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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L907	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Myotis csorbai	Myotis csorbai	Myotis csorbai and Myotis longipes longipes [synonym of]	Myotis longipes [synonym of]	Myotis csorbai	Myotis longipes longipes [synonym of]	Myotis longipes [synonym of]	Myotis longipes [synonym of]	 	Myotis longipes [synonym of]		[MSW3] Distinct from longipes; see Topál (1997).; [HMW] Myotis csorbai Topal, 1998 , about 30 km S of Pokhara town, 4 km E of Syangja, 1300 m , Syangja District, Nepal . Subgenus Myotis ; siligorensis species group. Myotis csorbai is morphologically very similar to M. longipes , and its sister-group is the capaccinii species group. Records from NE India ( Meghalaya ), S China ( Sichuan , Chongqing , Guizhou , and Guangxi ), and C Laos need additional taxonomic assessment to confirm their identity as M. csorbai . Monotypic.; [batnames2022] Distinct from longipes ; see Topál (1997).; [IUCN] <span lang="HU">Myotis csorbai is distinct from M. longipes (Topál 1997, Simmons 2005).															<span lang="HU">Myotis csorbai is distinct from M. longipes (Topál 1997, Simmons 2005).													N/A																																											Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis csorbai	Myotis		csorbai	Topál		1997		Acta Zool. Acad. Scient. Hungaricae	43	4	377		Csorba's Mouse-eared Myotis	Nepal, Syangja District, 4 km E of Syangja, about 30 km S of Pokhara town, 1,300 m.	Nepal.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Data Deficient.		Distinct from longipes; see Topál (1997).	4C3D87E8FF306A8FFF7794EB1F79B775	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	965	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF306A8FFF7794EB1F79B775.xml	Myotis csorbai	Vespertilionidae	Myotis	csorbai	Topal		Murin de Csorba @fr | Csorba-Langful Rfledermaus @de | Ratonero de Csorba @es | Csorba's Mouse-eared Bat @en | Csorba’s Mouse-eared Myotis @en	Myotis csorbai Topal, 1998 , about 30 km S of Pokhara town, 4 km E of Syangja, 1300 m , Syangja District, Nepal . Subgenus Myotis ; siligorensis species group. Myotis csorbai is morphologically very similar to M. longipes , and its sister-group is the capaccinii species group. Records from NE India ( Meghalaya ), S China ( Sichuan , Chongqing , Guizhou , and Guangxi ), and C Laos need additional taxonomic assessment to confirm their identity as M. csorbai . Monotypic.	Only definitely known from Nepal .	Head-body 37- 3-41 mm , tail 32-42 mm , ear 11-7-16- 4 mm , hindfoot 6-:6-8- 9 mm , forearm 33-7-37- 5 mm ; weight 3-4- 6 g . Csorba’s Myotis is generally similar to the Annamit Myotis (M. annamiticus ) and smaller than the Kashmir Cave Myotis ( M. longipes ). Upperparts of Csorba’s Myotis are dark brownish black; underparts are dull grayish. Ears and membranes are translucent smoky gray. Wing attaches at middle of metatarsus and is sparsely covered with thin hairs on upper side. Calcar is long and thick, without lobe. Uropatagium is hairless. Baculum is less than 0-5 mm long, 0-27 mm in maximum width, and almost triangular with moderate rounded notch at base. Upperside is slightly concaved on basal one-third and shallowly convexed in proximal one-third. Shallow depression occurs on ventral side of bone. Skull is small, with bulbous braincase that is distinctly elevated above flattened rostrum. Sagittal and lambdoid crests are scarcely evident. Zygomata are wide. Short coronoid process of mandible has vertical anterior border and horizontal posterior border. I? and I” are distinct and bicuspidate. C' is weak and about equal to height and ¢.50% of crown area of P*. P?> and P° are minute. P* is one-half the crown area of P% P? slightly displaced internally from tooth row. C, is smaller compared with C'. It is about twothirds the crown area of P,. Molars are characteristically larger. Condylo-canine lengths are 11- 1-12 mm ; maxillary tooth row lengths are 4:6-5- 5 mm .	Subtropical secondary forests at elevations of 1300-1700 m .	Csorba’s Myotis were observed hunting above a river and occasionally the riverbank. Flight typically involves slow circles at 10-30 cm above water surface, rarely above 1-3 m on the riverbank. Flight pattern is similar to that of Daubenton’s Myotis ( M. daubentonii ).	Most female Csorba’s Myotis captured in May had near-term embryos in right uterine horns. One was nulliparous, and some probably gave birth shortly after capture. Difference between average weight of pregnant and non-pregnant females was 1-4 g , suggesting very relatively high neonatal weight.	Csorba’s Myotis is nocturnal and roosts in caves. It was found roosting inside crevices and small holes. Some specimens were collected in a net in front of a cave, surrounded by subtropical secondary forest. Csorba’s Myotis left the cave early before total darkness in summer.	Holotype and four paratypes of Csorba’s Myotis were collected from a colony of ¢.50-60 individuals. Great Himalayan Leat-nosed Bats (Hipposideros armiger) and Schreibers’s Long-fingered Bats ( Miniopterus schreibersii ) also used the Kailash cave.	Classified as Data Deficient on The IUCN Red List. There is no recent information on extent of occurrence, ecological requirements, threats, and conservation status of Csorba’s Myotis . Its habitatis threatened by deforestation from logging, agriculture, and mining. It does not occur in any protected areas. There are public awareness campaigns and local conservation projects targeting the cave where the type series was collected.	Acharya et al. (2010) | Amador et al. (2018) | Bates & Harrison (1997) | Borisenko, Kruskop & Ivanova (2008) | Csorba & Thapa (2016) | Csorba et al. (1999) | Kruskop & Tsytsulina (2001) | Molur et al. (2002) | Pearch (2011) | Ruedi et al. (2013) | Simmons (2005) | Thapa, A. & Thapa (2010) | Thapa, S. (2014) | Thapa, S., Shrestha et al. (2012) | Topal (1998)	https://zenodo.org/record/6398912/files/figure.png	460. Csorba’s Myotis Myotis csorbai French: Murin de Csorba / German: Csorba-LangfulRfledermaus / Spanish: Ratonero de Csorba Other common names: Csorba's Mouse-eared Bat , Csorba’s Mouse-eared Myotis Taxonomy. Myotis csorbai Topal, 1998 , about 30 km S of Pokhara town, 4 km E of Syangja, 1300 m , Syangja District, Nepal . Subgenus Myotis ; siligorensis species group. Myotis csorbai is morphologically very similar to M. longipes , and its sister-group is the capaccinii species group. Records from NE India ( Meghalaya ), S China ( Sichuan , Chongqing , Guizhou , and Guangxi ), and C Laos need additional taxonomic assessment to confirm their identity as M. csorbai . Monotypic. Distribution. Only definitely known from Nepal . Descriptive notes. Head-body 37- 3-41 mm , tail 32-42 mm , ear 11-7-16- 4 mm , hindfoot 6-:6-8- 9 mm , forearm 33-7-37- 5 mm ; weight 3-4- 6 g . Csorba’s Myotis is generally similar to the Annamit Myotis (M. annamiticus ) and smaller than the Kashmir Cave Myotis ( M. longipes ). Upperparts of Csorba’s Myotis are dark brownish black; underparts are dull grayish. Ears and membranes are translucent smoky gray. Wing attaches at middle of metatarsus and is sparsely covered with thin hairs on upper side. Calcar is long and thick, without lobe. Uropatagium is hairless. Baculum is less than 0-5 mm long, 0-27 mm in maximum width, and almost triangular with moderate rounded notch at base. Upperside is slightly concaved on basal one-third and shallowly convexed in proximal one-third. Shallow depression occurs on ventral side of bone. Skull is small, with bulbous braincase that is distinctly elevated above flattened rostrum. Sagittal and lambdoid crests are scarcely evident. Zygomata are wide. Short coronoid process of mandible has vertical anterior border and horizontal posterior border. I? and I” are distinct and bicuspidate. C' is weak and about equal to height and ¢.50% of crown area of P*. P?> and P° are minute. P* is one-half the crown area of P% P? slightly displaced internally from tooth row. C, is smaller compared with C'. It is about twothirds the crown area of P,. Molars are characteristically larger. Condylo-canine lengths are 11- 1-12 mm ; maxillary tooth row lengths are 4:6-5- 5 mm . Habitat. Subtropical secondary forests at elevations of 1300-1700 m . Food and Feeding. Csorba’s Myotis were observed hunting above a river and occasionally the riverbank. Flight typically involves slow circles at 10-30 cm above water surface, rarely above 1-3 m on the riverbank. Flight pattern is similar to that of Daubenton’s Myotis ( M. daubentonii ). Breeding. Most female Csorba’s Myotis captured in May had near-term embryos in right uterine horns. One was nulliparous, and some probably gave birth shortly after capture. Difference between average weight of pregnant and non-pregnant females was 1-4 g , suggesting very relatively high neonatal weight. Activity patterns. Csorba’s Myotis is nocturnal and roosts in caves. It was found roosting inside crevices and small holes. Some specimens were collected in a net in front of a cave, surrounded by subtropical secondary forest. Csorba’s Myotis left the cave early before total darkness in summer. Movements, Home range and Social organization. Holotype and four paratypes of Csorba’s Myotis were collected from a colony of ¢.50-60 individuals. Great Himalayan Leat-nosed Bats (Hipposideros armiger) and Schreibers’s Long-fingered Bats ( Miniopterus schreibersii ) also used the Kailash cave. Status and Conservation. Classified as Data Deficient on The IUCN Red List. There is no recent information on extent of occurrence, ecological requirements, threats, and conservation status of Csorba’s Myotis . Its habitatis threatened by deforestation from logging, agriculture, and mining. It does not occur in any protected areas. There are public awareness campaigns and local conservation projects targeting the cave where the type series was collected. Bibliography. Acharya et al. (2010), Amador et al. (2018), Bates & Harrison (1997), Borisenko, Kruskop & Ivanova (2008), Csorba & Thapa (2016), Csorba et al. (1999), Kruskop & Tsytsulina (2001), Molur et al. (2002), Pearch (2011), Ruedi et al. (2013), Simmons (2005), Thapa, A. & Thapa (2010), Thapa, S. (2014), Thapa, S., Shrestha et al. (2012), Topal (1998).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis csorbai	Myotis	Unassigned-Myotis	csorbai	Top&aacute;l	1997	0	Acta Zool. Acad. Scient. Hungaricae	43(4): 377	Csorba's Mouse-eared Myotis	None.	Nepal, Syangja District, 4 km E of Syangja, about 30 km S of Pokhara town, 1,300 m.	Nepal.	Not listed.	Data Deficient	Distinct from longipes ; see Topál (1997).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2025	synonym of Myotis longipes																																													IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	29420	Myotis csorbai	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	csorbai	Topál, 1997	<span lang="HU">Myotis csorbai is distinct from M. longipes (Topál 1997, Simmons 2005).	20000000	Myotis csorbai	Data Deficient		2016	2015-12-26 00:00:00 UTC	3.1	English	This species is assessed as Data Deficient in view of the absence of recent information on its extent of occurrence, ecological requirements, threats and conservation status.	<p><span lang="HU">The species has been recorded from a cave surrounded by subtropical secondary forest; at other localities it was observed hunting above a river and occasionally the riverbank (Csorba et al . 1999).</p>	<p><span lang="HU">The habitat of this species is under threat from deforestation, generally resulting from logging operations and the conversion of land for agricultural purposes and mining operations (T.K. Shrestha pers. comm. January 2002; Molur et al . 2002).</p>	<p><span lang="HU">The holotype and four more paratypes were collected from a colony of approximately 50-60 individuals (Csorba, pers. comm). Subsequent visits to the very same cave in 2008 and 2009 evidenced the presence of the species; in 2009 at least 15 flying individuals were observed (Thapa, pers. comm).</p>	Unknown	<p><span lang="HU">This species is endemic to Nepal. The type series is originating from four localities around Pokhara (Topál 1997, Csorba et al. 1999), where it has been recorded at elevations between 1,300-1,700 m asl. Specimens collected in different places in the Kathmandu Valley and stored in the collections of Royal Ontario Museum (Pearch 2011) and in the Hokkaido University Natural History Museum are also referable to this species (Csorba pers. comm).</p>		Terrestrial	<p><span lang="HU">This species is not recorded from any protected areas. Surveys, ecological studies, population monitoring, habitat management and protection of the species are recommendations (Molur et al . 2002). There are public awareness campaigns and local conservation projects targeting the cave where the type series was collected (Thapa pers. comm).</p>	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 																																																																		Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393																																																	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586																		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505		Myotis csorbai; Myotis csorbai; Myotis csorbai; Myotis longipes; Myotis csorbai; ; Murin de Csorba; Csorba-Langful Rfledermaus; Ratonero de Csorba; Csorba's Mouse-eared Bat; Csorba’s Mouse-eared Myotis; Csorba's Mouse-eared Myotis;; M. longipes