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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L897	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis keeni	Myotis keenii	Myotis keenii	Myotis keenii	Myotis keenii	Myotis keenii	Myotis keenii	Myotis evotis keenii	Myotis keenii	Myotis keenii	Myotis keenii	Myotis keenii	Myotis keenii	Myotis keenii	Myotis keenii		[MSW2] Subgenus Myotis. Includes septentrionalis, possibly a separate species; see Van Zyll de Jong (1979). See Fitch and Shump (1979, Mammalian Species, 121).; [MSW3] Does not include septentrionalis; see van Zyll de Jong (1979) and Caceres and Barclay (2000). See Fitch and Shump (1979), but note that they included septentrionalis.; [HMW] Vespertilio subulatus keenii Merriam, 1895 , “Massett, Queen Charlotte Islands [= Haida Gwaii], British]. C[olumbia].,” Canada . Subgenus Pyzonix; lucifugus species group. See M. evotis . Genetic analyses suggest that M. keenii and M. evotis represent a single species with a morphometric cline that partially explains variation, but this issue remains unresolved. Monotypic.; [IUCN]  Myotis septentrionalis formerly was included in this species. Koopman (1993) included septentrionalis in M. keenii , but van Zyll de Jong (1985), and Simmons (2005) recognized M. keenii and M. septentrionalis as distinct species. Most literature references to M. keenii actually pertain to Myotis septentrionalis . A recent molecular study using mtDNA (cytochrome b gene) (Tanya Dewey, unpublished data) supports the close relationship of M. keenii and M. evotis and their distant relationship to M. septentrionalis . In fact, the molecular phylogeny suggests that M. keenii and M. evotis are conspecific. However, until further studies are completed and published, we here maintain M. keenii and M. evotis as different species.; [batnames2023] Does not include septentrionalis ; see van Zyll de Jong (1979) and Caceres and Barclay (2000). See Fitch and Shump (1979), but note that they included septentrionalis . There remains spirited debate over the status of keenii (see e.g, Nagorsen and Brigham, 1993; Van Zyll de Jong and Nagorsen, 1994; Nagorsen, 2002; Lung et al., 2013; Lausen et al, 2019; Morales et al., 2021; Lausen et al., 2021). For a summary of these issues, and to explain our decision to recognize keenii as a distinct species, see Upham and Francis (2022).; [MDD2023] status was re-affirmed by the Global Bat Taxonomy Working Group (Upham et al. 2022) after being questioned by Lausen et al. (2019), the questioning rebutted by Morales et al. (2021), and the rebuttal rebuted by Lausen et al. (2021). The burden of proof is on overturning the species-level status of Myotis keenii, which has been recognized consistently by taxonomic authorities since 1979 and is thus the prevailing usage in need of refutation.; [MDD2025_2.0] status was re-affirmed by the Global Bat Taxonomy Working Group (Upham et al. 2022) after being questioned by Lausen et al. (2019), the questioning rebutted by Morales et al. (2021), and the rebuttal rebuted by Lausen et al. (2021). The burden of proof is on overturning the species-level status of Myotis keenii, which has been recognized consistently by taxonomic authorities since 1979 and is thus the prevailing usage in need of refutation.; [batnames2025_1.7] Does not include septentrionalis; see van Zyll de Jong (1979) and Caceres and Barclay (2000). See Fitch and Shump (1979), but note that they included septentrionalis. There remains spirited debate over the status of keenii (see e.g, Nagorsen and Brigham, 1993; Van Zyll de Jong and Nagorsen, 1994; Nagorsen, 2002; Lung et al., 2013; Lausen et al, 2019; Morales et al., 2021; Lausen et al., 2021). For a summary of these issues, and to explain our decision to recognize keenii as a distinct species, see Upham and Francis (2022).; [MDD2025_2.2] status was re-affirmed by the Global Bat Taxonomy Working Group (Upham et al. 2022) after being questioned by Lausen et al. (2019), the questioning rebutted by Morales et al. (2021), and the rebuttal rebuted by Lausen et al. (2021). The burden of proof is on overturning the species-level status of Myotis keenii, which has been recognized consistently by taxonomic authorities since 1979 and is thus the prevailing usage in need of refutation.				septentrionalis	(septentrionalis)	septentrionalis.	septentrionalis, keenii							keenii	 Myotis septentrionalis formerly was included in this species. Koopman (1993) included septentrionalis in M. keenii , but van Zyll de Jong (1985), and Simmons (2005) recognized M. keenii and M. septentrionalis as distinct species. Most literature references to M. keenii actually pertain to Myotis septentrionalis . A recent molecular study using mtDNA (cytochrome b gene) (Tanya Dewey, unpublished data) supports the close relationship of M. keenii and M. evotis and their distant relationship to M. septentrionalis . In fact, the molecular phylogeny suggests that M. keenii and M. evotis are conspecific. However, until further studies are completed and published, we here maintain M. keenii and M. evotis as different species.			keenii	keenii, keeni			keenii (C. H. Merriam, 1895)|keeni Corbet & J. Edwards Hill, 1980 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Keen's myotis	Alaska – Washington, Manitoba – Newfoundland – Florida	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis keenii	Canada, British Columbia, Queen Charlotte Isis., Graham Isl., Massett.	Merriam	1895	Am. Nat., 29:860.	Distribution: Ranging across central North America from southeastern Alaska to Newfoundland and south to Florida.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Keen's myotis	Alaska – Washington, Manitoba – Newfoundland – Florida	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Merriam	1895	Am. Nat., 29:860.	Subgenus Myotis. Includes septentrionalis, possibly a separate species; see Van Zyll de Jong (1979). See Fitch and Shump (1979, Mammalian Species, 121).	Alaska Panhandle to W Washington (USA); Mackenzie to Prince Edward Isl (Canada), south to Arkansas and Florida Panhandle (USA).	Canada, British Columbia, Queen Charlotte Isis, Graham Isl, Massett.		MERRIAM	1895	Size medium (forearm length, 34-39 mm; condylobasal length, 13-15 mm). Ear fairly long but not black. Braincase fairly low and rostrum fairly short but relatively narrow. Nasal emargination medium. Middle upper premolar in toothrow. No fringe of hair on margin of uropatagium. Sagittal crest absent or poorly developed.	Distribution: Ranging across central North America from southeastern Alaska to Newfoundland and south to Florida.	Two subspecies (which may be separate species):	M. k. septentrionalis (Mackensie and Newfoundland to Florida), M. k. keenii (Alaska to Washington).	102	species	M. keenii	MERRIAM	1895	Myotis	subgenus	Myotis keenii				Size medium (forearm length, 34-39 mm; condylobasal length, 13-15 mm). Ear fairly long but not black. Braincase fairly low and rostrum fairly short but relatively narrow. Nasal emargination medium. Middle upper premolar in toothrow. No fringe of hair on margin of uropatagium. Sagittal crest absent or poorly developed.	Two subspecies (which may be separate species):		17. M. keenii (MERRIAM 1895) [evotis group].	17	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis keenii	Myotis		keenii	Merriam	y	1895		Am. Nat.	29		860		Keen's Myotis	Canada, British Columbia, Queen Charlotte Isls, Graham Isl, Massett.	Alaska Panhandle to W Washington (USA).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Does not include septentrionalis; see van Zyll de Jong (1979) and Caceres and Barclay (2000). See Fitch and Shump (1979), but note that they included septentrionalis.	4C3D87E8FF5D6AE2FA4A975D166CB2CD	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	928	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF5D6AE2FA4A975D166CB2CD.xml	Myotis keenii	Vespertilionidae	Myotis	keenii		1895	Murin de Keen @fr | Keen-Mausohr @de | Ratonero de Keen @es	Vespertilio subulatus keenii Merriam, 1895 , “Massett, Queen Charlotte Islands [= Haida Gwaii], British]. C[olumbia].,” Canada . Subgenus Pyzonix; lucifugus species group. See M. evotis . Genetic analyses suggest that M. keenii and M. evotis represent a single species with a morphometric cline that partially explains variation, but this issue remains unresolved. Monotypic.	Restricted in coastal Pacific Northwest, from extreme SE Alaska S along British Columbia , including Haida Gwaii and Vancouver I ( Canada ), to NW Washington ( USA ).	Head—-body 40-55 mm , tail 34-44 mm , ear 16-20 mm , hindfoot 8-10 mm , forearm 34- 6-39 mm ; weight 4-6 g . Keen’s Myotis is morphologically distinct from its North American congeners, except the Long-eared Myotis ( M. evotis ) and the Northern Myotis (M. septentrionalis ). Fur is long and silky. Dorsal hairs are bicolored, with blackish bases and golden brown, reddish, or olive-brown tips. Ventral hairs are strongly bicolored, with dark brown bases and pale yellowish tips. Ears are long, extending noticeably beyond tip of nose when laid forward; tragusis long, narrow, and pointed. Membranes and ears are blackish. Plagiopatagium is attached to feet by a broad band of membrane; uropatagium has minute intermittent hairs at its edge. Calcar has indistinct keel. Skull is large (greatest skull lengths 14-6-15- 6 mm ) and slender with steep forehead; sagittal and lambdoidal crests generally are present; and length of upper tooth row slightly exceeds greatest palatal breadth including molar.	Large coastal old growth forests but occasionally anthropogenic habitats. Maternity roosts and summer feeding areas in British Columbia are at elevations below 240 m , and hibernation sites occur above 400 m .	Keen's Myotis is forages in forests and near water. It has been observed catching insects in flight over hot spring pools and clearings above scrubby salal ( Gaultheria shallon, Ericaceae ). It prefers spiders ( Araneae ) and moths ( Lepidoptera ), but other flying insects such as midges, flies and mosquitoes ( Diptera ), and caddisflies (Trichoptera) are also eaten.	Keen's Myotis mates in autumn or early winter before entering hibernation; females store sperm until spring. Upon leaving hibernacula, pregnant females move to maternity roosts in early summer (April-May) where they stay until young are born. Gestation lasts ¢.40-60 days. Females have one offspring per year. Maternity roosts are known only from a hydrothermally heated system of rock crevices and a lodgepole pine ( Pinus contorta, Pinaceae ) snag.	Keen’s Myotis can be active at dusk and dawn. It hibernates during winter and becomes torpid during daytime roosting. It roosts under bark of large trees and in hollow trees, caves, rock crevices, and man-made structures (e.g. bridges and house attics).	Keen’s Myotis make small-scale movements between summer roosts and suitable hibernacula, but they do not make long-distance migrations. They tend to stay relatively close to roosting sites, and estimated daily movements are less than 1 km . They will return to specific foraging areas annually. Maternity roosts have 30-40 females, and hibernacula were reported as the residence for large colonies of individuals, even sharing with other species of bat.	Classified as Least Concern on The IUCN Red List. Keen's Myotis is widespread, presumably has a large population, occurs in some protected areas, and is unlikely to be declining at a rate to qualify for listing in a threatened category.	Fitch & Shump (1979) | Lausen et al. (2019) | Nagorsen & Brigham (1993) | Parker et al. (1997) | Simmons (2005) | Wilson & Ruff (1999) | van Zyll de Jong (1985)	https://zenodo.org/record/6398658/files/figure.png	377. Keen’s Myotis Myotis keenii French: Murin de Keen / German: Keen-Mausohr / Spanish: Ratonero de Keen Taxonomy. Vespertilio subulatus keenii Merriam, 1895 , “Massett, Queen Charlotte Islands [= Haida Gwaii], British]. C[olumbia].,” Canada . Subgenus Pyzonix; lucifugus species group. See M. evotis . Genetic analyses suggest that M. keenii and M. evotis represent a single species with a morphometric cline that partially explains variation, but this issue remains unresolved. Monotypic. Distribution. Restricted in coastal Pacific Northwest, from extreme SE Alaska S along British Columbia , including Haida Gwaii and Vancouver I ( Canada ), to NW Washington ( USA ). Descriptive notes. Head—-body 40-55 mm , tail 34-44 mm , ear 16-20 mm , hindfoot 8-10 mm , forearm 34- 6-39 mm ; weight 4-6 g . Keen’s Myotis is morphologically distinct from its North American congeners, except the Long-eared Myotis ( M. evotis ) and the Northern Myotis (M. septentrionalis ). Fur is long and silky. Dorsal hairs are bicolored, with blackish bases and golden brown, reddish, or olive-brown tips. Ventral hairs are strongly bicolored, with dark brown bases and pale yellowish tips. Ears are long, extending noticeably beyond tip of nose when laid forward; tragusis long, narrow, and pointed. Membranes and ears are blackish. Plagiopatagium is attached to feet by a broad band of membrane; uropatagium has minute intermittent hairs at its edge. Calcar has indistinct keel. Skull is large (greatest skull lengths 14-6-15- 6 mm ) and slender with steep forehead; sagittal and lambdoidal crests generally are present; and length of upper tooth row slightly exceeds greatest palatal breadth including molar. Habitat. Large coastal old growth forests but occasionally anthropogenic habitats. Maternity roosts and summer feeding areas in British Columbia are at elevations below 240 m , and hibernation sites occur above 400 m . Food and Feeding. Keen's Myotis is forages in forests and near water. It has been observed catching insects in flight over hot spring pools and clearings above scrubby salal ( Gaultheria shallon, Ericaceae ). It prefers spiders ( Araneae ) and moths ( Lepidoptera ), but other flying insects such as midges, flies and mosquitoes ( Diptera ), and caddisflies (Trichoptera) are also eaten. Breeding. Keen's Myotis mates in autumn or early winter before entering hibernation; females store sperm until spring. Upon leaving hibernacula, pregnant females move to maternity roosts in early summer (April-May) where they stay until young are born. Gestation lasts ¢.40-60 days. Females have one offspring per year. Maternity roosts are known only from a hydrothermally heated system of rock crevices and a lodgepole pine ( Pinus contorta, Pinaceae ) snag. Activity patterns. Keen’s Myotis can be active at dusk and dawn. It hibernates during winter and becomes torpid during daytime roosting. It roosts under bark of large trees and in hollow trees, caves, rock crevices, and man-made structures (e.g. bridges and house attics). Movements, Home range and Social organization. Keen’s Myotis make small-scale movements between summer roosts and suitable hibernacula, but they do not make long-distance migrations. They tend to stay relatively close to roosting sites, and estimated daily movements are less than 1 km . They will return to specific foraging areas annually. Maternity roosts have 30-40 females, and hibernacula were reported as the residence for large colonies of individuals, even sharing with other species of bat. Status and Conservation. Classified as Least Concern on The IUCN Red List. Keen's Myotis is widespread, presumably has a large population, occurs in some protected areas, and is unlikely to be declining at a rate to qualify for listing in a threatened category. Bibliography. Fitch & Shump (1979), Lausen et al. (2019), Nagorsen & Brigham (1993), Parker et al. (1997), Simmons (2005), Wilson & Ruff (1999), van Zyll de Jong (1985).			Myotis evotis keenii																Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis keenii	23	Keen's Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	keenii	Merriam	1895	1	Vespertilio_subulatus_keenii	Merriam, C. H. (1895). Bats of Queen Charlotte Islands, British Columbia. American Naturalist, 29, 860.	https://www.biodiversitylibrary.org/item/129692#page/276/mode/1up	USNM 72922		"Massett, Queen Charlotte Islands [= Haida Gwaii], B[ritish]. C[olumbia].," Canada.			keenii (Merriam, 1895)	NA	NA	Alaska|Canada|United States	North America	Nearctic	LC	0	0	0	Myotis_keenii	0	sciname match	Myotis_keenii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14171	Myotis keenii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	keenii	(Merriam, 1895)	 Myotis septentrionalis formerly was included in this species. Koopman (1993) included septentrionalis in M. keenii , but van Zyll de Jong (1985), and Simmons (2005) recognized M. keenii and M. septentrionalis as distinct species. Most literature references to M. keenii actually pertain to Myotis septentrionalis . A recent molecular study using mtDNA (cytochrome b gene) (Tanya Dewey, unpublished data) supports the close relationship of M. keenii and M. evotis and their distant relationship to M. septentrionalis . In fact, the molecular phylogeny suggests that M. keenii and M. evotis are conspecific. However, until further studies are completed and published, we here maintain M. keenii and M. evotis as different species.	20000000	Myotis keenii	Least Concern		2017	2016-09-30 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its wide distribution, presumed large population, it occurs at some protected areas, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	The species is known from 25 locations in Canada, where only a couple maternity colonies have been identified (COSEWIC 2003). The only known hibernacula in British Columbia are on northern Vancouver Island where M. keenii has been found in 8 caves from 3 separate areas (COSEWIC 2003). The species is known from just a few locations in southeast Alaska. The distributional range suggests an association with coastal forest habitat (van Zyll de Jong 1985, Nagorsen and Brigham, unpubl. manuscript). Apparently this bat is associated with mature forests, but it is not restricted to old growth (COSEWIC 2003). Across the range it has been found roosting in southwest-facing rock crevices, among geothermally heated rocks, in tree cavities, in bark crevices, and in buildings (D. Burles pers. comm., Nagorsen and Brigham 1993, Parker and Cook 1996). Tree cavities and loose bark are important natural roost sites and may be limiting in some parts of the range (British Columbia Ministry of Water, Land and Air Protection 2004). In British Columbia, one maternity colony (on Hot Springs Island in the Queen Charlotte Islands) is situated within geothermally heated rocks associated with hot spring activity (British Columbia Ministry of Water, Land and Air Protection 2004). The only other known maternity colony in British Columbia was suspected to be in a tree located in a low elevation, southwest-facing cliff at Knoll Hill near Tahsis, Vancouver Island (COSEWIC 2003). Known maternity roosts and summer feeding areas in British Columbia are at elevations below 240 meters, known hibernation sites occur above 400 meters in caves over 100 meters long (British Columbia Ministry of Water, Land and Air Protection 2004). These bats have been observed foraging over hot spring pools and clearings above scrubby salal (Gaultheria shallon ).	Habitat quantity and quality are presumed to be declining (British Columbia Ministry of Water, Land and Air Protection 2004), but the rate of decline is unknown. Population estimates made at a British Columbia maternity colony during the 1990s indicate that it remained stable between 1991 and 2000 (COSEWIC 2003). Logging activities and associated road building likely are reducing and fragmenting habitat, mineral extraction, forest fires, and pesticides also may be detrimental (Balcombe, 1988 COSEWIC report, British Columbia Ministry of Water, Land and Air Protection 2004). Disturbance during hibernation and while raising young is a major concern, disturbance may result from recreational activities (e.g., caving) or industrial activities (e.g., blasting for road construction) (British Columbia Ministry of Water, Land and Air Protection 2004). Current levels of timber harvest could have a detrimental effect on the Alaska population by altering forest structure important to bats (Parker et al.  1997). Bat activity is rare in clearcuts and second-growth forests of Southeast Alaska (Parker and Cook 1997, Parker et al . 1997).	Larger groups of this species form colonies to ;<a>hibernate</a> ;together in the winter, sometimes in the same ;<a>hibernaculum</a> ;as other ;<a>species</a> ;of bats. ;Total adult population size is unknown but presumably is at least several thousand. A maternity colony in British Columbia included about 40 reproductive females (COSEWIC 2003).	Stable	The range includes coastal Washington, British Columbia (including Vancouver Island, the Queen Charlotte Islands, and coastal mainland), and southeastern Alaska (e.g., Hoonah, Wrangell, Ketchikan) (Parker and Cook 1996, Simmons 2005). This species has one of the smallest distributional ranges of any North American bat (Van Zyll de Jong 1985).		Terrestrial	A maternity colony in British Columbia is protected within Gwaii Haanas National Park Reserve. Weymer Cave and White Ridge Provincial Parks protect areas that support M. keenii hibernacula and roosting habitat. The former park has a draft management plan that identifies the need to protect the bat roosts, but also identifies recreational caving, hiking, and tourism as acceptable park activities (COSEWIC 2003). Range-wide population surveys are needed. Coordinated surveys should be conducted in Alaska, British Columbia, and Washington. See British Columbia Ministry of Water, Land and Air Protection (2004) for specific protection needs and management recommendations. This species is known only from a limited number of museum specimens. Virtually nothing is known about its biology (van Zyll de Jong 1985). Present research needs include the following: 1. Assess habitat preferences, including roosting and feeding areas, activity patterns, and food requirements. 2. Determine if there is any seasonal movement within the population. 3. Assessing whether this species hibernates, and if so, where and under what conditions. 4. Research the reproductive capacity of the species and evaluate its long-term population viability.	Nearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	keenii	Merriam	1895	1	Am. Nat.	43:20:00	Keen's Myotis	None.	Canada, British Columbia, Queen Charlotte Islands [= Haida Gwaii], Massett	United States (SE Alaska and NW Washington) and British Columbia (including Haida Gwaii and Vancouver Isls; Canada)	Not listed.	Least Concern	Does not include septentrionalis ; see van Zyll de Jong (1979) and Caceres and Barclay (2000). See Fitch and Shump (1979), but note that they included septentrionalis . There remains spirited debate over the status of keenii (see e.g, Nagorsen and Brigham, 1993; Van Zyll de Jong and Nagorsen, 1994; Nagorsen, 2002; Lung et al., 2013; Lausen et al, 2019; Morales et al., 2021; Lausen et al., 2021). For a summary of these issues, and to explain our decision to recognize keenii as a distinct species, see Upham and Francis (2022).	Myotis keenii	1005425	23	Keen's Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	keenii	Merriam	1895	1	Vespertilio_subulatus_keenii	Merriam, C. H. (1895). Bats of Queen Charlotte Islands, British Columbia. American Naturalist, 29, 860.	https://www.biodiversitylibrary.org/item/129692#page/276/mode/1up	USNM 72922		"Massett, Queen Charlotte Islands [= Haida Gwaii], B[ritish]. C[olumbia].," Canada.			keenii (Merriam, 1895)	status was re-affirmed by the Global Bat Taxonomy Working Group (Upham et al. 2022) after being questioned by Lausen et al. (2019), the questioning rebutted by Morales et al. (2021), and the rebuttal rebuted by Lausen et al. (2021). The burden of proof is on overturning the species-level status of Myotis keenii, which has been recognized consistently by taxonomic authorities since 1979 and is thus the prevailing usage in need of refutation.	Upham, N. S. and Francis, C. M. on behalf of the G. B. T. W. G. of the IUCN SSC Bat Specialist Group (2022). On the taxonomy of Myotis keenii and Myotis evotis. https://doi.org/10.5281/zenodo.7336341|Lausen, C. L., Proctor, M., Nagorsen, D. W., Burles, D., Paetkau, D., Harmston, E., Blejwas, K., Govindarajulu, P., & Friis, L. (2019). Population genetics reveal Myotis keenii (Keen’s myotis) and Myotis evotis (long-eared myotis) to be a single species. Canadian Journal of Zoology, 97(3), 267–279. https://doi.org/10.1139/cjz-2018-0113|Morales, A. E., Fenton, M. B., Carstens, B. C., & Simmons, N. B. (2021). Comment on “Population genetics reveal Myotis keenii (Keen’s myotis) and Myotis evotis (long-eared myotis) to be a single species.” Canadian Journal of Zoology, 99(5), 415–422. https://doi.org/10.1139/cjz-2020-0048|Lausen, C. L., Proctor, M. F., Paetkau, D., Nagorsen, D. W., Govindarajulu, P., Burles, D., & Blejwas, K. (2021). Reply to the comment by Morales et al. On “Population genetics reveal Myotis keenii (Keen’s myotis) and Myotis evotis (long-eared myotis) to be a single species.” Canadian Journal of Zoology, 99(5), 423–432. https://doi.org/10.1139/cjz-2020-0266			USA(AK,WA)	Alaska|Canada|United States	North America	Nearctic	LC	0	0	0	Myotis_keenii	0	sciname match	Myotis_keenii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_keenii	1005425	23	Keen's Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	keenii	C. H. Merriam	1	Vespertilio subulatus keenii	Merriam, C.H. 1895-09-01. Bats of Queen Charlotte Islands, British Columbia. The American Naturalist 29(345):860-861.	https://www.biodiversitylibrary.org/page/41521071	USNM:MAMM:72922	holotype	http://n2t.net/ark:/65665/3001b129d-06f0-4aa6-abd2-b2abcee5ea66	"Massett, Queen Charlotte Islands [= Haida Gwaii], B[ritish]. C[olumbia].," Canada.			status was re-affirmed by the Global Bat Taxonomy Working Group (Upham et al. 2022) after being questioned by Lausen et al. (2019), the questioning rebutted by Morales et al. (2021), and the rebuttal rebuted by Lausen et al. (2021). The burden of proof is on overturning the species-level status of Myotis keenii, which has been recognized consistently by taxonomic authorities since 1979 and is thus the prevailing usage in need of refutation.	Upham, N. S. and Francis, C. M. on behalf of the G. B. T. W. G. of the IUCN SSC Bat Specialist Group (2022). On the taxonomy of Myotis keenii and Myotis evotis. https://doi.org/10.5281/zenodo.7336341|Lausen, C. L., Proctor, M., Nagorsen, D. W., Burles, D., Paetkau, D., Harmston, E., Blejwas, K., Govindarajulu, P., & Friis, L. (2019). Population genetics reveal Myotis keenii (Keen’s myotis) and Myotis evotis (long-eared myotis) to be a single species. Canadian Journal of Zoology, 97(3), 267–279. https://doi.org/10.1139/cjz-2018-0113|Morales, A. E., Fenton, M. B., Carstens, B. C., & Simmons, N. B. (2021). Comment on “Population genetics reveal Myotis keenii (Keen’s myotis) and Myotis evotis (long-eared myotis) to be a single species.” Canadian Journal of Zoology, 99(5), 415–422. https://doi.org/10.1139/cjz-2020-0048|Lausen, C. L., Proctor, M. F., Paetkau, D., Nagorsen, D. W., Govindarajulu, P., Burles, D., & Blejwas, K. (2021). Reply to the comment by Morales et al. On “Population genetics reveal Myotis keenii (Keen’s myotis) and Myotis evotis (long-eared myotis) to be a single species.” Canadian Journal of Zoology, 99(5), 423–432. https://doi.org/10.1139/cjz-2020-0266			USA(AK,WA)	Canada|United States	North America	Nearctic	LC	0	0	0	Myotis_keenii	0	sciname match	Myotis_keenii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	keenii	Merriam	1895	1	Am. Nat.	43:20:00	Keen's Myotis	None.	Canada, British Columbia, Queen Charlotte Islands [= Haida Gwaii], Massett	United States (SE Alaska and NW Washington) and British Columbia (including Haida Gwaii and Vancouver Isls; Canada)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14171/22055579/' target='_blank'>Least Concern</a>	Does not include septentrionalis; see van Zyll de Jong (1979) and Caceres and Barclay (2000). See Fitch and Shump (1979), but note that they included septentrionalis. There remains spirited debate over the status of keenii (see e.g, Nagorsen and Brigham, 1993; Van Zyll de Jong and Nagorsen, 1994; Nagorsen, 2002; Lung et al., 2013; Lausen et al, 2019; Morales et al., 2021; Lausen et al., 2021). For a summary of these issues, and to explain our decision to recognize keenii as a distinct species, see Upham and Francis (2022).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis keenii; Myotis keenii; Myotis evotis keenii; Myotis keenii; Myotis keenii; Myotis keenii; keenii; Murin de Keen; Keen-Mausohr; Ratonero de Keen; Keen's Myotis; Keen's Myotis; Keen's Myotis; M. keenii