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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L915	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus	Myotis macrotarsus		[MSW2] Subgenus Leuconoe.; [MSW3] May include stalkeri; see Findley (1972) and Corbet and Hill (1992). Apparently closely related to hasseltii and horsfieldii; see Ruedi and Mayer (2001).; [HMW] of M. macrotarsus should be reviewed. Two subspecies recognized.; [batnames2022] May include stalkeri ; see Findley (1972) and Corbet and Hill (1992). Apparently closely related to hasseltii and horsfieldii ; see Ruedi and Mayer (2001).; [IUCN] The taxonomy of this species should be reviewed (L. Heaney pers. comm. 2006).; [batnames2023] May include stalkeri ; see Findley (1972) and Corbet and Hill (1992). Apparently closely related to hasseltii and horsfieldii ; see Ruedi and Mayer (2001).; [batnames2025_1.7] May include stalkeri; see Findley (1972) and Corbet and Hill (1992). Apparently closely related to hasseltii and horsfieldii; see Ruedi and Mayer (2001).						saba.	saba, macrotarsus	macrotarsus, saba		macrotarsus, saba		macrotarsus, saba		macrotarsus, saba	The taxonomy of this species should be reviewed (L. Heaney pers. comm. 2006).	macrotarsus, saba		macrotarsus, saba 	macrotarsus, saba 	macrotarsus, saba		macrotarsus (G. R. Waterhouse, 1845)|saba D. D. Davis, 1962		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Philippines, Borneo	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis macrotarsus	Philippines.	Waterhouse	1845	Proc. Zool. Soc. Lond., 1845:5.	Distribution: Known only from Borneo and the Philippines.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Philippines, Borneo	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Waterhouse	1845	Proc. Zool. Soc. Lond., 1845:5.	Subgenus Leuconoe.	Philippines, N Borneo.	Philippines.		WATERHOUSE	1845	Size fairly large (forearm length, 45-49 mm; condylobasal length, 16-17 mm). Foot unusually large. Margin of plagiopatagium attached to distal end of tibia. No fringe of hair on margin of uropatagium. Middle upper premolar more or less displaced medially from the toothrow.	Distribution: Known only from Borneo and the Philippines.	Two subspecies:	M. m. saba (Borneo), M. m. macrotarsus (Philippines).	108	species	M. macrotarsus	WATERHOUSE	1845	Leuconoe	subgenus	Myotis macrotarsus				Size fairly large (forearm length, 45-49 mm; condylobasal length, 16-17 mm). Foot unusually large. Margin of plagiopatagium attached to distal end of tibia. No fringe of hair on margin of uropatagium. Middle upper premolar more or less displaced medially from the toothrow.	Two subspecies:		78. M. macrotarsus (WATERHOUSE 1845) [macrotarsus group].	78	_M. m. macrotarsus_ (Waterhouse, 1845); _M. m. saba_ Davis, 1962			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis macrotarsus	Myotis		macrotarsus	Waterhouse	y	1845		Proc. Zool. Soc. Lond.	1845	3	5		Pallid Large-footed Myotis	Philippines.	Philippines, N Borneo.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	saba Davis, 1962.	May include stalkeri; see Findley (1972) and Corbet and Hill (1992). Apparently closely related to hasseltii and horsfieldii; see Ruedi and Mayer (2001).	4C3D87E8FF356A8AFA6297AE18C7B149	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	968	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF356A8AFA6297AE18C7B149.xml	Myotis macrotarsus	Vespertilionidae	Myotis	macrotarsus		1845	Murin des Philippines @fr | Blasse Langful3fledermaus @de | Ratonero de las Filipinas @es | Philippine Large-footed Myotis @en	of M. macrotarsus should be reviewed. Two subspecies recognized.	M.m.macrotarsusWaterhouse,1845—Phil-ippines,includingPalawanandSuluAr-chipelago(Tawi-Tawi).ItprobablyoccursthroughoutthePhilippines. M. m. saba D. D. Davis, 1962 — N Borneo, on Banggi and Belambangan Is and Madai Caves in Sabah . May be more widespread on coastal region of Sabah .	Head—body 55 mm (type), tail 45-59 mm , ear 18-19 mm , hindfoot 14-17 mm , forearm 46-50 mm ; weight 11-16 g . Dorsal hairs are pale grayish brown, with gray bases. Ventral pelage is pale yellow or nearly white. Separation of upperparts and underparts is sharp at the level of wing insertion. Fur is very short, scarcely extending to wing or uropatagium. Wing membranes are pinkish gray. No fringe of hair occurs on margin of uropatagium. Wing attaches above the ankle, not on side of foot. Feet are unusually large, with black claws, contrasting with pale integuments. Ears are slightly longer than head, with long and slender tragus. Subspecies saba from Borneo is bigger than nominate subspecies, and it lacks reddish elements. Horsfield’s Myotis ( M. horsfieldii ) is darker and smaller (forearm 35-38 mm ) and has smaller ( 9-11 mm ) feet, and base of wing attaches at base of foot. Skull is medium-sized (condylo-basal length 16-17 mm ). P? is usually a little intruded into tooth row. Condylo-canine length is 17 mm ; maxillary tooth row length is 7- 3 mm . Chromosomal complement has 2n = 44 and FN = 50 (Negros Island).	Mixed agricultural/regenerating areas and primary lowland forest from sea level up to elevations of ¢. 750 m .	The Pallid Large-footed Myotis possibly trawls for fish and insects over calm waters, as do other congeners with large feet such as Gray Large-footed Myotis ( M. adversus ), Lesser Large-footed Myotis (M. hasseltir), or Southern Myotis ( M. macropus ). Borneo is a center of diversity for fishing bats, with up to five species trawling for insects over freshwaterrivers or saltwater lagoons.	No information.	The Pallid Large-footed Myotis is nocturnal and roosts in caves. It is locally common but dependent on locations with caves and rivers.	On Palawan , Pallid Large-footed Myotis were captured in small numbers in caves.	Classified as Near Threatened on The IUCN Red List. Extent of occurrence of the Pallid Large-footed Myotis is more than 20,000 km 2, but it 1s uncommon and thought to be declining probably at close to 30% over the next 15 years primarily due to ongoing loss of cave habitats. Cave disturbance is a major threat. Better regulations of cave use and comprehensive surveys of populations are needed. It occurs in some protected areas.	Allen (1922) | Amador et al. (2018) | Boitani et al. (2006) | Campbell (2011) | Corbet & Hill (1992) | Dobson (1878) | Esselstyn, Widmann & Heaney (2004) | Findley (1972) | Heaney, Balete, Dolar et al. (1998) | Heaney, Balete & Rickart (2016) | Hollister (1913a) | Koopman (1993, 1994) | MBCFI (2018) | Mudar & Allen (1986) | Nor (1996) | Payne et al. (1985) | Rickart et al. (1999) | Rosell-Ambal & Tabaranza (2008) | Ruedi & Mayer (2001) | Ruedi et al. (2013) | Simmons (2005) | Stadelmann, Jacobs et al. (2004) | Stadelmann, Lin Liangkong et al. (2007) | Tate (1941d) | Taylor (1934)	https://zenodo.org/record/6398934/files/figure.png	469. Pallid Large-footed Myotis Myotis macrotarsus French: Murin des Philippines / German: Blasse Langful3fledermaus / Spanish: Ratonero de las Filipinas Other common names: Philippine Large-footed Myotis Taxonomy. Vespertilio macrotarsus Waterhouse, 1845 , “ Philippine Islands .” Subgenus Myotis ; horsfieldii species group. See M. ridleyi , M. horsfieldii , and M. stalkeri . Taxonomy of M. macrotarsus should be reviewed. Two subspecies recognized. Subspecies and Distribution. M.m.macrotarsusWaterhouse,1845—Phil-ippines,includingPalawanandSuluAr-chipelago(Tawi-Tawi).ItprobablyoccursthroughoutthePhilippines. M. m. saba D. D. Davis, 1962 — N Borneo, on Banggi and Belambangan Is and Madai Caves in Sabah . May be more widespread on coastal region of Sabah . Descriptive notes. Head—body 55 mm (type), tail 45-59 mm , ear 18-19 mm , hindfoot 14-17 mm , forearm 46-50 mm ; weight 11-16 g . Dorsal hairs are pale grayish brown, with gray bases. Ventral pelage is pale yellow or nearly white. Separation of upperparts and underparts is sharp at the level of wing insertion. Fur is very short, scarcely extending to wing or uropatagium. Wing membranes are pinkish gray. No fringe of hair occurs on margin of uropatagium. Wing attaches above the ankle, not on side of foot. Feet are unusually large, with black claws, contrasting with pale integuments. Ears are slightly longer than head, with long and slender tragus. Subspecies saba from Borneo is bigger than nominate subspecies, and it lacks reddish elements. Horsfield’s Myotis ( M. horsfieldii ) is darker and smaller (forearm 35-38 mm ) and has smaller ( 9-11 mm ) feet, and base of wing attaches at base of foot. Skull is medium-sized (condylo-basal length 16-17 mm ). P? is usually a little intruded into tooth row. Condylo-canine length is 17 mm ; maxillary tooth row length is 7- 3 mm . Chromosomal complement has 2n = 44 and FN = 50 (Negros Island). Habitat. Mixed agricultural/regenerating areas and primary lowland forest from sea level up to elevations of ¢. 750 m . Food and Feeding. The Pallid Large-footed Myotis possibly trawls for fish and insects over calm waters, as do other congeners with large feet such as Gray Large-footed Myotis ( M. adversus ), Lesser Large-footed Myotis (M. hasseltir), or Southern Myotis ( M. macropus ). Borneo is a center of diversity for fishing bats, with up to five species trawling for insects over freshwaterrivers or saltwater lagoons. Breeding. No information. Activity patterns. The Pallid Large-footed Myotis is nocturnal and roosts in caves. It is locally common but dependent on locations with caves and rivers. Movements, Home range and Social organization. On Palawan , Pallid Large-footed Myotis were captured in small numbers in caves. Status and Conservation. Classified as Near Threatened on The IUCN Red List. Extent of occurrence of the Pallid Large-footed Myotis is more than 20,000 km 2, but it 1s uncommon and thought to be declining probably at close to 30% over the next 15 years primarily due to ongoing loss of cave habitats. Cave disturbance is a major threat. Better regulations of cave use and comprehensive surveys of populations are needed. It occurs in some protected areas. Bibliography. Allen (1922), Amador et al. (2018), Boitani et al. (2006), Campbell (2011), Corbet & Hill (1992), Dobson (1878), Esselstyn, Widmann & Heaney (2004), Findley (1972), Heaney, Balete, Dolar et al. (1998), Heaney, Balete & Rickart (2016), Hollister (1913a), Koopman (1993, 1994), MBCFI (2018), Mudar & Allen (1986), Nor (1996), Payne et al. (1985), Rickart et al. (1999), Rosell-Ambal & Tabaranza (2008), Ruedi & Mayer (2001), Ruedi et al. (2013), Simmons (2005), Stadelmann, Jacobs et al. (2004), Stadelmann, Lin Liangkong et al. (2007), Tate (1941d), Taylor (1934).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis macrotarsus	Myotis	Unassigned-Myotis	macrotarsus	Waterhouse	1845	1	Proc. Zool. Soc. Lond.	1845(3): 5	Pallid Large-footed Myotis	<b> saba </b>Davis, 1962.	Philippines.	Philippines, N Borneo.	Not listed.	Least Concern	May include stalkeri ; see Findley (1972) and Corbet and Hill (1992). Apparently closely related to hasseltii and horsfieldii ; see Ruedi and Mayer (2001).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis macrotarsus	23	Pallid Large-footed Myotis	Philippine Large-footed Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	macrotarsus	Waterhouse	1845	1						"Philippine Islands."			macrotarsus (Waterhouse, 1845)|saba D. D. Davis, 1962	NA	NA	Philippines|Malaysia	Asia	Indomalaya	LC	0	0	0	Myotis_macrotarsus	0	sciname match	Myotis_macrotarsus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14178	Myotis macrotarsus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	macrotarsus	(Waterhouse, 1845)	The taxonomy of this species should be reviewed (L. Heaney pers. comm. 2006).	20000000	Myotis macrotarsus	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	The species is listed as Least Concerned as it has a moderately broad distribution and a substantial number of locality records across several major islands in the Philippines have been confirmed for the species since the last assessment. Furthermore, the species appears to be highly tolerant of disturbances as most records were from degraded habitats. As such, current available data for the species does not substantiate a 30% decline over the next 15 years.	In the Philippines the species has been recorded roosting in caves near sea level (L. Heaney and R. Utzurrumunpubl. data) and foraging in highly disturbed river habitats near the coast (P. Alviola, Pers. Comm. 2018), over freshwater, rivers, and in agricultural areas. The species has also been recorded in a shallow cave surrounded by brushlands in Pangasinan and in a secondary growth forest in Initao, Misamis Oriental (M.R. Duya, pers. Comm. 2018).	Cave disturbance is a major threat to the species. In the Philippines, caves have been blocked in Misamis Oriental where the species was once abundant (Warguez pers. comm. 2006). On Negros, caves are disturbed by guano miners (L. Heaney pers. comm. 2006) and in other areas they are dammed for water. On Cebu, large numbers of people partake in unregulated spelunking and often open up new areas to the sport.	This is a moderately widespread but uncommon species across its distribution. It roosts in caves or their immediate vicinity (Heaney et al. 1998). On Palawan Island, the species was captured in small numbers in caves (Esselstyn et al. 2004). Current available data for the species does not substantiate a 30% decline over the next 15 years.	Stable	This species occurs from Borneo, Balambangam (Nor 1996) and Banggi islands, to the Philippines. It probably occurs throughout the Philippines, there are records from Bantayan (ROM), Bohol (ROM), Cebu (L. Paguntalan pers. comm. 2006), Guimaras, Luzon (Bulacan, Camarines Sur (FMNH), Quirino (FMNH), Agno, Pangasinan (M.R. Duya, pers. comm. 2018), Rizal Province and Mount Makiling between Batangas and Laguna provinces (N. Ingle pers. comm. 2006)), Marinduque, Mindanao (Lanao del Norte Province, Misamis Oriental (M.R. Duya, pers. Comm. 2018), and Zamboanga Province), Mindoro (MCZ), Negros, Palawan, Polillo, Sicogon (P.G. Jakosalem, pers. comm. 2018), and Tawi-tawi (Hollister 1913, Lawrence 1939, Taylor 1934, Heaney et al. 1998).		Terrestrial	This species occurs in a number of protected areas. Better regulation of cave use is required for this uncommon and cave dependent species, and comprehensive surveys of populations are required.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	macrotarsus	Waterhouse	1845	1	Proc. Zool. Soc. Lond.	1845(3): 5	Pallid Large-footed Myotis	<b> saba </b>Davis, 1962.	Philippines.	Philippines, N Borneo.	Not listed.	Least Concern	May include stalkeri ; see Findley (1972) and Corbet and Hill (1992). Apparently closely related to hasseltii and horsfieldii ; see Ruedi and Mayer (2001).	Myotis macrotarsus	1005435	23	Pallid Large-footed Myotis	Philippine Large-footed Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	macrotarsus	Waterhouse	1845	1						"Philippine Islands."			macrotarsus (Waterhouse, 1845)|saba D. D. Davis, 1962	NA	NA				Philippines|Malaysia	Asia	Indomalaya	LC	0	0	0	Myotis_macrotarsus	0	sciname match	Myotis_macrotarsus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_macrotarsus	1005435	23	Pallid Large-footed Myotis	Philippine Large-footed Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	macrotarsus	G. R. Waterhouse	1	Vespertilio macrotarsus	Waterhouse, G.R. 1845-04. Descriptions of species of Bats collectd in the Philippine Islands, and presented to the Society by H. Cuming, Esq. Proceedings of the Zoological Society of London 1845:3-10.	https://www.biodiversitylibrary.org/page/12862501	BMNH:Mamm:1855.12.6.259	holotype		"Philippine Islands."			NA	NA				Philippines|Malaysia	Asia	Indomalaya	LC	0	0	0	Myotis_macrotarsus	0	sciname match	Myotis_macrotarsus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	macrotarsus	Waterhouse	1845	1	Proc. Zool. Soc. Lond.	1845(3): 5	Pallid Large-footed Myotis	saba Davis, 1962.	Philippines.	Philippines, N Borneo.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14178/22065997/' target='_blank'>Least Concern</a>	May include stalkeri; see Findley (1972) and Corbet and Hill (1992). Apparently closely related to hasseltii and horsfieldii; see Ruedi and Mayer (2001).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis macrotarsus; Myotis macrotarsus; Myotis macrotarsus; Myotis macrotarsus; Myotis macrotarsus; Myotis macrotarsus; macrotarsus; saba; macrotarsus; saba; saba; macrotarsus; saba; Murin des Philippines; Blasse Langful3fledermaus; Ratonero de las Filipinas; Philippine Large-footed Myotis; Pallid Large-footed Myotis; Philippine Large-footed Myotis; Pallid Large-footed Myotis; Pallid Large-footed Myotis; M. macrotarsus