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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L889	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi	Myotis ikonnikovi		[MSW2] Subgenus Selysius. Probably a subspecies of muricola; see Corbet (1978c:48), who rejected European records.; [MSW3] Revised by Tsytsulina (2001); also see Corbet (1978c), Yoshiyuki (1989), and Benda and Tsytsulina (2000). Molecular sequence data support placement of fujiensis in ikonnikovi (Kawai et al., 2003).; [HMW] Myotis tkonnikovi Ognev, 1912 , Euseevka, Dalnerechen District, Primorsky Krai (= Ussuri Region), Russia . Subgenus Myotis ; mystacinus species group (4 species), which may besister to muricola species group. This species appears to be sister to M. altarium . Previously treated as a race of M. muricola , but split on genetic and morphological grounds. Monotypic.; [batnames2022] Includes hosonoi,  ozensis and yesoensis; see Abe et al. (2005). Revised by Tsytsulina (2001); also see Corbet (1978 c ), Yoshiyuki (1989), and Benda and Tsytsulina (2000). Molecular sequence data support placement of fujiensis in ikonnikovi (Kawai et al., 2003).; [MDD2022] includes hosonoi, ozensis, and yesoensis; [IUCN] Belongs to the group “mystacinus ”. Not conspecific with S Asian M. muricola Gray, 1846 (Tsytsulina 2001); includes Japanese fujiensis Imaizumi, 1954 (Kawai et al. 2003).; [batnames2023] Includes hosonoi,  ozensis and yesoensis; see Abe et al. (2005). Revised by Tsytsulina (2001); also see Corbet (1978 c ), Yoshiyuki (1989), and Benda and Tsytsulina (2000). Molecular sequence data support placement of fujiensis in ikonnikovi (Kawai et al., 2003).; [MDD2023] includes hosonoi, ozensis, and yesoensis; [MDD2025_2.0] includes hosonoi, ozensis, and yesoensis; [batnames2025_1.7] Includes hosonoi, ozensis and yesoensis; see Abe et al. (2005). Revised by Tsytsulina (2001); also see Corbet (1978c), Yoshiyuki (1989), and Benda and Tsytsulina (2000). Molecular sequence data support placement of fujiensis in ikonnikovi (Kawai et al., 2003).; [MDD2025_2.2] includes hosonoi, ozensis, and yesoensis									fujiensis			ikonnikovi 	ikonnikovi - fujiensis, hosonoi, ozensis, yesoensis	ikonnikovi, fujiensis, hosonoi, ozensis, yesoensis	Belongs to the group “mystacinus ”. Not conspecific with S Asian M. muricola Gray, 1846 (Tsytsulina 2001); includes Japanese fujiensis Imaizumi, 1954 (Kawai et al. 2003).	ikonnikovi 	ikonnikovi - fujiensis, hosonoi, ozensis, yesoensis	ikonnikovi, fujiensis, hosonoi, ozensis, yesoensis	ikonnikovi, fujiensis, hosonoi, ozensis, yesoensis	ikonnikovi	ikonnikovi - fujiensis, hosonoi, ozensis, yesoensis	ikonnikovi Ognev, 1912|fujiensis Imaizumi, 1954|hosonoi Imaizumi, 1954|ozensis Imaizumi, 1954|yesoensis Yoshiyuki, 1984|oxensis Corbet & J. Edwards Hill, 1991 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		E Siberia, N Korea, Sakhalin, Hokkaido	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis ikonnikovi	U.S.S.R., Primorsk. Krai (= Ussuri Region), Iman Dist.	Ognev	1912	Ann. Mus. Zool. Acad. Sci. St. Petersb., 16:477.	Distribution: Ranging from southeastern Siberia to Mongolia and Korea, also Sakhalin and Hokkaido in Japan.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		E Siberia, N Korea, Sakhalin, Hokkaido	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Ognev	1912	Ann. Mus. Zool. Acad. Imp. Sci. St. Petersbourg, 16:477.	Subgenus Selysius. Probably a subspecies of muricola; see Corbet (1978c:48), who rejected European records.	Ussuri region and N Korea to Lake Baikal (Russia), the Altai Mtns, and Mongolia, NE China; Sakhalin Isl (Russia) and Hokkaido Isl (Japan).	Russia, Primorsk. Krai (= Ussuri Region), Dalnerechen Dist., Euseevka.		OGNEV	1912	Size small (forearm length, 32-35 mm). Keel on calcar poorly developed. Tail relatively long but extending to edge of uropatagium. Braincase fairly high. Middle upper premolar in toothrow. Posterior border of ear pinna with a distinct emargination, tragus relatively straight.	Distribution: Ranging from southeastern Siberia to Mongolia and Korea, also Sakhalin and Hokkaido in Japan.	No subspecies. May be a subspecies of M. muricola.		103	species	M. ikonnikovi	OGNEV	1912	Selysius	subgenus	Myotis ikonnikovi				Size small (forearm length, 32-35 mm). Keel on calcar poorly developed. Tail relatively long but extending to edge of uropatagium. Braincase fairly high. Middle upper premolar in toothrow. Posterior border of ear pinna with a distinct emargination, tragus relatively straight.	No subspecies. May be a subspecies of M. muricola.		28. M. ikonnikovi OGNEV 1912 [muricola group].	28	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis ikonnikovi	Myotis		ikonnikovi	Ognev		1912		Ann. Mus. Zool. Acad. Imp. Sci. St. Petersbourg	16		477		Ikonnikov's Myotis	Russia, Primorsk. Krai (= Ussuri Region), Dalnerechen Dist., Euseevka.	Ussuri region and N Korea to Lake Baikal (Russia), the Altai Mtns, and Mongolia, NE China; Sakhalin Isl (Russia) and Honshû and Hokkaido Isls (Japan).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	fujiensis Imaizumi, 1954.	Revised by Tsytsulina (2001); also see Corbet (1978c), Yoshiyuki (1989), and Benda and Tsytsulina (2000). Molecular sequence data support placement of fujiensis in ikonnikovi (Kawai et al., 2003).	4C3D87E8FF3A6A85FA8492171909B6A6	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	955	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF3A6A85FA8492171909B6A6.xml	Myotis ikonnikovi	Vespertilionidae	Myotis	ikonnikovi	Ognev	1912	Murin d'lkonnikov @fr | Kurzohr-Bartfledermaus @de | Ratonero de lkonnikov @es | lkonnikov's Bat @en | lkonnikov's Whiskered Bat @en	Myotis tkonnikovi Ognev, 1912 , Euseevka, Dalnerechen District, Primorsky Krai (= Ussuri Region), Russia . Subgenus Myotis ; mystacinus species group (4 species), which may besister to muricola species group. This species appears to be sister to M. altarium . Previously treated as a race of M. muricola , but split on genetic and morphological grounds. Monotypic.		Head-body 36-55 mm , tail 29-41 mm , ear 8-15 mm , hindfoot 6-9- 5 mm , forearm 30-36 mm . Dorsal pelage of Ikonnikov’s Myotis is uniformly dark brown; ventral pelage slightly paler with slightly darker patch on each side of chest. Bare portions of ears, face, and membranes are dark grayish brown. Ears are relatively long and rather slender with narrowly rounded tip, shallow concavity on posterior borderjust below tip, outward-turned anterior border in middle, and are lightly furred on the inner surface; tragus is just over half ear-height and is slightly concave or straight on anterior margin and convex on posterior margin with pointed tip. Wings are attached to base of outer toe, and uropatagium lacks fringe of hairs on posterior border; calcar is keeled. Skull is small and has pointed rostrum; braincase is broad and low with abruptly rising forehead; interorbital breadth is greater than distance between canines. P? is slightly smaller than P? and is within tooth row, and not leaning inward, P,is small and within tooth row. Chromosomal complement has 2n = 44 and FNa = 52.	Ikonnikov’s Myotis is mainly found in mountain forests at elevations of 600— 1000 m , although it has been recorded as low as 200 m in Aomori . Known from most forest types throughoutits range and also found in agricultural land.	Seen foraging for aerial insects low along forest roads or streams.	Births are known to occur from late June to late July; lactating females and juveniles recorded from late July to early August. Females give birth to single young.	Ikonnikov’s Myotis roosts by day under bark,in tree cavities, between lianas on tree trunks, in old buildings, under bridges, and in farm roofs, rarely tunnels or other subterranean habitats. During the day, it enters torpid state at its roosts, leaving in the evening to forage through the night. It hibernates through winter in holes or crevices inside caves. Calls are a steep FM sweep with an average start frequency of 90 kHz (55-2-118-1 kHz), end frequency 43-2 kHz (37-8-50-1 kHz), peak frequency 50-6 kHz (47-3-55-9 kHz), and duration 2 milliseconds (0-9-3-4 milliseconds) recorded in Hokkaido , Japan . Recordings from South Korea had average start frequencies of 83-4-132-2 kHz, end frequencies 26-6-44-2 kHz, peak frequencies 46-7-62-6 kHz, interpulse intervals 53-121-3 milliseconds, and durations 2:6—4 milliseconds.	In summer, Ikonnikov’s Myotis forms maternity colonies ranging in size from tens to over a hundred. Ikonnikov’s Myotis shares roosts with Greater Japanese Horseshoe Bats (Rhinolophus nippon), Far Eastern Myotis (M. bombinus ), and Big-footed Myotis ( M. macrodactylus ). While hibernating however, it usually roost alone. It switches roosts every 1-2 days and may return to a roost several times. Radio-telemetry shows thatit flies 69-360 m each night while foraging.	Classified as Least Concern on The IUCN Red List. Tkonnikov’s Myotis is apparently widespread, with no major threats known.	Abe et al. (2005) | Bannikov (1954) | Findley (1972) | Fukui , Agetsuma & Hill (2004) | Fukui , Hill et al. (2015) | Harada & Yoshida (1978) | Jo Yeong-Seok et al. (2018) | Kawai, Kondo et al. (2006) | Kawai, Nikaido et al. (2003) | Kondo & Sasaki (2005) | Ohdachi et al. (2009) | Ono & Obara (1994) | Smith & Xie Yan (2008) | Stubbe, Ariunbold, Buuveibaatar, Dorjderem, Monkhzul, Otgonbaatar & Tsogbadrakh (2008b) | Tsytsulina (2001) | Tsytsulina et al. (2012) | Yasui et al. (2004)	https://zenodo.org/record/6398861/files/figure.png	438. Ikonnikov’'s Myotis Myotis ikonnikovi French: Murin d'lkonnikov / German: Kurzohr-Bartfledermaus / Spanish: Ratonero de lkonnikov Other common names: lkonnikov's Bat , lkonnikov's Whiskered Bat Taxonomy. Myotis tkonnikovi Ognev, 1912 , Euseevka, Dalnerechen District, Primorsky Krai (= Ussuri Region), Russia . Subgenus Myotis ; mystacinus species group (4 species), which may besister to muricola species group. This species appears to be sister to M. altarium . Previously treated as a race of M. muricola , but split on genetic and morphological grounds. Monotypic. Distribution. NE Kazakhstan , SC & SE Russia (including Sakhalin I), N Mongolia , NE Cin a (Nei Mongol, TE Jilin, and Liaoning), Korean Peninsula (including Jeju I), much ofJapan (Hokkaido, Honshu, Okushiri, and Rishirito Is), and S Kuril Is (Kunashir) Descriptive notes. Head-body 36-55 mm , tail 29-41 mm , ear 8-15 mm , hindfoot 6-9- 5 mm , forearm 30-36 mm . Dorsal pelage of Ikonnikov’s Myotis is uniformly dark brown; ventral pelage slightly paler with slightly darker patch on each side of chest. Bare portions of ears, face, and membranes are dark grayish brown. Ears are relatively long and rather slender with narrowly rounded tip, shallow concavity on posterior borderjust below tip, outward-turned anterior border in middle, and are lightly furred on the inner surface; tragus is just over half ear-height and is slightly concave or straight on anterior margin and convex on posterior margin with pointed tip. Wings are attached to base of outer toe, and uropatagium lacks fringe of hairs on posterior border; calcar is keeled. Skull is small and has pointed rostrum; braincase is broad and low with abruptly rising forehead; interorbital breadth is greater than distance between canines. P? is slightly smaller than P? and is within tooth row, and not leaning inward, P,is small and within tooth row. Chromosomal complement has 2n = 44 and FNa = 52. Habitat. Ikonnikov’s Myotis is mainly found in mountain forests at elevations of 600— 1000 m , although it has been recorded as low as 200 m in Aomori . Known from most forest types throughoutits range and also found in agricultural land. Food and Feeding. Seen foraging for aerial insects low along forest roads or streams. Breeding. Births are known to occur from late June to late July; lactating females and juveniles recorded from late July to early August. Females give birth to single young. Activity patterns. Ikonnikov’s Myotis roosts by day under bark,in tree cavities, between lianas on tree trunks, in old buildings, under bridges, and in farm roofs, rarely tunnels or other subterranean habitats. During the day, it enters torpid state at its roosts, leaving in the evening to forage through the night. It hibernates through winter in holes or crevices inside caves. Calls are a steep FM sweep with an average start frequency of 90 kHz (55-2-118-1 kHz), end frequency 43-2 kHz (37-8-50-1 kHz), peak frequency 50-6 kHz (47-3-55-9 kHz), and duration 2 milliseconds (0-9-3-4 milliseconds) recorded in Hokkaido , Japan . Recordings from South Korea had average start frequencies of 83-4-132-2 kHz, end frequencies 26-6-44-2 kHz, peak frequencies 46-7-62-6 kHz, interpulse intervals 53-121-3 milliseconds, and durations 2:6—4 milliseconds. Movements, Home range and Social organization. In summer, Ikonnikov’s Myotis forms maternity colonies ranging in size from tens to over a hundred. Ikonnikov’s Myotis shares roosts with Greater Japanese Horseshoe Bats (Rhinolophus nippon), Far Eastern Myotis (M. bombinus ), and Big-footed Myotis ( M. macrodactylus ). While hibernating however, it usually roost alone. It switches roosts every 1-2 days and may return to a roost several times. Radio-telemetry shows thatit flies 69-360 m each night while foraging. Status and Conservation. Classified as Least Concern on The IUCN Red List. Tkonnikov’s Myotis is apparently widespread, with no major threats known. Bibliography. Abe et al. (2005), Bannikov (1954), Findley (1972), Fukui , Agetsuma & Hill (2004), Fukui , Hill et al. (2015), Harada & Yoshida (1978), Jo Yeong-Seok et al. (2018), Kawai, Kondo et al. (2006), Kawai, Nikaido et al. (2003), Kondo & Sasaki (2005), Ohdachi et al. (2009), Ono & Obara (1994), Smith & Xie Yan (2008), Stubbe, Ariunbold, Buuveibaatar, Dorjderem, Monkhzul, Otgonbaatar & Tsogbadrakh (2008b), Tsytsulina (2001), Tsytsulina et al. (2012), Yasui et al. (2004).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis ikonnikovi	Myotis	Unassigned-Myotis	ikonnikovi	Ognev	1912	0	Ann. Mus. Zool. Acad. Imp. Sci. St. Petersbourg	0.9979	Ikonnikov's Myotis	 fujiensis Imaizumi, 1954; hosonoi Imaizumi, 1954; ozensis Imaizumi, 1954; yesoensis Yoshiyuki, 1984.	Russia, Primorsk. Krai (= Ussuri Region), Dalnerechen Dist., Euseevka.	Ussuri region and N Korea to Lake Baikal (Russia), the Altai Mtns, and Mongolia, NE China; Sakhalin Isl (Russia) and Honshû and Hokkaido Isls (Japan).	Not listed.	Least Concern	Includes hosonoi,  ozensis and yesoensis; see Abe et al. (2005). Revised by Tsytsulina (2001); also see Corbet (1978 c ), Yoshiyuki (1989), and Benda and Tsytsulina (2000). Molecular sequence data support placement of fujiensis in ikonnikovi (Kawai et al., 2003).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis ikonnikovi	23	Ikonnikov's Myotis	Ikonnikov's Bat|Ikonnikov's Whiskered Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	ikonnikovi	Ognev	1912	0	Myotis_ikonnikovi			ZMMU S-96372 [syntype]		Euseevka, Dalnerechen District, Primorsky Krai (= Ussuri Region), Russia.			ikonnikovi Ognev, 1912|fujiensis Imaizumi, 1954|hosonoi Imaizumi, 1954|ozensis Imaizumi, 1954|yesoensis Yoshiyuki, 1984	includes hosonoi, ozensis, and yesoensis	Ohdachi, S. D. I, Ishibashi, Y., Iwasa, M. A., & Saitoh, Takashi (2009). The Wild Mammals of Japan, Shoukadoh, Kyoto.	Kazakhstan|Russia|Mongolia|China|North Korea|South Korea|Japan	Asia	Palearctic	LC	0	0	0	Myotis_ikonnikovi	0	sciname match	Myotis_ikonnikovi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14168	Myotis ikonnikovi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	ikonnikovi	Ognev, 1912	Belongs to the group “mystacinus ”. Not conspecific with S Asian M. muricola Gray, 1846 (Tsytsulina 2001); includes Japanese fujiensis Imaizumi, 1954 (Kawai et al. 2003).	20000000	Myotis ikonnikovi	Least Concern		2020	2019-07-21 00:00:00 UTC	3.1	English	Myotis ikonnikovi is confirmed as Least Concern, because it is widespread, and there are few major threats to the species across its range. However, the Japanese populations are threatened by loss of old-growth forest. Although the species global population is suspected to be declining, it is not at a rate that qualifies under a threatened criterion.	The species is widely distributed in flat and mountain forests and forest-steppe. The species roosts in tree hollows in the summer and is known to occasionally breed in houses on Honshu (Abe et al. 2005). In continental part of the range it will also roost in rock cracks as well. It hibernates in various underground shelters including caves. It feeds on flying insects low above ground and forest rivers.	The primary threats for the species include deforestation and forest fires which reduce the available habitat for the species. The loss of old-growth forests in Japan is a problem for the species. Recreational cavers can also cause disturbance and mortality in winter roosts.	Myotis ikonnikovi is a naturally rare, but widespread species. ;The species global population is suspected to be in decline due to forest fragmentation and loss of key roost sites, but not at a rate to qualify as a threatened species.	Decreasing	This species ranges from the forested Altai Mountains in Kazakhstan eastwards through Siberia, Cisbaikalia, Transbaikalia, and Lake Baikal to the Ussuri region of Russia, and southwards in to Mongolia, China, Democratic People's Republic of Korea, Republic of Korea, and to Sakhalin Island (Russia) and Japan (Bobrinskiy et al. ;1965, Tiunov 1997, Botvinkin 2002, Kawai 2009, Zhigalin and Khritankov 2014, Zhigalin et al. ;2019). In Japan, the species is found on Honshu and Hokkaido (Abe et al. 2005). In Mongolia it is known from northern mountain habitats in the Ikh Hyangan Mountain Range (Bannikov 1954). The northern extent of the species is ca. 53–55Ëš N Latitude.		Terrestrial	It occurs in protected areas throughout its range. In Japan. M. i. Yezoensis is listed as Endangered, M. i. Hosonoi as Vulnerable, and M. i. Fujiensis as Near Threatened on the Japanese Red List.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	ikonnikovi	Ognev	1912	0	Ann. Mus. Zool. Acad. Imp. Sci. St. Petersbourg	0.997917	Ikonnikov's Myotis	 fujiensis Imaizumi, 1954; hosonoi Imaizumi, 1954; ozensis Imaizumi, 1954; yesoensis Yoshiyuki, 1984.	Russia, Primorsk. Krai (= Ussuri Region), Dalnerechen Dist., Euseevka.	Ussuri region and N Korea to Lake Baikal (Russia), the Altai Mtns, and Mongolia, NE China; Sakhalin Isl (Russia) and Honshû and Hokkaido Isls (Japan).	Not listed.	Least Concern	Includes hosonoi,  ozensis and yesoensis; see Abe et al. (2005). Revised by Tsytsulina (2001); also see Corbet (1978 c ), Yoshiyuki (1989), and Benda and Tsytsulina (2000). Molecular sequence data support placement of fujiensis in ikonnikovi (Kawai et al., 2003).	Myotis ikonnikovi	1005420	23	Ikonnikov's Myotis	Ikonnikov's Bat|Ikonnikov's Whiskered Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	ikonnikovi	Ognev	1912	0	Myotis_ikonnikovi			ZMMU S-96372 [syntype]		Euseevka, Dalnerechen District, Primorsky Krai (= Ussuri Region), Russia.			ikonnikovi Ognev, 1912|fujiensis Imaizumi, 1954|hosonoi Imaizumi, 1954|ozensis Imaizumi, 1954|yesoensis Yoshiyuki, 1984	includes hosonoi, ozensis, and yesoensis	Ohdachi, S. D. I, Ishibashi, Y., Iwasa, M. A., & Saitoh, Takashi (2009). The Wild Mammals of Japan, Shoukadoh, Kyoto.				Kazakhstan|Russia|Mongolia|China|North Korea|South Korea|Japan	Asia	Palearctic	LC	0	0	0	Myotis_ikonnikovi	0	sciname match	Myotis_ikonnikovi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_ikonnikovi	1005420	23	Ikonnikov's Myotis	Ikonnikov's Bat|Ikonnikov's Whiskered Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	ikonnikovi	Ognev	0	Myotis ikonnikovi	Ognev, S.I. 1912-03. О коллекции млекопитающихъ изъ Уссурийскаго края. Annuaire du Musée Zoologique de l'Académie Imperiale des Sciences de St.-Pétersbourg 16:475-511.	https://www.biodiversitylibrary.org/page/8484938	ZMMU S-96372	lectotype	https://zmmu.msu.ru/dbs/list_record.php?id=S-96372	Euseevka, Dalnerechen District, Primorsky Krai (= Ussuri Region), Russia.			includes hosonoi, ozensis, and yesoensis	Ohdachi, S. D. I, Ishibashi, Y., Iwasa, M. A., & Saitoh, Takashi (2009). The Wild Mammals of Japan, Shoukadoh, Kyoto.				Kazakhstan|Russia|Mongolia|China|North Korea|South Korea|Japan	Asia	Palearctic	LC	0	0	0	Myotis_ikonnikovi	0	sciname match	Myotis_ikonnikovi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	ikonnikovi	Ognev	1912	0	Ann. Mus. Zool. Acad. Imp. Sci. St. Petersbourg	0.997917	Ikonnikov's Myotis	fujiensis Imaizumi, 1954; hosonoi Imaizumi, 1954; ozensis Imaizumi, 1954; yesoensis Yoshiyuki, 1984.	Russia, Primorsk. Krai (= Ussuri Region), Dalnerechen Dist., Euseevka.	Ussuri region and N Korea to Lake Baikal (Russia), the Altai Mtns, and Mongolia, NE China; Sakhalin Isl (Russia) and Honshû and Hokkaido Isls (Japan).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14168/22057122/' target='_blank'>Least Concern</a>	Includes hosonoi, ozensis and yesoensis; see Abe et al. (2005). Revised by Tsytsulina (2001); also see Corbet (1978c), Yoshiyuki (1989), and Benda and Tsytsulina (2000). Molecular sequence data support placement of fujiensis in ikonnikovi (Kawai et al., 2003).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis ikonnikovi; Myotis ikonnikovi; Myotis ikonnikovi; Myotis ikonnikovi; Myotis ikonnikovi; Myotis ikonnikovi; fujiensis; fujiensis; hosonoi; ozensis; yesoensis; ikonnikovi; fujiensis; hosonoi; ozensis; yesoensis; Murin d'lkonnikov; Kurzohr-Bartfledermaus; Ratonero de lkonnikov; lkonnikov's Bat; lkonnikov's Whiskered Bat; Ikonnikov's Myotis; Ikonnikov's Bat; Ikonnikov's Whiskered Bat; Ikonnikov's Myotis; Ikonnikov's Myotis; M. ikonnikovi