified as Myotis simus . Monotypic.	Known only from Bolivia ( Beni and Santa Cruz departments) and Paraguay ( Presidente Hayes Departament).	Head—body ¢. 50-59 mm , tail 36-41 mm , ear 12-15 mm , hindfoot 8-11 mm , forearm 38-2—40- 7 mm ; weight 6-11 g . The Golden Myotis is morphologically similar to the Velvety Myotis ( M. simus ) but easily distinctive to other New World Myotis . It differs from all other South American myotine bats (except the Velvety Myotis ) by having plagiopatagium attached at toes by narrow band of membrane (less than 1-5 mm ). Pelage is woolly, extremely short (less than 5 mm ), and bright golden yellow. Bolivian samples are ocherous orange dorsally, with slightly paler venter. A few specimens are slightly paler, with mixed blocks of grayish and golden fur. Ears are short, extending forward halfway from eye to nostril. Antitragal notch is barely evident; tragus is pointed, slightly curving outward above and convex below, with small triangular lobule at outer base. Ears and membranes are blackish. Skull is moderately long (greatest skull lengths 13-9-15- 1 mm ), and occipital region is flattened. Sagittal and occipital crests are present and vary from low to moderately high. C' is large in length and basal cross section, and there is short distance between C' and P*, resulting in P* and P* being close or in actual contact, displacing P? lingually so that P? is usually not visible in labially. Chromosomal complement has 2n = 44 and FN = 50, with three large and one small metacentric and 17 medium to small acrocentric pairs of autosomes. X-chromosome is medium submetacentric, and Y-chromosome is small acrocentric.	Semideciduous forest (Cerrado ecoregion, Bolivia ) and savanna (Chaco ecoregion, Paraguay ) at elevations of ¢. 150-250 m .	The Golden Myotis is an aerial insectivore. Its diet contains Orthoptera ( Gryllidae ), Hemiptera ( Corixidae ), Hemiptera ( Cicadellidae , Delphacidae ), Coleoptera ( Carabidae , Dytiscidae , Staphylinidae , Scarabaeidae , Curculionidae ), Lepidoptera (at least three families), and Diptera (Nematocera) . Nine of ten prey items in stomachs were less than 10 mm long. Dietary items indicate that it forages in forest interiors and near water bodies.	Ten pregnant Golden Myotis , each with one embryo, were caught in September in Bolivia , and four pregnant females were caught in October in Paraguay .	The Golden Myotis emerges just before sunset. It roosts in hollow trees and holes in the ground and under thatched roofs in Bolivia .	The Golden Myotis roosts with the Argentine Serotine (Eptesicusfurinalis), the Lesser Bulldog Bat ( Noctilio albiventris), and Pallas’s Mastiff Bat (Molossus molossus). Golden Myotis and Lesser Bulldog Bats habiting the same roost had similarly colored fur, texture, and smell. Data from these roosts suggest that Golden Myotis are solitary or form very small groups.	Not assessed on The IUCN Red List.	Aguirre (1994) | Aguirre, Lens & Matthysen (2003) | Anderson (1997) | Espinoza (2007) | Moratelli & Wilson (2014a) | Moratelli, Idarraga & Wilson (2015) | Tarifa & Aguirre (2009)	https://zenodo.org/record/6398751/files/figure.png	391. Golden Myous Myotis midastactus French: Murin midas / German: Goldgelbes Mausohr / Spanish: Ratonero dorado Other common names: Midas Myotis Taxonomy. Myotis midastactus Moratelli & Wilson, 2014 , “Cercado, Rio Mamoré, Beni , Bolivia , about 23 km W of San Javier (1434S, 64°B5"W).” Subgenus Pyzonix; ruber species group. Myotis midastactus was recently described from 28 specimens originally identified as Myotis simus . Monotypic. Distribution. Known only from Bolivia ( Beni and Santa Cruz departments) and Paraguay ( Presidente Hayes Departament). Descriptive notes. Head—body ¢. 50-59 mm , tail 36-41 mm , ear 12-15 mm , hindfoot 8-11 mm , forearm 38-2—40- 7 mm ; weight 6-11 g . The Golden Myotis is morphologically similar to the Velvety Myotis ( M. simus ) but easily distinctive to other New World Myotis . It differs from all other South American myotine bats (except the Velvety Myotis ) by having plagiopatagium attached at toes by narrow band of membrane (less than 1-5 mm ). Pelage is woolly, extremely short (less than 5 mm ), and bright golden yellow. Bolivian samples are ocherous orange dorsally, with slightly paler venter. A few specimens are slightly paler, with mixed blocks of grayish and golden fur. Ears are short, extending forward halfway from eye to nostril. Antitragal notch is barely evident; tragus is pointed, slightly curving outward above and convex below, with small triangular lobule at outer base. Ears and membranes are blackish. Skull is moderately long (greatest skull lengths 13-9-15- 1 mm ), and occipital region is flattened. Sagittal and occipital crests are present and vary from low to moderately high. C' is large in length and basal cross section, and there is short distance between C' and P*, resulting in P* and P* being close or in actual contact, displacing P? lingually so that P? is usually not visible in labially. Chromosomal complement has 2n = 44 and FN = 50, with three large and one small metacentric and 17 medium to small acrocentric pairs of autosomes. X-chromosome is medium submetacentric, and Y-chromosome is small acrocentric. Habitat. Semideciduous forest (Cerrado ecoregion, Bolivia ) and savanna (Chaco ecoregion, Paraguay ) at elevations of ¢. 150-250 m . Food and Feeding. The Golden Myotis is an aerial insectivore. Its diet contains Orthoptera ( Gryllidae ), Hemiptera ( Corixidae ), Hemiptera ( Cicadellidae , Delphacidae ), Coleoptera ( Carabidae , Dytiscidae , Staphylinidae , Scarabaeidae , Curculionidae ), Lepidoptera (at least three families), and Diptera (Nematocera) . Nine of ten prey items in stomachs were less than 10 mm long. Dietary items indicate that it forages in forest interiors and near water bodies. Breeding. Ten pregnant Golden Myotis , each with one embryo, were caught in September in Bolivia , and four pregnant females were caught in October in Paraguay . Activity patterns. The Golden Myotis emerges just before sunset. It roosts in hollow trees and holes in the ground and under thatched roofs in Bolivia . Movements, Home range and Social organization. The Golden Myotis roosts with the Argentine Serotine (Eptesicusfurinalis), the Lesser Bulldog Bat ( Noctilio albiventris), and Pallas’s Mastiff Bat (Molossus molossus). Golden Myotis and Lesser Bulldog Bats habiting the same roost had similarly colored fur, texture, and smell. Data from these roosts suggest that Golden Myotis are solitary or form very small groups. Status and Conservation. Not assessed on The IUCN Red List. Bibliography. Aguirre (1994), Aguirre, Lens & Matthysen (2003), Anderson (1997), Espinoza (2007), Moratelli & Wilson (2014a), Moratelli, Idarraga & Wilson (2015), Tarifa & Aguirre (2009).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis midastactus	Myotis	Unassigned-Myotis	midastactus	Moratelli and Wilson	2014	0	J.Mammal.	95(4): E19	Golden Myotis	None.	Bolivia	Bolivia, Argentina, SE Brazil, Uruguay.	Not listed.	Not Evaluated	Specimens of midastactus were previously identified as Myotis simus ; see Moratelli and Wilson (2014).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis midastactus	23	Golden Myotis	Midas Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	midastactus	Moratelli & D. E. Wilson	2014	0	Myotis_midastactus	Moratelli, R., & Wilson, D. E. (2014). A new species of Myotis (Chiroptera, Vespertilionidae) from Bolivia. Journal of Mammalogy, 95(4), E19.	https://academic.oup.com/jmammal/article/95/4/E17/889615	AMNH 21156		"Cercado, Río Mamoré, Beni, Bolivia, about 23 km W of San Javier (14°34'S, 64°55'W)."	-14.57	-64.92	midastactus Moratelli & D. E. Wilson, 2014	recently described	Moratelli, R., & Wilson, D. E. (2014). A new species of Myotis (Chiroptera, Vespertilionidae) from Bolivia. Journal of Mammalogy, 95(4), E17-E25.	Bolivia|Paraguay	South America	Neotropic	NA	0	0	0	Myotis_midastactus	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136697	Myotis macropus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	macropus	(Gould, 1855)		20000000	Myotis macropus	Least Concern		2021	2020-11-10 00:00:00 UTC	3.1	English	Myotis macropus is assessed as Least Concern because, although the full global distribution of the species is still unclear, it has a wide distribution within Australia, presumed large population, and it is not believed to be declining fast enough to qualify for listing in a threatened category.	This species is highly associated with aquatic habitats ranging from estuaries to streams, rivers, lakes and reservoirs (Richards et al. 2008, Gonsalves and Law 2017). Within this specialised habitat, this species has a patchy distribution and was absent from ecologically diverse riparian forests with abundant surface water (Law and Anderson 1999). This small bat displays morphological adaptations for foraging over water by having long, pointed wings with slightly high aspect ratio and low wing loading (Norberg and Rayner 1987, Jones and Rayner 1991, Campbell 2011), as well as large feet to rake the surface of the water for aquatic prey (Campbell 2011, Law et al. 2020). Dietary analysis identified a diet dominated by aquatic insects and to a lesser extent, fish (Jones and Rayner 1991, Law and Urquhart 2000). The distribution of the species has been affected by heavy metal contamination and sediment pollution (Clarke-Wood et al. 2016, Gonsalves and Law 2017).  Myotis macropus roosts in tree hollows in riparian vegetation located within 100 m of permanent water (Campbell 2009). This species has also been recorded roosting in caves, and in artificial structures like tunnels and bridges also within 100 m of permanent water (Richards et al. 2008, Campbell 2009, Law et al. 2020). This species is found in urban environments where it has been recorded roosting in jetties (Gonsalves and Law 2017) and road culverts (Gorecki et al. 2020). A study on culvert roost availability found these roost types were limited, and this may restrict the species’ distribution and abundance in urban environments, whilst increasing their vulnerability to roost disturbance (Gorecki et al. 2020). Females form maternity colonies from October to March, giving birth to a single pup two to three times per breeding season (Dwyer 1970, Lloyd et al. 1999, Gorecki et al. 2020). Dispersal distances are unknown for this species, however, it has been tracked foraging up to 22 km from roost sites (Campbell 2009). Population structure has been identified between populations 12–15 km apart (Campbell et al. 2009, Law et al. 2020), and female genetic relatedness was not detected beyond 30 km, indicating restricted gene flow between even geographically close populations (VGorecki, unpublished data).	This species is spatially restricted due to its association with aquatic habitats, which are highly susceptible to changes in water quality caused by sedimentation, chemical pollutants, and altered flow regimes (Clarke-Wood et al. 2016, Gonsalves and Law 2017). Changes in water quality may further restrict this species’ distribution through the reduction in suitable foraging grounds. Roosting colonies are threatened by disturbance to roost sites, particularly in culverts and bridges during maternity season. Disturbance to maternity colonies has the potential to impact survival and breeding success. The loss of artificial roost sites has the potential to restrict the species’ distribution and abundance in urban environments due to the limited availability of suitable artificial roosts (Gorecki et al. 2020). Many hollow-bearing trees along riparian corridors were destroyed during the extensive 2019/20 wildfires, and water quality may have been affected.	Population size is expected to be greater than 10,000 mature individuals based on the extent of occurrence although population densities vary across its distribution (Law and Anderson 1999, Campbell 2011). Population numbers are likely to have declined significantly due to long-term drought combined with the extent of riparian areas burnt during the 2019/20 bushfires across a large proportion of its restricted habitat in south-eastern Australia.	Unknown	This species occurs in Australia where it is present in a broad coastal band and along large inland rivers, from South Australia to the Northern Territory (Richards et al. 2008, Churchill 2008). Although the species has a broad geographic distribution, it is uncommon, and has a patchy distribution within this range (Law and Anderson 1999, Campbell 2011). ; ; Populations previously identified as Myotis moluccarum (or called M. adversus ) in Australia, have been shown to be M. macropus although population substructure was identified between northern and southern M. macropus (Cooper et al. 2001). Morphologic and mitochondrial markers indicate this species may be present in Papua New Guinea (Cooper et al. 2001). This species is known from sea level to 840 m, but most records below 300 m in Victoria at least (L. Lumsden pers. comm.).		Terrestrial	The species has been recorded from a number of protected areas. Further studies are needed into the taxonomy, distribution, abundance, and threats to this species, especially the impact of water pollution and severe wildfires. Long-term monitoring is required to determine population trends over time and to quantify the impacts of roost disturbance to urban populations.	Australasian		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	midastactus	Moratelli and Wilson	2014	0	J.Mammal.	95(4): E19	Golden Myotis	None.	Bolivia	Bolivia, Argentina, SE Brazil, Uruguay.	Not listed.	Not Evaluated	Specimens of midastactus were previously identified as Myotis simus ; see Moratelli and Wilson (2014).	Myotis midastactus	1005438	23	Golden Myotis	Midas Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	midastactus	Moratelli & D. E. Wilson	2014	0	Myotis_midastactus	Moratelli, R., & Wilson, D. E. (2014). A new species of Myotis (Chiroptera, Vespertilionidae) from Bolivia. Journal of Mammalogy, 95(4), E19.	https://academic.oup.com/jmammal/article/95/4/E17/889615	AMNH 21156		"Cercado, Río Mamoré, Beni, Bolivia, about 23 km W of San Javier (14°34'S, 64°55'W)."	-14.5667	-64.9167	midastactus Moratelli & D. E. Wilson, 2014	recently described	Moratelli, R., & Wilson, D. E. (2014). A new species of Myotis (Chiroptera, Vespertilionidae) from Bolivia. Journal of Mammalogy, 95(4), E17-E25.				Bolivia|Paraguay	South America	Neotropic	NA	0	0	0	Myotis_midastactus	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_midastactus	1005438	23	Golden Myotis	Midas Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	midastactus	Moratelli & D. E. Wilson	0	Myotis midastactus	Moratelli, R. and Wilson, D.E. 2014-08-22. A new species of _Myotis_ (Chiroptera, Vespertilionidae) from Bolivia. Journal of Mammalogy 95(4):E17-E25.	https://doi.org/10.1644/14-MAMM-149	AMNH M-211156	holotype	http://portal.vertnet.org/o/amnh/mammals?id=urn-catalog-amnh-mammals-m-211156	"Cercado, Río Mamoré, Beni, Bolivia, about 23 km W of San Javier (14°34'S, 64°55'W)."	-14.5667	-64.9167	recently described	Moratelli, R., & Wilson, D. E. (2014). A new species of Myotis (Chiroptera, Vespertilionidae) from Bolivia. Journal of Mammalogy, 95(4), E17-E25.				Bolivia|Paraguay	South America	Neotropic	NE	0	0	0	Myotis_midastactus	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	midastactus	Moratelli & Wilson	2014	0	J.Mammal.	95(4): E19	Golden Myotis	None.	Bolivia	Bolivia, Argentina, SE Brazil, Uruguay.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	Not Evaluated	Specimens of midastactus were previously identified as Myotis simus; see Moratelli and Wilson (2014).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis midastactus; Myotis midastactus; Myotis midastactus; Myotis midastactus; Myotis macropus; Myotis midastactus; midastactus; Murin midas; Goldgelbes Mausohr; Ratonero dorado; Midas Myotis; Golden Myotis; Midas Myotis; Gould's Large-footed Myotis; Golden Myotis; M. midastactus