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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L901	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Myotis lavali	Myotis lavali	Myotis lavali	Myotis lavali	Myotis lavali	Myotis lavali	Myotis lavali	Myotis lavali	Myotis lavali		[HMW] Myotis lavali Moratelli et al, 2011 , “6.km S of Exu ( 7°30'S , 30°43'W ), Pernambuco State , Brazil , 523 m above sea level.” Subgenus Pizonyx; albescens species group. See M. nigricans . Monotypic.; [MDD2022] recently described; [MDD2023] split from M. nesopolus; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														lavali				larensis	lavali			lavali Moratelli, A. L. Peracchi, Dias, & J. A. de Oliveira, 2011						N/A																																								NA																											4C3D87E8FF4F6AF0FF4092421FF3B09D	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	942	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF4F6AF0FF4092421FF3B09D.xml	Myotis lavali	Vespertilionidae	Myotis	lavali	Moratelli et al.	2011	Murin de La Val @fr | La Val-Mausohr @de | Ratonero de La Val @es	Myotis lavali Moratelli et al, 2011 , “6.km S of Exu ( 7°30'S , 30°43'W ), Pernambuco State , Brazil , 523 m above sea level.” Subgenus Pizonyx; albescens species group. See M. nigricans . Monotypic.	From NE Brazil to Paraguay and NW Argentina .	Head-body c.41 50 mm , tail 31-42 mm , ear 11-14 mm , hindfoot 5-8 mm , forearm 31: 5-37 mm ; weight 3-7 g . Fur is long and silky (dorsal fur ¢. 7 mm ; ventral fur ¢. 6 mm ). Dorsal hairs are strongly bicolored, with medium brown bases (two-thirds the total length) and light-brown tips; ventral hairs are strongly bicolored, with dark brown bases (two-thirds the total length) and cinnamon buff tips. Specimens from Atlantic Forest can have darker dorsal fur and less contrast between bases and tips. Ears are short, extending forward halfway from eye to nostril. Antitragal notch is barely evident. Tragusis pointed, slightly curving outward above and convex below, with small (length 7-9 mm ) triangular lobule at outer base. Plagiopatagium is attached to toes by a broad band of membrane. Fringe of hairs along trailing edge of uropatagium is absent, although some hairs can be present in a few specimens. Membranes are mummy brown. Skull is small to moderate in size (greatest length of skull 13-2-15- 2 mm ), with long and upwardly oriented rostrum; postorbital constriction is narrow; forehead slopes steeply relative to skull; supraoccipital region is rounded; occipital projects beyond posterior limit of occipital condyles; sagittal and lambdoid crests are generally present and low; and P? is generally in tooth row.	Brazilian Caatinga and Cerrado and Paraguayan Alto Chaco ecoregions, predominantly with semiarid and savanna formations, with peripheral records in seasonal semideciduous forests in adjacent Atlantic Forest at elevations of 15-900 m (apparently more frequent at ¢. 350-550 m .	[L.aVal’s Myotis forages in open areas and over water, capturing prey in flight. Diets include various insects, particularly Coleoptera , Hymenoptera , and Lepidoptera .	L.aVal’s Myotis from Brazilian Caatinga breeds year-round, with no peaks of pregnancy or lactation in rainy seasons.	[.aVal’s Myotis emerges just before sunset. In Brazilian Caatinga,its activity 1s concentrated in the first five hours after sunset, and it roosted in horizontal crevices in sandstone rock, small cave in a sandstone plateau with xeric shrub vegetation, and rooftops of abandoned human dwelling. Wing morphology is typical of aerial insectivore that uses cluttered spaces.	In Brazilian Caatinga, LLaVal’s Myotis formed colonies of 16-20 individuals.	Classified as Least Concern on The IUCN Red List. L.aVal’s Myotis is widespread and presumably has a large population.	Barquez et al. (2017) | Moratelli & Wilson (2013) | Moratelli, Peracchi, Dias & Oliveira (2011) | Solari (2017) | Willig (1985a, 1985c)	https://zenodo.org/record/6398783/files/figure.png	405. LaVal's Myotis Myotis lavali French: Murin de LaVal / German: LaVal-Mausohr / Spanish: Ratonero de LaVal Taxonomy. Myotis lavali Moratelli et al, 2011 , “6.km S of Exu ( 7°30'S , 30°43'W ), Pernambuco State , Brazil , 523 m above sea level.” Subgenus Pizonyx; albescens species group. See M. nigricans . Monotypic. Distribution. From NE Brazil to Paraguay and NW Argentina . Descriptive notes. Head-body c.41 50 mm , tail 31-42 mm , ear 11-14 mm , hindfoot 5-8 mm , forearm 31: 5-37 mm ; weight 3-7 g . Fur is long and silky (dorsal fur ¢. 7 mm ; ventral fur ¢. 6 mm ). Dorsal hairs are strongly bicolored, with medium brown bases (two-thirds the total length) and light-brown tips; ventral hairs are strongly bicolored, with dark brown bases (two-thirds the total length) and cinnamon buff tips. Specimens from Atlantic Forest can have darker dorsal fur and less contrast between bases and tips. Ears are short, extending forward halfway from eye to nostril. Antitragal notch is barely evident. Tragusis pointed, slightly curving outward above and convex below, with small (length 7-9 mm ) triangular lobule at outer base. Plagiopatagium is attached to toes by a broad band of membrane. Fringe of hairs along trailing edge of uropatagium is absent, although some hairs can be present in a few specimens. Membranes are mummy brown. Skull is small to moderate in size (greatest length of skull 13-2-15- 2 mm ), with long and upwardly oriented rostrum; postorbital constriction is narrow; forehead slopes steeply relative to skull; supraoccipital region is rounded; occipital projects beyond posterior limit of occipital condyles; sagittal and lambdoid crests are generally present and low; and P? is generally in tooth row. Habitat. Brazilian Caatinga and Cerrado and Paraguayan Alto Chaco ecoregions, predominantly with semiarid and savanna formations, with peripheral records in seasonal semideciduous forests in adjacent Atlantic Forest at elevations of 15-900 m (apparently more frequent at ¢. 350-550 m . Food and Feeding. [L.aVal’s Myotis forages in open areas and over water, capturing prey in flight. Diets include various insects, particularly Coleoptera , Hymenoptera , and Lepidoptera . Breeding. L.aVal’s Myotis from Brazilian Caatinga breeds year-round, with no peaks of pregnancy or lactation in rainy seasons. Activity patterns. [.aVal’s Myotis emerges just before sunset. In Brazilian Caatinga,its activity 1s concentrated in the first five hours after sunset, and it roosted in horizontal crevices in sandstone rock, small cave in a sandstone plateau with xeric shrub vegetation, and rooftops of abandoned human dwelling. Wing morphology is typical of aerial insectivore that uses cluttered spaces. Movements, Home range and Social organization. In Brazilian Caatinga, LLaVal’s Myotis formed colonies of 16-20 individuals. Status and Conservation. Classified as Least Concern on The IUCN Red List. L.aVal’s Myotis is widespread and presumably has a large population. Bibliography. Barquez et al. (2017), Moratelli & Wilson (2013), Moratelli, Peracchi, Dias & Oliveira (2011), Solari (2017), Willig (1985a, 1985c).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis lavali	Myotis	Unassigned-Myotis	lavali	Moratelli, Peracchi, Dias & Oliveira	2011	0	Z. S&auml;ugetierkd.	76(5): 602	LaVal&apos;s Myotis	None.	Brazil, Pernambuco State, 6 km south of Exu.	NE Brazil	Not listed.	Least Concern		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis lavali	23	LaVal's Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	lavali	Moratelli, Peracchi, Dias, & J. A. Oliveira	2011	0	Myotis_lavali	Moratelli, R., Peracchi, A. L., Dias, D., & de Oliveira, J. A. (2011). Geographic variation in South American populations of Myotis nigricans (Schinz, 1821)(Chiroptera, Vespertilionidae), with the description of two new species. Mammalian Biology, 76(5), 602.	https://link.springer.com/article/10.1016/j.mambio.2011.01.003	MZUSP 18762		"6 km S of Exu (7°30'S, 39°43'W), Pernambuco State, Brazil, 523 m above sea level."	-7.5	-39.72	lavali Moratelli, Peracchi, Dias, & J. A. Oliveira, 2011	recently described	Moratelli, R., Peracchi, A. L., Dias, D., & de Oliveira, J. A. (2011). Geographic variation in South American populations of Myotis nigricans (Schinz, 1821)(Chiroptera, Vespertilionidae), with the description of two new species. Mammalian Biology, 76(5), 592-607.	Brazil|Paraguay|Argentina	South America	Neotropic	LC	0	0	0	Myotis_lavali	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Myotis lavali	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	lavali	Moratelli, Peracchi, Dias &; de Oliveira, 2011		90000000	Myotis lavali	Least Concern		2017	2017-03-10 00:00:00 UTC	3.1	English	Myotis lavali is listed as Least Concern because its inferred distribution encompasses a wide region (the Brazilian Caatinga and Cerrado), that is not highly restricted or threatened; protected areas through this geographic range could be effective for its long term survival. Field data suggests this species shows high plasticity and could be locally common. Finally, it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	After a careful review of museum specimens, M. lavali seems to occurs along a diagonal corridor composed of the Brazilian Caatinga and Cerrado and the Paraguayan Alto Chaco, encompassing localities predominantly composed of semiarid (Caatinga and Alto Chaco) and savanna (Cerrado) formations, with a few peripheral records in adjacent Atlantic Forest localities (Moratelli and Wilson 2013). Regarding the reproductive cycle, Willig (1985) found evidence of continuous breeding year-round, with no peaks of pregnancy or lactation detected. The species appears to occur predominantly in semiarid and savanna ecosystems in South America. Despite this data, almost nothing is known about its natural history.	Threats to this species are unknown.	Specimens from Caatinga occur in deciduous, drought-adapted formations and open-forest savanna formations. In most of these localities M. lavali is a common and abundant species, with sexes occurring in similar proportions (Willig 1983, 1985). Almost 30 individuals were caught at the type locality, for other localities between 3 to 5 individuals were recorded at a single sampling event (Moratelli ;et al.  ;2011, Moratelli and Wilson 2013). It seems like the species may be locally common at its dominant habitat (deciduous forests). The species frequently was found roosting in roofs of abandoned buildings, but without forming aggregations of any size (Willig 1983). Records of M. lavali must be examined in a more detailed and accurate geographic scale within these ecosystems to determine habitat preferences (Moratelli and Wilson 2013). Few data exist on natural history of the species thus preventing a complete assessment of its spacial or temporal dynamics.	Unknown	<p>Myotis lavali apparently occurs in a diagonal corridor composed of the Brazilian Caatinga and Cerrado and the Paraguayan Alto Chaco, encompassing localities predominantly composed of semiarid (Caatinga and Alto Chaco) and savanna (Cerrado) formations, with peripheral records in adjacent Atlantic Forest localities in northeastern Brazil and Paraguay (Moratelli ;et al. ;2011, Moratelli and Wilson 2013)</p>		Terrestrial	Only a few, small protected areas exist at the vicinity of the northern localities for the species. However, its distribution may overlap that of the Chapada do Araripe Protection Area, which includes a National Forest. In the central portion of the Caatinga, National Parks and Protection areas may be covering part of the range. Also, in Paraguay, the Gran Chaco is protected by a National Park and a Management Natural area. It can be said that a number of protected areas occur along the entire distribution of the species, securing populations through its geographic range.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	lavali	Moratelli, Peracchi, Dias & Oliveira	2011	0	Z. S&auml;ugetierkd.	76(5): 602	LaVal&apos;s Myotis	None.	Brazil, Pernambuco State, 6 km south of Exu.	NE Brazil	Not listed.	Least Concern		Myotis larensis	1006534	23	Lara Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	larensis	LaVal	1973	0						“Río Tocuyo, Lara, Venezuela”.			larensis LaVal, 1973	split from M. nesopolus	Novaes, R. L. M., Cláudio, V. C., Larsen, R. J., Wilson, D. E., Weksler, M., & Moratelli, R. (2021). The taxonomic status of Myotis nesopolus larensis (Chiroptera, Vespertilionidae) and new insights on the diversity of Caribbean Myotis. ZooKeys, 1015, 145-167.				Colombia|Venezuela	South America	Neotropic	NA	0	0	0	NA	1	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_lavali	1005427	23	LaVal's Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	lavali	Moratelli, A. L. Peracchi, Dias, & J. A. de Oliveira	0	Myotis lavali	Moratelli, R., Peracchi, A.L., Dias, D. and Oliveira, J.A. de. 2011-09. Geographic variation in South American populations of _Myotis nigricans_ (Schinz, 1821) (Chiroptera, Vespertilionidae), with the description of two new species. Mammalian Biology 76(5):592-607.	https://doi.org/10.1016/j.mambio.2011.01.003	MZUSP 18762	holotype		"6 km S of Exu (7°30'S, 39°43'W), Pernambuco State, Brazil, 523 m above sea level."	-7.5	-39.7167	recently described	Moratelli, R., Peracchi, A. L., Dias, D., & de Oliveira, J. A. (2011). Geographic variation in South American populations of Myotis nigricans (Schinz, 1821)(Chiroptera, Vespertilionidae), with the description of two new species. Mammalian Biology, 76(5), 592-607.				Brazil|Paraguay|Argentina	South America	Neotropic	LC	0	0	0	Myotis_lavali	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	lavali	Moratelli, Peracchi, Dias & Oliveira	2011	0	Z. S&auml;ugetierkd.	76(5): 602	LaVal&apos;s Myotis	None.	Brazil, Pernambuco State, 6 km south of Exu.	NE Brazil	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/88151601/88151604/' target='_blank'>Least Concern</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis lavali; Myotis lavali; Myotis lavali; Myotis lavali; Myotis lavali; lavali; Murin de La Val; La Val-Mausohr; Ratonero de La Val; LaVal's Myotis; LaVal&apos;s Myotis; M. lavali