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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L916	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis	Myotis martiniquensis		[MSW2] Subgenus Selysius.; [MSW3] See Masson and Breuil (1992) and Timm and Genoways (2003).; [HMW] Myotis martiniquensis LaVal, 1973 , 6 km east of La Trinité, Tartane, Martinique , Lesser Antilles. Subgenus Pizonyx; albescens species group. Records from Barbados and Guadeloupe were assigned to M. nyctor and those from Dominica to M. dominicensis . Monotypic.; [batnames2022] Does not include nyctor; see Larsen et al. (2012). See Masson and Breuil (1992) and Timm and Genoways (2003).; [MDD2022] previously included M. nyctor; [IUCN] <span>Myotis martiniquensis previously included M. m. nyctor now raised to species level (Larsen et al. 2012).; [batnames2023] Does not include nyctor; see Larsen et al. (2012). See Masson and Breuil (1992) and Timm and Genoways (2003).; [MDD2023] previously included M. nyctor; [MDD2025_2.0] previously included M. nyctor; [batnames2025_1.7] Does not include nyctor; see Larsen et al. (2012). See Masson and Breuil (1992) and Timm and Genoways (2003).; [MDD2025_2.2] previously included M. nyctor						nyctor.	nyctor, martiniquensis	martiniquensis, nyctor						martiniquensis	<span>Myotis martiniquensis previously included M. m. nyctor now raised to species level (Larsen et al. 2012).			martiniquensis 	martiniquensis 			martiniquensis LaVal, 1973		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Martinique, Lesser Antilles	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis martiniquensis	Martinique (Lesser Antilles), Tartane, 6 km E. La Trinite (France).	LaVal	1973	Bull. Los Ang. Cty. Mus. Nat. Hist. Sci. Soc., 15:35.	Distribution: Confined to the southern Lesser Antilles.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Schwarz's myotis	Martinique, Barbados, Lesser Antilles	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	LaVal	1973	Bull. Los Angeles Cty. Mus. Nat. Hist. Sci. Soc., 15:35.	Subgenus Selysius.	Martinique, Barbados (Lesser Antilles).	Martinique (Lesser Antilles), Tartane, 6 km E La Trinite (France).		LA VAL	1973	Size medium (forearm length, 34-39 mm; condylobasal length, 13-14 mm). Braincase fairly high and occiput not raised above it.	Distribution: Confined to the southern Lesser Antilles.	Two subspecies are recognized:	M. m. nyctor (Barbados), M. m. martiniquensis (Martinique).	105	species	M. martiniquensis	LA VAL	1973	Selysius	subgenus	Myotis martiniquensis				Size medium (forearm length, 34-39 mm; condylobasal length, 13-14 mm). Braincase fairly high and occiput not raised above it.	Two subspecies are recognized:		48. M. martiniquensis LA VAL 1973 [nigricans group].	48	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis martiniquensis	Myotis		martiniquensis	LaVal		1973		Bull. Los Angeles Cty. Mus. Nat. Hist. Sci. Soc.	15		35		Schwartz's Myotis	Martinique (Lesser Antilles), Tartane, 6 km E La Trinité (France).	Martinique, Barbados (Lesser Antilles).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	nyctor LaVal and Schwartz, 1975.	See Masson and Breuil (1992) and Timm and Genoways (2003).	4C3D87E8FF4F6AF1FA489A2A1C27BD8D	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	942	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF4F6AF1FA489A2A1C27BD8D.xml	Myotis martiniquensis	Vespertilionidae	Myotis	martiniquensis	LaVal	1973	Murin de Martinique @fr | Schwartz-Mausohr @de | Ratonero de Schwartz @es	Myotis martiniquensis LaVal, 1973 , 6 km east of La Trinité, Tartane, Martinique , Lesser Antilles. Subgenus Pizonyx; albescens species group. Records from Barbados and Guadeloupe were assigned to M. nyctor and those from Dominica to M. dominicensis . Monotypic.	Martinique I, Lesser Antilles.	Forearm 36-539- 6 mm ; weight 5-8 g . Fur is medium to long (dorsal fur 6-9 mm ; ventral fur 4-6 mm ) and woolly. Ventral hairs are bicolored, with dark brown bases and yellowish brown tips. Bases and tips of dorsal fur contrast weakly, varying from medium to dark brown. Ears are comparatively short, extending forward halfway from eye to nostril. Membranes are dark brown; plagiopatagium is attached to foot at toes by a broad band of membrane; fur on dorsum of uropatagium extends well past knees but not onto tibia; and remainder of membranes is very sparsely furred. Skull is medium-sized (greatest length of skull 14-2-14- 8 mm ); sagittal and lambdoidal crests are generally present, ranging from low to high. P* is aligned in tooth row and visible in labial view. Bacula measurements are 0-65— 0-58 mm long, ¢. 0-29 mm deep, and 0-36- 0-42 mm wide.	Forested habitats.	Schwartz's Myotis is an aerial insectivore that forages in forested habitats and over water.	No information.	Schwartz's Myotis emerges just before sunset. It has been reported roosting in a rocky tunnel, crevice of a large cliff, and crevice below a concrete bridge. [t has also been observed roosting c¢. 5 m aboveground under a leaf of Coccoloba pubescens ( Polygonaceae ) on a steep slope in dry tropical forest in south-western Martinique . A little group roosted under two huge umbrella-like leaves along a mangrove trail in Caravelle Nature Reserve, north-eastern Martinique . Wing morphology and echolocation are typical of aerial insectivore that uses cluttered spaces. Mean call parameters are initial frequency of 94-4 kHz, terminal frequency of 43-9 kHz, frequency of maximum energy of 54-3 kHz, call duration of 4-2 milliseconds, and pulse interval of 82-2 milliseconds.	No information.	Classified as Near Threatened on The IUCN Red List. Schwartz's Myotis has a restricted distribution and faces regular threats from hurricanes and severe weather and habitat decline from expanding tourist facilities.	Barataud etal. (2015) | Dinets (2017b) | Larsen (2016b) | Larsen, Larsen et al. (2012) | LaVal (1973b) | Moratelli et al. (2017) | Simmons (2005)	https://zenodo.org/record/6398789/files/figure.png	408. Schwartz's Myotis Myotis martiniquensis French: Murin de Martinique / German: Schwartz-Mausohr / Spanish: Ratonero de Schwartz Taxonomy. Myotis martiniquensis LaVal, 1973 , 6 km east of La Trinité, Tartane, Martinique , Lesser Antilles. Subgenus Pizonyx; albescens species group. Records from Barbados and Guadeloupe were assigned to M. nyctor and those from Dominica to M. dominicensis . Monotypic. Distribution. Martinique I, Lesser Antilles. Descriptive notes. Forearm 36-539- 6 mm ; weight 5-8 g . Fur is medium to long (dorsal fur 6-9 mm ; ventral fur 4-6 mm ) and woolly. Ventral hairs are bicolored, with dark brown bases and yellowish brown tips. Bases and tips of dorsal fur contrast weakly, varying from medium to dark brown. Ears are comparatively short, extending forward halfway from eye to nostril. Membranes are dark brown; plagiopatagium is attached to foot at toes by a broad band of membrane; fur on dorsum of uropatagium extends well past knees but not onto tibia; and remainder of membranes is very sparsely furred. Skull is medium-sized (greatest length of skull 14-2-14- 8 mm ); sagittal and lambdoidal crests are generally present, ranging from low to high. P* is aligned in tooth row and visible in labial view. Bacula measurements are 0-65— 0-58 mm long, ¢. 0-29 mm deep, and 0-36- 0-42 mm wide. Habitat. Forested habitats. Food and Feeding. Schwartz's Myotis is an aerial insectivore that forages in forested habitats and over water. Breeding. No information. Activity patterns. Schwartz's Myotis emerges just before sunset. It has been reported roosting in a rocky tunnel, crevice of a large cliff, and crevice below a concrete bridge. [t has also been observed roosting c¢. 5 m aboveground under a leaf of Coccoloba pubescens ( Polygonaceae ) on a steep slope in dry tropical forest in south-western Martinique . A little group roosted under two huge umbrella-like leaves along a mangrove trail in Caravelle Nature Reserve, north-eastern Martinique . Wing morphology and echolocation are typical of aerial insectivore that uses cluttered spaces. Mean call parameters are initial frequency of 94-4 kHz, terminal frequency of 43-9 kHz, frequency of maximum energy of 54-3 kHz, call duration of 4-2 milliseconds, and pulse interval of 82-2 milliseconds. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Near Threatened on The IUCN Red List. Schwartz's Myotis has a restricted distribution and faces regular threats from hurricanes and severe weather and habitat decline from expanding tourist facilities. Bibliography. Barataud etal. (2015), Dinets (2017b), Larsen (2016b), Larsen, Larsen et al. (2012), LaVal (1973b), Moratelli et al. (2017), Simmons (2005).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis martiniquensis	Myotis	Unassigned-Myotis	martiniquensis	LaVal	1973	0	Bull. Los Angeles Cty. Mus. Nat. Hist. Sci. Soc.	15:35	Schwartz's Myotis	None.	Martinique (Lesser Antilles), Tartane, 6 km E La Trinité (France).	Guadeloupe, Dominica, Martinique, and Grenada (Lesser Antilles).	Not listed.	Near Threatened	Does not include nyctor; see Larsen et al. (2012). See Masson and Breuil (1992) and Timm and Genoways (2003).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis martiniquensis	23	Schwartz's Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	martiniquensis	LaVal	1973	0	Myotis_martiniquensis	LaVal, R.K. (1973). A revision of the Neotropical bats of the genus Myotis. Science Bulletin of Los Angeles County Museum, 15, 35.		AMNH 214062		6 km east of La Trinité, Tartane, Martinique, Lesser Antilles.			martiniquensis LaVal, 1973	previously included M. nyctor	Larsen, R. J., Larsen, P. A., Genoways, H. H., Catzeflis, F. M., Geluso, K., Kwiecinski, G. G., ... & Baker, R. J. (2012). Evolutionary history of Caribbean species of Myotis, with evidence of a third Lesser Antillean endemic. Mammalian Biology-Zeitschrift für Säugetierkunde, 77(2), 124-134.	Martinique	North America	Neotropic	NT	0	0	0	Myotis_martiniquensis	0	sciname match	Myotis_martiniquensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Myotis martiniquensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	martiniquensis	LaVal, 1973	<span>Myotis martiniquensis previously included M. m. nyctor now raised to species level (Larsen et al. 2012).	20000000	Myotis martiniquensis	Near Threatened		2016	2015-03-16 00:00:00 UTC	3.1	English	This species is listed as Near Threatened as it is close to being threatened based on criterion B1ab(iii). It is only found on four small Caribbean islands of the Lesser Antilles (less than five locations) and the extent of occurrence is c. 27,000 km², with no exact places of record, so this is an estimate. Hurricanes and severe weather are the main threats to island populations, also its habitat is declining due to expansion of tourist facilities.	This species roosts in caves (Timm and Genoways 2003) and it can also be found in anthropogenic habitats (Rodriguez Duran and Kunz 2001). It is insectivorous (Nowak 1999). The little information available on its habitat and ecology is mostly from unpublished studies on the island of Martinique by M. Barataud and collaborators (<a href="http://www.ecologieacoustique.fr/">http://www.ecologieacoustique.fr/)</a>. It shows an apparent fidelity to its hunting areas, being more active at the first two hours after sunset.	Populations might be declining due to habitat loss associate to tourist facilities development and severe weather (hurricanes).	The species is not very common or easily captured.	Unknown	This species occurs on the islands of Martinique, Dominica, Guadeloupe and Barbados (Lesser Antilles) (Larsen et al.  2012, Pedersen et al . 2013).		Terrestrial	Research actions are needed to protect the primary forests this species uses to hunt insects.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	martiniquensis	LaVal	1973	0	Bull. Los Angeles Cty. Mus. Nat. Hist. Sci. Soc.	15:35	Schwartz's Myotis	None.	Martinique (Lesser Antilles), Tartane, 6 km E La Trinité (France).	Guadeloupe, Dominica, Martinique, and Grenada (Lesser Antilles).	Not listed.	Near Threatened	Does not include nyctor; see Larsen et al. (2012). See Masson and Breuil (1992) and Timm and Genoways (2003).	Myotis martiniquensis	1005436	23	Schwartz's Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	martiniquensis	LaVal	1973	0	Myotis_martiniquensis	LaVal, R.K. (1973). A revision of the Neotropical bats of the genus Myotis. Science Bulletin of Los Angeles County Museum, 15, 35.		AMNH 214062		6 km east of La Trinité, Tartane, Martinique, Lesser Antilles.			martiniquensis LaVal, 1973	previously included M. nyctor	Larsen, R. J., Larsen, P. A., Genoways, H. H., Catzeflis, F. M., Geluso, K., Kwiecinski, G. G., ... & Baker, R. J. (2012). Evolutionary history of Caribbean species of Myotis, with evidence of a third Lesser Antillean endemic. Mammalian Biology-Zeitschrift für Säugetierkunde, 77(2), 124-134.				Martinique	North America	Neotropic	NT	0	0	0	Myotis_martiniquensis	0	sciname match	Myotis_martiniquensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_martiniquensis	1005436	23	Schwartz's Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	martiniquensis	LaVal	0	Myotis martiniquensis	LaVal, R.K. 1973-02-14. A revision of the Neotropical bats of the genus _Myotis_. Natural History Museum of Los Angeles County, Science Bulletin 15:1-54.		AMNH M-214062	holotype	http://portal.vertnet.org/o/amnh/mammals?id=urn-catalog-amnh-mammals-m-214062	6 km east of La Trinité, Tartane, Martinique, Lesser Antilles.	14.75	-60.9	previously included M. nyctor	Larsen, R. J., Larsen, P. A., Genoways, H. H., Catzeflis, F. M., Geluso, K., Kwiecinski, G. G., ... & Baker, R. J. (2012). Evolutionary history of Caribbean species of Myotis, with evidence of a third Lesser Antillean endemic. Mammalian Biology-Zeitschrift für Säugetierkunde, 77(2), 124-134.				Martinique	North America	Neotropic	NT	0	0	0	Myotis_martiniquensis	0	sciname match	Myotis_martiniquensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	martiniquensis	LaVal	1973	0	Bull. Los Angeles Cty. Mus. Nat. Hist. Sci. Soc.	15:35	Schwartz's Myotis	None.	Martinique (Lesser Antilles), Tartane, 6 km E La Trinité (France).	Guadeloupe, Dominica, Martinique, and Grenada (Lesser Antilles).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/76435251/22066280/' target='_blank'>Near Threatened</a>	Does not include nyctor; see Larsen et al. (2012). See Masson and Breuil (1992) and Timm and Genoways (2003).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis martiniquensis; Myotis martiniquensis; Myotis martiniquensis; Myotis martiniquensis; Myotis martiniquensis; Myotis martiniquensis; martiniquensis; nyctor; martiniquensis; Murin de Martinique; Schwartz-Mausohr; Ratonero de Schwartz; Schwartz's Myotis; Schwartz's Myotis; Schwartz's Myotis; M. martiniquensis