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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L904	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Myotis levis [synonym of]	Myotis levis dinellii	Myotis levis dinellii	Myotis dinellii	Myotis dinellii	Myotis dinellii	Myotis dinellii	Myotis dinellii	Myotis dinellii	Myotis dinellii	Myotis dinellii	Myotis levis [synonym of]		[HMW] Myotis dinellii Thomas, 1902 , “ Tucuman ,” Tucuman , Argentina . Subgenus Pizonyx; albescens species group. Myotis dinelliiwas considered a subspecies of M. levis , but taxonomic reviews confirmed its status as a distinct specific. Records for southern Brazil were invalidated because specimens were M. albescens . Monotypic.; [batnames2022] Formerly included in M. levis but recognized as distinct by Miranda et al. (2013).; [MDD2022] split from M. levis; [IUCN] It was a subspecies of levis ; Barquez et al. (2006) treat as species.; [batnames2023] Formerly included in M. levis but recognized as distinct by Miranda et al. (2013).; [MDD2023] split from M. levis; [MDD2025_2.0] split from M. levis; [batnames2025_1.7] Formerly included in M. levis but recognized as distinct by Miranda et al. (2013); but see Carrión Bonilla et al. (2024).														dinellii	It was a subspecies of levis ; Barquez et al. (2006) treat as species.			dinellii 	dinellii, dinelli									N/A																																																																			4C3D87E8FF486AF7FA9797F217F3B393	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	941	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF486AF7FA9797F217F3B393.xml	Myotis dinellii	Vespertilionidae	Myotis	dinellii	Thomas		Murin de Dinelli @fr | Dinelli-Mausohr @de | Ratonero de Dinelli @es	Myotis dinellii Thomas, 1902 , “ Tucuman ,” Tucuman , Argentina . Subgenus Pizonyx; albescens species group. Myotis dinelliiwas considered a subspecies of M. levis , but taxonomic reviews confirmed its status as a distinct specific. Records for southern Brazil were invalidated because specimens were M. albescens . Monotypic.	From C Bolivia S to Argentina and extreme E Chile .	Head-body c. 45-55 mm , tail 22-44 mm , ear 7-14 mm , hindfoot 4-11 mm , forearm 34-3-38- 5 mm ; weight 4-8 g . Fur is long (dorsal fur 6-10 mm ; ventral fur 5-8 mm ) and silky. Ventral hairs are bicolored, with black bases, yellowish tips, and strong contrast between bases and tips. Dorsal hairs are bicolored, with black bases, reddish yellow or rufous tips, and strong contrast between bases and tips. Ears are comparatively short, extending forward halfway from eye to nostril. Rostrum, membranes, and ears are black in strong contrast to body color. Plagiopatagium is broadly attached to foot at base oftoes. Fringe of hairs along trailing edge of uropatagium is present but with scattered hairs; upper and lower surfaces of uropatagium are barely covered with hair. Skull is moderate in size (greatest length of skull 13- 8-15 mm ) but quite robust; rostrum is elongated; and postorbital constriction narrow, generally less than 4 mm . Sagittal and lambdoidal crests are generally absent, but when present, they are poorly developed; P* is generally aligned with P* and is visible in profile view. Baculum is rather broad and stout, with average length of 0-65 mm , depth of 0-32 mm , and width of 0-40 mm . Chromosomal complement has 2n = 44 and FN = 50, with three large pairs and one small pair of metacentric and 17 pairs of acrocentric autosomes. Large X-chromosome and small Y-chromosome are submetacentric.	Wide variety of habitats, including deciduousforests, savannas, semiarid open environments, and agricultural landscapes at elevations of 350-3890 m .	Dinelli’s Myotis is an aerial insectivore that forages in open areas and near water bodies.	Two subadult Dinelli’s Myotis were collected in March and September in Tucuman , Argentina .	Dinelli’s Myotis emerges just before sunset. Wing morphology is typical of aerial insectivore that uses cluttered spaces.	A small group of Dinelli’s Myotis roosted with Silver-tipped Myotis ( M. albescens ) and Brazilian Free-tailed Bat (Tadanda brasiliensis ) in an abandoned house in Argentina .	Classified as Least Concern on The IUCN Red List. Dinelli’s Myotisis widespread and presumably has a large population.	Barquez (2006) | Barquez & Diaz (2016e) | Barquez et al. (1999) | Gamboa et al. (2017) | LaVal (1973b) | Miranda et al. (2013) | Sandoval & Barquez (2013) | Thomas (1902c) | Vargas (2007) | Wilson (2008b)	https://zenodo.org/record/6398779/files/figure.png	403. Dinelli’s Myotis Myotis dinellii French: Murin de Dinelli / German: Dinelli-Mausohr / Spanish: Ratonero de Dinelli Taxonomy. Myotis dinellii Thomas, 1902 , “ Tucuman ,” Tucuman , Argentina . Subgenus Pizonyx; albescens species group. Myotis dinelliiwas considered a subspecies of M. levis , but taxonomic reviews confirmed its status as a distinct specific. Records for southern Brazil were invalidated because specimens were M. albescens . Monotypic. Distribution. From C Bolivia S to Argentina and extreme E Chile . Descriptive notes. Head-body c. 45-55 mm , tail 22-44 mm , ear 7-14 mm , hindfoot 4-11 mm , forearm 34-3-38- 5 mm ; weight 4-8 g . Fur is long (dorsal fur 6-10 mm ; ventral fur 5-8 mm ) and silky. Ventral hairs are bicolored, with black bases, yellowish tips, and strong contrast between bases and tips. Dorsal hairs are bicolored, with black bases, reddish yellow or rufous tips, and strong contrast between bases and tips. Ears are comparatively short, extending forward halfway from eye to nostril. Rostrum, membranes, and ears are black in strong contrast to body color. Plagiopatagium is broadly attached to foot at base oftoes. Fringe of hairs along trailing edge of uropatagium is present but with scattered hairs; upper and lower surfaces of uropatagium are barely covered with hair. Skull is moderate in size (greatest length of skull 13- 8-15 mm ) but quite robust; rostrum is elongated; and postorbital constriction narrow, generally less than 4 mm . Sagittal and lambdoidal crests are generally absent, but when present, they are poorly developed; P* is generally aligned with P* and is visible in profile view. Baculum is rather broad and stout, with average length of 0-65 mm , depth of 0-32 mm , and width of 0-40 mm . Chromosomal complement has 2n = 44 and FN = 50, with three large pairs and one small pair of metacentric and 17 pairs of acrocentric autosomes. Large X-chromosome and small Y-chromosome are submetacentric. Habitat. Wide variety of habitats, including deciduousforests, savannas, semiarid open environments, and agricultural landscapes at elevations of 350-3890 m . Food and Feeding. Dinelli’s Myotis is an aerial insectivore that forages in open areas and near water bodies. Breeding. Two subadult Dinelli’s Myotis were collected in March and September in Tucuman , Argentina . Activity patterns. Dinelli’s Myotis emerges just before sunset. Wing morphology is typical of aerial insectivore that uses cluttered spaces. Movements, Home range and Social organization. A small group of Dinelli’s Myotis roosted with Silver-tipped Myotis ( M. albescens ) and Brazilian Free-tailed Bat (Tadanda brasiliensis ) in an abandoned house in Argentina . Status and Conservation. Classified as Least Concern on The IUCN Red List. Dinelli’s Myotisis widespread and presumably has a large population. Bibliography. Barquez (2006), Barquez & Diaz (2016e), Barquez et al. (1999), Gamboa et al. (2017), LaVal (1973b), Miranda et al. (2013), Sandoval & Barquez (2013), Thomas (1902c), Vargas (2007), Wilson (2008b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis dinellii	Myotis	Unassigned-Myotis	dinellii	Thomas	1902	0	Ann. Mag. Nat. Hist.	ser. 7, 10: 493	Dinelli's Myotis	None.		Brazil	Not listed.	Least Concern	Formerly included in M. levis but recognized as distinct by Miranda et al. (2013).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis dinellii	23	Dinelli's Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	dinellii	O. Thomas	1902	0	Myotis_Dinellii	Thomas, O. (1902). On Azara's "Chauve-souris onzième" (Myotis ruber, Geoff.) and a new species allied to it. Annals and Magazine of Natural History, ser. 7, 10, 493.	https://www.biodiversitylibrary.org/item/63688#page/527/mode/1up	BM 1900.7.9.4		"Tucuman," Tucumán, Argentina.			dinellii O. Thomas, 1902	split from M. levis	Miranda, J., Bernardi, I. P., Sponchiado, J., & Passos, F. C. (2013). The taxonomic status of Myotis levis levis and Myotis levis dinellii (Mammalia: Chiroptera: Vespertilionidae). Zoologia (Curitiba), 30(5), 513-518.	Bolivia|Argentina|Chile	South America	Neotropic	LC	0	0	0	Myotis_dinellii	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136204	Myotis dinellii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	dinellii	I. Geoffroy, 1824	It was a subspecies of levis ; Barquez et al. (2006) treat as species.	20000000	Myotis dinellii	Least Concern		2016	2016-07-28 00:00:00 UTC	3.1	English	This species is listed as Least Concern in because of its presumed large population, wide distribution, protection at the distinct levels at several section of its expected distribution, and the fact that no current threats seem to be affecting their populations to put them close to one threatened category.	This is an aerial insectivore. It has been found in all kind of roosts (Barquez, pers. comm.), including man-made constructions (Passos et al . 2010).	None known that could be affecting this species.	The species is common to abundant at part of its geographic range.	Stable	This species is present in Argentina, southern Bolivia and southern Brazil (Barquez et al.  2006, Passos et al. 2010, Miranda et al. 2013), and could be present in eastern Chile (Wilson 2008).		Terrestrial	Its distribution extends over several protected areas, including Biosphere Reserves, national, provincial and natural reserves. Research actions are necessary to understand the differences with M. levis , and the exact limits of its current distribution.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	dinellii	Thomas	1902	0	Ann. Mag. Nat. Hist.	ser. 7, 10: 493	Dinelli's Myotis	None.		Brazil	Not listed.	Least Concern	Formerly included in M. levis but recognized as distinct by Miranda et al. (2013).	Myotis dinellii	1005399	23	Dinelli's Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	dinellii	O. Thomas	1902	0	Myotis_Dinellii	Thomas, O. (1902). On Azara's "Chauve-souris onzième" (Myotis ruber, Geoff.) and a new species allied to it. Annals and Magazine of Natural History, ser. 7, 10, 493.	https://www.biodiversitylibrary.org/item/63688#page/527/mode/1up	BM 1900.7.9.4		"Tucuman," Tucumán, Argentina.			dinellii O. Thomas, 1902	split from M. levis	Miranda, J., Bernardi, I. P., Sponchiado, J., & Passos, F. C. (2013). The taxonomic status of Myotis levis levis and Myotis levis dinellii (Mammalia: Chiroptera: Vespertilionidae). Zoologia (Curitiba), 30(5), 513-518.				Bolivia|Argentina|Chile	South America	Neotropic	LC	0	0	0	Myotis_dinellii	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393																																																	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	dinellii	Thomas	1902	0	Ann. Mag. Nat. Hist.	ser. 7, 10: 493	Dinelli's Myotis	None.		Brazil	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136204/22009702/' target='_blank'>Least Concern</a>	Formerly included in M. levis but recognized as distinct by Miranda et al. (2013); but see Carrión Bonilla et al. (2024).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505		Myotis dinellii; Myotis dinellii; Myotis dinellii; Myotis dinellii; Myotis dinellii; dinellii; Murin de Dinelli; Dinelli-Mausohr; Ratonero de Dinelli; Dinelli's Myotis; Dinelli's Myotis; M. dinellii