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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L898	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Myotis daubentoni [synonym of]	N/A	Myotis daubentoni [synonym of]	Myotis daubentoni laniger	Myotis laniger	Myotis laniger	Myotis laniger	Myotis laniger	Myotis laniger	Myotis laniger	Myotis laniger	Myotis laniger	Myotis laniger	Myotis laniger		[MSW3] Included in daubentonii by many authors, but see Topál (1997) and Bates et al. (1999).; [HMW] Vespertilio laniger Peters in Swinhoe, 1871 , Amoy, Fujian , China . Subgenus Myotis ; siligorensis species group (7 species). Myotis laniger was formerly included under M. daubentonii but is generally recognized as a distinct species now based on substantial morphological and molecular data. It likely represents a species complex, and comparison of specimens from throughout the distribution is needed. Monotypic.; [batnames2022] Included in daubentonii by many authors, but see Topál (1997) and Bates et al. (1999). Restricted to specimens from Fujian and Henanprovinces in China and Taiwan by Ruedi et al. (2015), but may have a more extensive range into Vietnam and E India.; [MDD2022] includes annamiticus based on low molecular divergence; [batnames2023] Includes annamiticus ; see Ruedi et al. (2021). Included in daubentonii by many authors, but see Topál (1997) and Bates et al. (1999). Restricted to specimens from Fujian and Henan provinces in China and Taiwan by Ruedi et al. (2015), but may have a more extensive range into Vietnam and E India.; [MDD2023] includes annamiticus based on low molecular divergence; [MDD2025_2.0] includes annamiticus based on low molecular divergence; [batnames2025_1.7] Includes annamiticus; see Ruedi et al. (2021). Included in daubentonii by many authors, but see Topál (1997) and Bates et al. (1999). Restricted to specimens from Fujian and Henan provinces in China and Taiwan by Ruedi et al. (2015), but may have a more extensive range into Vietnam and E India.; [MDD2025_2.2] includes annamiticus based on low molecular divergence														laniger, annamiticus		laniger	laniger - annamiticus	laniger, annamiticus	laniger, annamiticus	laniger	laniger - annamitiicus	laniger (W. C. H. Peters in Swinhoe, 1871)|annamiticus Kruskop & K. A. Tsytsulina, 2001						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis laniger	Myotis		laniger	Peters		1870	1871	Proc. Royal Soc. Lond.	3 (1870)		617		Chinese Water Myotis	China, Fujian, Amoy.	S China including Tibet, Vietnam, E India.	IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001).		Included in daubentonii by many authors, but see Topál (1997) and Bates et al. (1999).	4C3D87E8FF316A8FFA5190A81FC6B95B	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	964	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF316A8FFA5190A81FC6B95B.xml	Myotis laniger	Vespertilionidae	Myotis	laniger		1871	Murin laineux @fr | China-Wasserfledermaus @de | Ratonero chino lanudo @es | Chinese Water Bat @en | Chinese Myotis @en | @en | ndochinese Myotis @en	Vespertilio laniger Peters in Swinhoe, 1871 , Amoy, Fujian , China . Subgenus Myotis ; siligorensis species group (7 species). Myotis laniger was formerly included under M. daubentonii but is generally recognized as a distinct species now based on substantial morphological and molecular data. It likely represents a species complex, and comparison of specimens from throughout the distribution is needed. Monotypic.	S & E China ( Hunan , Fujian , and Hainan ), Taiwan , NE India ( Meghalaya ), NE & C Laos , and N Vietnam .	Head-body 36- 3-44 mm , tail 30- 3-40 mm , ear 11- 8-17 mm , hindfoot 7-2-11- 7 mm , forearm 31-37- 1 mm ; weight 3- 3-4 g . The Chinese Water Myotis has dense, woolly fur and distinctly hairy face. Dorsal pelage is grayish brown; venteris creamy whitish, with hair extending onto ventral membranes and limbs up to knees and elbows; there is a distinct color morph that is rufescent dorsally and ventrally. Bare parts of skin around eyesare a lighter fleshy color. Ears are long, narrow, and relatively pointed, with inconspicuous notch along posterior edges; tragus is relatively straight and narrow and is ¢.50% of ear length. Wing membrane attaches close to ankle but is prolonged by narrow strip of membrane to base of metatarsus. Uropatagium stretches from ankle to nearly tip of tail (except last vertebrae, which is free). Calcar is long at three-quarters the distance from ankle to tail tip and lacks lobe or keel. Feet are relatively large and hairy. Penis is club-shaped. Skull profile typically rises sharply after postorbital constriction, while being nearly horizontal above, and is rounded in occipital region; surface of braincase is smooth and lacks sagittal crest; teeth are typically weak; C' barely reach size of P4, C. are even smaller than C!; P? is smaller than P? and in tooth row; inferior teeth are weak; and lower molars are myotodont. Chromosomal complement has on = 48 and FNa = 54 ( China ).	Forested regions near water. Chinese Water Myotis were captured in agricultural areas with small ponds in Cuc Phuong, Vietnam , and disturbed pristine forest in Hoang Lien, Vietnam , at an elevation of ¢. 1850 m .	Chinese Water Myotis are insectivorous and forage low above water. In Guizhou , diet included mostly Diptera and their larvae (78:7%) but also smaller amounts of Coleoptera (6-4%), Lepidoptera (4:3%), and Trichoptera (1-9%).	No information.	Chinese Water Myotis are nocturnal and roost in caves during the day.	Chinese Water Myotis probably roost in small colonies. It was reported roosting in the same cave as the Fringed Longfooted Myotis ( M. fimbriatus ) in mainland China .	Classified as Least Concern on The IUCN Red List. The Chinese Water Myotis is widespread and does not seem to face any major threats, but it is relatively uncommon throughout much of its distribution.	Bates & Harrison (1997) | Bates et al. (1999) | Csorba & Bates (2008a) | Hu Kailiang et al. (2012) | Kruskop (2013a) | Kruskop & Tsytsulina (2001) | Ruedi et al. (2015) | Smith & Xie Yan (2008) | Tu Vuong Tan etal. (2016) | Zhang Weidao (1984)	https://zenodo.org/record/6398910/files/figure.png	459. Chinese Water Myotis Myotis laniger French: Murin laineux / German: China-Wasserfledermaus / Spanish: Ratonero chino lanudo Other common names: Chinese Water Bat , Chinese Myotis , Indochinese Myotis Taxonomy. Vespertilio laniger Peters in Swinhoe, 1871 , Amoy, Fujian , China . Subgenus Myotis ; siligorensis species group (7 species). Myotis laniger was formerly included under M. daubentonii but is generally recognized as a distinct species now based on substantial morphological and molecular data. It likely represents a species complex, and comparison of specimens from throughout the distribution is needed. Monotypic. Distribution. S & E China ( Hunan , Fujian , and Hainan ), Taiwan , NE India ( Meghalaya ), NE & C Laos , and N Vietnam . Descriptive notes. Head-body 36- 3-44 mm , tail 30- 3-40 mm , ear 11- 8-17 mm , hindfoot 7-2-11- 7 mm , forearm 31-37- 1 mm ; weight 3- 3-4 g . The Chinese Water Myotis has dense, woolly fur and distinctly hairy face. Dorsal pelage is grayish brown; venteris creamy whitish, with hair extending onto ventral membranes and limbs up to knees and elbows; there is a distinct color morph that is rufescent dorsally and ventrally. Bare parts of skin around eyesare a lighter fleshy color. Ears are long, narrow, and relatively pointed, with inconspicuous notch along posterior edges; tragus is relatively straight and narrow and is ¢.50% of ear length. Wing membrane attaches close to ankle but is prolonged by narrow strip of membrane to base of metatarsus. Uropatagium stretches from ankle to nearly tip of tail (except last vertebrae, which is free). Calcar is long at three-quarters the distance from ankle to tail tip and lacks lobe or keel. Feet are relatively large and hairy. Penis is club-shaped. Skull profile typically rises sharply after postorbital constriction, while being nearly horizontal above, and is rounded in occipital region; surface of braincase is smooth and lacks sagittal crest; teeth are typically weak; C' barely reach size of P4, C. are even smaller than C!; P? is smaller than P? and in tooth row; inferior teeth are weak; and lower molars are myotodont. Chromosomal complement has on = 48 and FNa = 54 ( China ). Habitat. Forested regions near water. Chinese Water Myotis were captured in agricultural areas with small ponds in Cuc Phuong, Vietnam , and disturbed pristine forest in Hoang Lien, Vietnam , at an elevation of ¢. 1850 m . Food and Feeding. Chinese Water Myotis are insectivorous and forage low above water. In Guizhou , diet included mostly Diptera and their larvae (78:7%) but also smaller amounts of Coleoptera (6-4%), Lepidoptera (4:3%), and Trichoptera (1-9%). Breeding. No information. Activity patterns. Chinese Water Myotis are nocturnal and roost in caves during the day. Movements, Home range and Social organization. Chinese Water Myotis probably roost in small colonies. It was reported roosting in the same cave as the Fringed Longfooted Myotis ( M. fimbriatus ) in mainland China . Status and Conservation. Classified as Least Concern on The IUCN Red List. The Chinese Water Myotis is widespread and does not seem to face any major threats, but it is relatively uncommon throughout much of its distribution. Bibliography. Bates & Harrison (1997), Bates et al. (1999), Csorba & Bates (2008a), Hu Kailiang et al. (2012), Kruskop (2013a), Kruskop & Tsytsulina (2001), Ruedi et al. (2015), Smith & Xie Yan (2008), Tu Vuong Tan etal. (2016), Zhang Weidao (1984).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis laniger	Myotis	Unassigned-Myotis	laniger	Peters	1871	1	Proc. Royal Soc. Lond.	3 (1870): 617	Chinese Water Myotis	None.	China, Fujian, Amoy.	Fujian and Henan provinces (China) and Taiwan, but possibly more widely distributed into Vietnam and E India (e.g., Topal 1997; see Ruedi et al., 2015).	Not listed.	Least Concern	Included in daubentonii by many authors, but see Topál (1997) and Bates et al. (1999). Restricted to specimens from Fujian and Henanprovinces in China and Taiwan by Ruedi et al. (2015), but may have a more extensive range into Vietnam and E India.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis laniger	23	Chinese Water Myotis	Chinese Water Bat|Chinese Myotis|Indochinese Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	laniger	W. Peters	1871	1	Vespertilio_laniger	Peters, W. C. H. (1871). In Swinhoe, R. Catalogue of the mammals of China (south of the River Yangtse) and of the Island of Formosa. Proceedings of the zoological Society of London, 1871 [for 1870], 617.	https://www.biodiversitylibrary.org/item/90543#page/707/mode/1up	ZMB 4146 [lectotype]		Amoy, Fujian, China.			laniger (W. Peters, 1870)|annamiticus Kruskop & Tsytsulina, 2001	includes annamiticus based on low molecular divergence	Ruedi, M., Saikia, U., Thabah, A., Görföl, T., Thapa, S., & Csorba, G. (2021). Molecular and morphological revision of small Myotinae from the Himalayas shed new light on the poorly known genus Submyotodon (Chiroptera: Vespertilionidae). Mammalian Biology, 101(4), 465-480.|Csorba, G., & Furey, N. M. (2022). From greener times: a new species of Thick-thumbed Myotis from Phnom Penh, Cambodia. Acta Zoologica Academiae Scientiarum Hungaricae, 68(1), 85-97.	India|China|Laos|Vietnam|Taiwan	Asia	Palearctic	LC	0	0	0	Myotis_laniger	0	sciname match	Myotis_laniger	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136429	Myotis laniger	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	laniger	Peters, 1871		20000000	Myotis laniger	Least Concern		2019	2018-08-29 00:00:00 UTC	3.1	English	Listed as Least Concern, although there is very little recent information on its extent of occurrence, population status and ecological requirements, it has a very wide range, and its populations are unlikely to be declining fast enough for listing in a more threatened category.	Little is known about the habitat or ecology of this species except that it is a forest dwelling species that may roost in caves and tree hollows (C. Srinivasulu pers. comm., Bates and Harrison 1997, Smith et al . 2008). In China, the species normally was captured at caves (T.Jiang pers. comm. August 2018)	The threats to this species are unknown.	The abundance, population size and trends for this species are not known.	Unknown	This species is widespread in central and south east China (Smith et al. 2008), including Shandong, Jiangsu, Anhui, Zhejiang, Fujiang, Jiangxi, Guangdong, Xianggang, Hainan, Guizhou, Chongqin, Yunnan, Xizang, Sichuang and Shannxi (Jiang et al. 2015), extending to eastern India, where it is known from Meghalaya (Srinivasulu et al. in press), and Viet Nam.		Terrestrial	There are no direct conservation measures in place for this species. It has not been recorded from any protected areas. Further studies are needed into the taxonomy, distribution, abundance, threats and ecology of this species.	Indomalayan|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	laniger	Peters	1871	1	Proc. Royal Soc. Lond.	3 (1870): 617	Chinese Water Myotis	annamitiicus	China, Fujian, Amoy	Fujian, Henan, and Hainan provinces (China) and Taiwan, Vietnam, Meghalaya (India), NE & C Laos, N & C Vietnam	Not listed.	Least Concern	Includes annamiticus ; see Ruedi et al. (2021). Included in daubentonii by many authors, but see Topál (1997) and Bates et al. (1999). Restricted to specimens from Fujian and Henan provinces in China and Taiwan by Ruedi et al. (2015), but may have a more extensive range into Vietnam and E India.	Myotis laniger	1005426	23	Chinese Water Myotis	Chinese Water Bat|Chinese Myotis|Indochinese Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	laniger	W. Peters	1871	1	Vespertilio_laniger	Peters, W. C. H. (1871). In Swinhoe, R. Catalogue of the mammals of China (south of the River Yangtse) and of the Island of Formosa. Proceedings of the zoological Society of London, 1871 [for 1870], 617.	https://www.biodiversitylibrary.org/item/90543#page/707/mode/1up	ZMB 4146 [lectotype]		Amoy, Fujian, China.			laniger (W. Peters, 1870)|annamiticus Kruskop & Tsytsulina, 2001	includes annamiticus based on low molecular divergence	Ruedi, M., Saikia, U., Thabah, A., Görföl, T., Thapa, S., & Csorba, G. (2021). Molecular and morphological revision of small Myotinae from the Himalayas shed new light on the poorly known genus Submyotodon (Chiroptera: Vespertilionidae). Mammalian Biology, 101(4), 465-480.|Csorba, G., & Furey, N. M. (2022). From greener times: a new species of Thick-thumbed Myotis from Phnom Penh, Cambodia. Acta Zoologica Academiae Scientiarum Hungaricae, 68(1), 85-97.				India|China|Laos|Vietnam|Taiwan	Asia	Palearctic	LC	0	0	0	Myotis_laniger	0	sciname match	Myotis_laniger	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_laniger	1005426	23	Chinese Water Myotis	Chinese Water Bat|Chinese Myotis|Indochinese Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	laniger	W. C. H. Peters in Swinhoe	1	Vespertilio laniger	Swinhoe, R. 1871-04. Catalogue of the mammals of China (south of the river Yangtsze) and of the island of Formosa. Proceedings of the Zoological Society of London 1870(3):615-653.	https://www.biodiversitylibrary.org/page/28555479	ZMB 4146	lectotype		Amoy, Fujian, China.			includes annamiticus based on low molecular divergence	Ruedi, M., Saikia, U., Thabah, A., Görföl, T., Thapa, S., & Csorba, G. (2021). Molecular and morphological revision of small Myotinae from the Himalayas shed new light on the poorly known genus Submyotodon (Chiroptera: Vespertilionidae). Mammalian Biology, 101(4), 465-480.|Csorba, G., & Furey, N. M. (2022). From greener times: a new species of Thick-thumbed Myotis from Phnom Penh, Cambodia. Acta Zoologica Academiae Scientiarum Hungaricae, 68(1), 85-97.|Liu, T., Jia, J., Liu, L., Wang, J., Chen, W., Miao, G., ... & Feng, J. (2023). New Insights into the Taxonomy of Myotis Bats in China Based on Morphology and Multilocus Phylogeny. Diversity, 15(7), 805.				India|China|Laos|Vietnam|Taiwan	Asia	Palearctic	LC	0	0	0	Myotis_laniger	0	sciname match	Myotis_laniger	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	laniger	Peters	1871	1	Proc. Royal Soc. Lond.	3 (1870): 617	Chinese Water Myotis	annamitiicus	China, Fujian, Amoy	Fujian, Henan, and Hainan provinces (China) and Taiwan, Vietnam, Meghalaya (India), NE & C Laos, N & C Vietnam	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136429/21984685/' target='_blank'>Least Concern</a>	Includes annamiticus; see Ruedi et al. (2021). Included in daubentonii by many authors, but see Topál (1997) and Bates et al. (1999). Restricted to specimens from Fujian and Henan provinces in China and Taiwan by Ruedi et al. (2015), but may have a more extensive range into Vietnam and E India.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis laniger; Myotis laniger; Myotis laniger; Myotis laniger; Myotis laniger; Myotis laniger; laniger; annamiticus; Murin laineux; China-Wasserfledermaus; Ratonero chino lanudo; Chinese Water Bat; Chinese Myotis; ndochinese Myotis; Chinese Water Myotis; Chinese Water Bat; Chinese Myotis; Indochinese Myotis; Chinese Water Myotis; Chinese Water Myotis; M. laniger