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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L883	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis horsfieldi	Myotis horsfieldii	Myotis horsfieldii	Myotis horsfieldii	Myotis horsfieldii	Myotis horsfieldii	Myotis horsfieldii	Myotis horsfieldii	Myotis horsfieldii	Myotis horsfieldii	Myotis horsfieldii	Myotis horsfieldii	Myotis horsfieldii	Myotis horsfieldii	Myotis horsfieldii		[MSW2] Subgenus Leuconoe.; [MSW3] Reviewed by Hill (1983), Bates and Harrison (1997), Bates et al. (1999), and Hendrichsen et al. (2001). Apparently closely related to macrotarsus and hasseltii; see Ruedi and Mayer (2001).; [HMW] Vespertilio horsfieldii Temminck, 1840 , Mount Gede, Java , Indonesia . Subgenus Myotis ; horsfieldii species group. See M. ridleyi . Myotis horsfieldit was previously included under M. adversus and later found closely related to M. macrotarsus and M. hasseltii , with M. browni in a more basal position. Myotis horsfieldii represents a species complex in need of deep revision. Five subspecies recognized.; [batnames2022] Reviewed by Hill (1983), Bates and Harrison (1997), Bates et al. (1999), and Hendrichsen et al. (2001). Apparently closely related to macrotarsus and hasseltii ; see Ruedi and Mayer (2001).; [IUCN] This ;species ;represents a ;complex ;that ;requires ;revision, ;with ;some closely related species ;indistinguishable (G. ;Csorba ;pers. comm.). Earlier included under ;Myotis ;adversus ; Horsfield, 1824 (Ellerman and Morrison-Scott 1951), the taxa ;dryas ; Andersen, 1907 and ;peshwa ; Thomas, 1915, are considered subspecies of this taxon (Hill 1983, Corbet and Hill 1992, Bates and Harrison 1997, Simmons 2005).; [batnames2023] Reviewed by Hill (1983), Bates and Harrison (1997), Bates et al. (1999), and Hendrichsen et al. (2001). Apparently closely related to macrotarsus and hasseltii ; see Ruedi and Mayer (2001).; [batnames2025_1.7] Reviewed by Hill (1983), Bates and Harrison (1997), Bates et al. (1999), and Hendrichsen et al. (2001). Apparently closely related to macrotarsus and hasseltii; see Ruedi and Mayer (2001).				lepidus and deignani	(dryas) (peshwa)	deignani, dryas, jeannei, lepidus, peshwa (see Hill, 1983).	peshwa, deignani, dryas, horsfieldii, jeannei	horsfieldii, deignani, dryas, jeannei, peshwa	lepidus	horsfieldii, deignani, dryas, jeannei, peshwa		horsfieldii, deignani, dryas, jeannei, peshwa	horsfieldii - lepidus 	horsfieldii, dryas, lepidus, peshwa, jeannei , deignani	This ;species ;represents a ;complex ;that ;requires ;revision, ;with ;some closely related species ;indistinguishable (G. ;Csorba ;pers. comm.). Earlier included under ;Myotis ;adversus ; Horsfield, 1824 (Ellerman and Morrison-Scott 1951), the taxa ;dryas ; Andersen, 1907 and ;peshwa ; Thomas, 1915, are considered subspecies of this taxon (Hill 1983, Corbet and Hill 1992, Bates and Harrison 1997, Simmons 2005).	horsfieldii, deignani, dryas, jeannei, peshwa	horsfieldii - lepidus 	horsfieldii, dryas, lepidus, peshwa, jeannei, deignani 	horsfieldii, dryas, lepidus, peshwa, jeannei, deignani, horsfieldi	deignani, dryas, horsfieldii, jeannei, peshwa	horsfieldii - lepidus	horsfieldii (Temminck, 1840)|horsfieldi (Jerdon, 1867) [incorrect subsequent spelling]|dryas Andersen, 1907|lepidus (O. Thomas, 1915)|peshwa (O. Thomas, 1915)|jeannei E. H. Taylor, 1934|deignani Shamel, 1942		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		S China – Java, Bali, Borneo, Celebes; ref. 4.61	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis horsfieldii	Indonesia, Java.	Temminck	1840	Monogr. Mamm., 2:226.	Distribution: Ranging from In- dia and southeastern China to the Philippines and Celebes.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	(? jeannei)	India – S China, Java, Bali, Sulawesi, ? Philippine Is	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Temminck	1840	Monogr. Mamm., 2:226.	Subgenus Leuconoe.	SE China, Thailand, India (including Andaman Isis), perhaps Sri Lanka, W Malaysi, Java, Bali, and Sulawesi, Borneo, Philippines.	Indonesia, Java.		TEMMINCK	1840	Size medium (forearm length, 35-40 mm; condylobasal length, 13-15 mm). Margin of plagiopatagium attached to side of foot, which is fairly large. Rostrum of medium width. Middle upper premolar in toothrow or somewhat displaced medially. Uropatagium without a fringe of hair. Postpalatal extension of skull well developed.	Distribution: Ranging from India and southeastern China to the Philippines and Celebes.	Five subspecies are currently recognized:	M. h. peshwa (India), M. h. deignani (southeastern China, northern Thailand), M. h. dryas (Andaman islands), M. h. horsfieldii (Malaya to Bali and Celebes), M. h.jeannei (Philippines).	106	species	M. horsfieldii	TEMMINCK	1840	Leuconoe	subgenus	Myotis horsfieldii				Size medium (forearm length, 35-40 mm; condylobasal length, 13-15 mm). Margin of plagiopatagium attached to side of foot, which is fairly large. Rostrum of medium width. Middle upper premolar in toothrow or somewhat displaced medially. Uropatagium without a fringe of hair. Postpalatal extension of skull well developed.	Five subspecies are currently recognized:		50. M. horsfieldii (TEMMINCK 1840) [adversus group].	50	_M. h. deignani_ Shamel, 1942; _M. h. dryas_ Andersen, 1907; _M. h. horsfieldii_ (Temminck, 1840) (synonyms: _lepidus_ (Thomas, 1915)); _M. h. jeannei_ Taylor, 1934; _M. h. peshwa_ (Thomas, 1915)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis horsfieldii	Myotis		horsfieldii	Temminck	y	1840		Monogr. Mamm.	2		226		Horsfield's Myotis	Indonesia, Java, Mount Gede.	India (including Andaman Isls, SE China, Thailand, Burma, Laos, Vietnam, W Malaysia, Java, Bali, Sulawesi, Borneo, Philippines.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	lepidus Thomas, 1915; deignani Shamel, 1942; dryas K. Andersen, 1907; jeannei Taylor, 1934; peshwa Thomas, 1915.	Reviewed by Hill (1983), Bates and Harrison (1997), Bates et al. (1999), and Hendrichsen et al. (2001). Apparently closely related to macrotarsus and hasseltii; see Ruedi and Mayer (2001).	4C3D87E8FF366A8AFA799C2F1687BA9D	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	967	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF366A8AFA799C2F1687BA9D.xml	Myotis horsfieldii	Vespertilionidae	Myotis	horsfieldii		1840	Murin de Horsfield @fr | Horsfield-Langfu Rfledermaus @de | Ratonero de Horsfield @es | Common Asiatic Myotis @en | Horsfield's Bat @en | Lesser Large-tooth Bat @en	Vespertilio horsfieldii Temminck, 1840 , Mount Gede, Java , Indonesia . Subgenus Myotis ; horsfieldii species group. See M. ridleyi . Myotis horsfieldit was previously included under M. adversus and later found closely related to M. macrotarsus and M. hasseltii , with M. browni in a more basal position. Myotis horsfieldii represents a species complex in need of deep revision. Five subspecies recognized.	M.h.horsfieldiiTemminck,1840—PeninsularMalaysia(includingLangkawiandPen-angIs),Singapore,Borneo,SSumatra(Lampung),Java,Bali,Lombok,andSulawesi. M.h.deignaniShamel,1942—SEChina(Guangdong,Hongkong,andHainan)andmainlandSEAsia. M.h.dryasK.Andersen,1907—AndamanandNicobarIs. M.h.jeanne:E.H.Taylor,1934—Philippines. M. h. peshwa Thomas, 1915 — C, W & NE India ( Madhya Pradesh , Assam , Meghalaya , Maharashtra , Goa , Karnataka , Kerala , and Tamil Nadu ).	Head-body 44-51 mm ,tail 33-43 mm , ear 13-5—- 17 mm , hindfoot 7-11 mm , forearm 33-8-41- 5 mm ; weight 5-7- 5 g . Specimens from India (peshwa) are slightly larger (forearm 36-5—41- 5 mm ) and have browner pelage. Subspecies dryas from the Andaman Islands is on average smaller (forearm 34-8-37- 1 mm ). Fur is soft and dense,feet large, and ears long. Upperparts are dark brown to almost black, with dark brown hair tips that can be grizzled with gray in some individuals. Underparts are gray to dark gray, with pale gray or buffy hair tips and very dark roots. Fur surrounding anal region is nearly white. Ears are dark brown to black and relatively tall compared with size of head and have smoothly convex anterior borders,slightly concave posterior borders, and bluntly pointed tips. Tragus is narrowly pointed and is less than one-half the height of pinna. Feet are moderately large and exceed one-half the length of tibia. Claws are small. Interfemoral and wing membranes are chocolate-brown. Wing membrane attaches to metatarsus below ankle. Subspecies deignani has tricolored hairs: broad band of dark basal hairs, followed by pale buff band, and then narrow dark brown band, tipped with flaxen. Mingling of these colors gives it a grizzled appearance compared with individuals from Java, which does not have a tricolored appearance and is larger in external and cranial measurements. Baculum is small and saddle-shaped, with rounded tip (specimen from India ). Of two penial bones from Vietnam , one (probably immature specimen) was similar to that described from India . The second was larger, c. 0-72 mm , with almost parallel sides, abruptly narrowing to blunt tip; its upper side had abrupt tooth-like elevation on its basal onequarter. Skull is medium-sized, with moderately broad (7-2-7- 7 mm ) braincase. Rostrum is robust, with shallow depression in its midline. Sagittal crest is absent or very weak. Supraocciptal is bulbous. Zygomata are well developed and outwardly flared. C! is nearly twice the height of P*. P? is ¢.50% of crown area of P?, usually lies in tooth row oris slightly displaced internally, and is in close contact with P*. Upper molars are typically myotodont. P, is ¢.50% the height of the canine, which is higher than P. P,, which occasionally can be displaced internally, is sometimes in close contact with P,. Condylo-basal lengths are 14-3-14- 8 mm ; maxillary tooth row lengths are 5-8- 6 mm . Chromosomal complement has 2n = 44 and FN = 52 ( Thailand , Peninsular Malaysia , and Sabah in Borneo) and 2n = 44 and FN = 50 ( Hainan , China ). This difference in numbers of chromosomal arms was due to use of different criteria for classification of chromosome types.	Wide variety of habitats including wooded areas with local supply of freshwater, mixed and hill forests, and tea estates ( India ); lowland forests and agricultural areas, adjacent to limestone karst ( Myanmar ); dry dipterocarp and dry evergreen forests ( Thailand ); lowland forests,riverine forests, disturbed forests, and open land near villages ( Vietnam ); lowland and hill forest ( Malaysia ); lowland forests, close to large rivers and streams (Borneo and south-western Sumatra); secondary and primary lowland, montane, and mossy forests and agricultural areas ( Philippines ); and secondary forests (Sulawesi) from sea level up to elevations of ¢. 1450 m .	Horsfield’s Myotis typically flies in circles c. 10 cm above water, similar to European Daubenton’s Myotis ( M. daubentonii ). It was observed foraging near forest streams, skimming insects from water surface in some instances. On Da Lat plateau, Vietnam , it foraged in the air, similar to the Nepalese Whiskered Myotis ( M. muricola ).	In India ( Madhya Pradesh ), pregnant Horsfield’s Myotis were caught in February-March. In Malaysia , pregnant females were recorded in January-June and October; a few females had twin fetuses.	Horsfield’s Myotis is nocturnal. On Pulau Langkawi, Malaysia , it has been found roosting in limestone caves. In India , it often roosts in culverts, tunnels, and houses and under bridges. In Madhya Pradesh , it was found in leaves of a palmyra palm. On the Philippines , it roosted beneath a large rock over a stream. It might depend on caves near water sources; it is usually absent where streams are lacking. It disappears when water dries up. Echolocation calls are steeply FM without any CF component; frequencies are 45-100 kHz. Call characteristics imply that it can detect prey close to substrate or over water.	Horsfield’s Myotis is often the most common insectivorous bat species in some areas. It was the most frequently captured species in tunneltraps, representing 45% and 66% of tunnel trap captures on the west and south slopes of Mount Isarog, Luzon. It normally roosts in small (2-5) to mediumsized groups. One roost had more than 100 individuals. In Assam , it was netted while coming out of a culvert where a group of 4-5 individuals was roosting. Horsfield’s Myotis often share roosts with species of Rhinolophus, Hipposideros, and Miniopterus .	Classified as Least Concern on The IUCN Red List. Horsfield’s Myotis is widespread, presumably has a large population, occurs in a number of protected areas, tolerates some habitat modification, and is unlikely to be declining fast enough to qualify for listing in a more threatened category. In South Asia,its habitat is being deforested for timber, firewood, and agriculture. It is also threatened by disturbance of roosting sites.	Abramov et al. (2010) | Amador et al. (2018) | Bates & Harrison (1997) | Bates, Hendrichsen et al. (1999) | Bates, Nwe Tin et al. (2005) | Boitani et al. (2006) | Borisenko & Kruskop (2003) | Boro et al. (2018) | Corbet & Hill (1992) | Ellerman & Morrison-Scott (1951) | Francis (2008a) | Francis, Borisenko et al. (2010) | Francis, Guillén-Servent & Robinson (1999) | Harada & Kobayashi (1980) | Heaney, Balete, Dolar et al. (1998) | Heaney, Balete & Rickart (2016) | Hendrichsen, Bates, Hayes & Walston (2001) | Hill (1983) | Honacki etal. (1982) | Huang, J.C.C. etal. (2014) | Hughes et al. (2011) | Keng etal. (2014) | Khajuria (1984) | Kingston et al. (2006) | Kock & Dobat (2000) | Koopman (1994) | Kruskop (2013a, 2013b) | Lim etal. (2017) | Medway (1983) | Molur et al. (2002) | Payne et al. (1985) | Phillipps & Phillipps (2016) | Robinson et al. (1995) | Rosell-Ambal, Tabaranza, Heaney, Gonzales et al. (2008) | Ruedi & Mayer (2001) | Ruedi et al. (2013) | Sedlock (2001) | Sedlock et al. (2008) | Shamel (1942) | Simmons (2005) | Smith & Xie Yan (2008) | Srinivasulu & Srinivasulu (2012) | Srinivasulu et al. (2017) | Stadelmann, Jacobs et al. (2004) | Stadelmann, Lin Liangkong et al. (2007) | Supanuam etal. (2013) | Vanitharani (2006) | Volleth & Heller (2012) | Wordley et al. (2014) | Wu Yi, Motokawa et al. (2009)	https://zenodo.org/record/6398930/files/figure.png	468. Horsfield’s Myotis Myotis horsfieldii French: Murin de Horsfield / German: Horsfield-LangfuRfledermaus / Spanish: Ratonero de Horsfield Other common names: Common Asiatic Myotis , Horsfield's Bat , Lesser Large-tooth Bat Taxonomy. Vespertilio horsfieldii Temminck, 1840 , Mount Gede, Java , Indonesia . Subgenus Myotis ; horsfieldii species group. See M. ridleyi . Myotis horsfieldit was previously included under M. adversus and later found closely related to M. macrotarsus and M. hasseltii , with M. browni in a more basal position. Myotis horsfieldii represents a species complex in need of deep revision. Five subspecies recognized. Subspecies and Distribution. M.h.horsfieldiiTemminck,1840—PeninsularMalaysia(includingLangkawiandPen-angIs),Singapore,Borneo,SSumatra(Lampung),Java,Bali,Lombok,andSulawesi. M.h.deignaniShamel,1942—SEChina(Guangdong,Hongkong,andHainan)andmainlandSEAsia. M.h.dryasK.Andersen,1907—AndamanandNicobarIs. M.h.jeanne:E.H.Taylor,1934—Philippines. M. h. peshwa Thomas, 1915 — C, W & NE India ( Madhya Pradesh , Assam , Meghalaya , Maharashtra , Goa , Karnataka , Kerala , and Tamil Nadu ). Descriptive notes. Head-body 44-51 mm ,tail 33-43 mm , ear 13-5—- 17 mm , hindfoot 7-11 mm , forearm 33-8-41- 5 mm ; weight 5-7- 5 g . Specimens from India (peshwa) are slightly larger (forearm 36-5—41- 5 mm ) and have browner pelage. Subspecies dryas from the Andaman Islands is on average smaller (forearm 34-8-37- 1 mm ). Fur is soft and dense,feet large, and ears long. Upperparts are dark brown to almost black, with dark brown hair tips that can be grizzled with gray in some individuals. Underparts are gray to dark gray, with pale gray or buffy hair tips and very dark roots. Fur surrounding anal region is nearly white. Ears are dark brown to black and relatively tall compared with size of head and have smoothly convex anterior borders,slightly concave posterior borders, and bluntly pointed tips. Tragus is narrowly pointed and is less than one-half the height of pinna. Feet are moderately large and exceed one-half the length of tibia. Claws are small. Interfemoral and wing membranes are chocolate-brown. Wing membrane attaches to metatarsus below ankle. Subspecies deignani has tricolored hairs: broad band of dark basal hairs, followed by pale buff band, and then narrow dark brown band, tipped with flaxen. Mingling of these colors gives it a grizzled appearance compared with individuals from Java, which does not have a tricolored appearance and is larger in external and cranial measurements. Baculum is small and saddle-shaped, with rounded tip (specimen from India ). Of two penial bones from Vietnam , one (probably immature specimen) was similar to that described from India . The second was larger, c. 0-72 mm , with almost parallel sides, abruptly narrowing to blunt tip; its upper side had abrupt tooth-like elevation on its basal onequarter. Skull is medium-sized, with moderately broad (7-2-7- 7 mm ) braincase. Rostrum is robust, with shallow depression in its midline. Sagittal crest is absent or very weak. Supraocciptal is bulbous. Zygomata are well developed and outwardly flared. C! is nearly twice the height of P*. P? is ¢.50% of crown area of P?, usually lies in tooth row oris slightly displaced internally, and is in close contact with P*. Upper molars are typically myotodont. P, is ¢.50% the height of the canine, which is higher than P. P,, which occasionally can be displaced internally, is sometimes in close contact with P,. Condylo-basal lengths are 14-3-14- 8 mm ; maxillary tooth row lengths are 5-8- 6 mm . Chromosomal complement has 2n = 44 and FN = 52 ( Thailand , Peninsular Malaysia , and Sabah in Borneo) and 2n = 44 and FN = 50 ( Hainan , China ). This difference in numbers of chromosomal arms was due to use of different criteria for classification of chromosome types. Habitat. Wide variety of habitats including wooded areas with local supply of freshwater, mixed and hill forests, and tea estates ( India ); lowland forests and agricultural areas, adjacent to limestone karst ( Myanmar ); dry dipterocarp and dry evergreen forests ( Thailand ); lowland forests,riverine forests, disturbed forests, and open land near villages ( Vietnam ); lowland and hill forest ( Malaysia ); lowland forests, close to large rivers and streams (Borneo and south-western Sumatra); secondary and primary lowland, montane, and mossy forests and agricultural areas ( Philippines ); and secondary forests (Sulawesi) from sea level up to elevations of ¢. 1450 m . Food and Feeding. Horsfield’s Myotis typically flies in circles c. 10 cm above water, similar to European Daubenton’s Myotis ( M. daubentonii ). It was observed foraging near forest streams, skimming insects from water surface in some instances. On Da Lat plateau, Vietnam , it foraged in the air, similar to the Nepalese Whiskered Myotis ( M. muricola ). Breeding. In India ( Madhya Pradesh ), pregnant Horsfield’s Myotis were caught in February-March. In Malaysia , pregnant females were recorded in January-June and October; a few females had twin fetuses. Activity patterns. Horsfield’s Myotis is nocturnal. On Pulau Langkawi, Malaysia , it has been found roosting in limestone caves. In India , it often roosts in culverts, tunnels, and houses and under bridges. In Madhya Pradesh , it was found in leaves of a palmyra palm. On the Philippines , it roosted beneath a large rock over a stream. It might depend on caves near water sources; it is usually absent where streams are lacking. It disappears when water dries up. Echolocation calls are steeply FM without any CF component; frequencies are 45-100 kHz. Call characteristics imply that it can detect prey close to substrate or over water. Movements, Home range and Social organization. Horsfield’s Myotis is often the most common insectivorous bat species in some areas. It was the most frequently captured species in tunneltraps, representing 45% and 66% of tunnel trap captures on the west and south slopes of Mount Isarog, Luzon. It normally roosts in small (2-5) to mediumsized groups. One roost had more than 100 individuals. In Assam , it was netted while coming out of a culvert where a group of 4-5 individuals was roosting. Horsfield’s Myotis often share roosts with species of Rhinolophus, Hipposideros, and Miniopterus . Status and Conservation. Classified as Least Concern on The IUCN Red List. Horsfield’s Myotis is widespread, presumably has a large population, occurs in a number of protected areas, tolerates some habitat modification, and is unlikely to be declining fast enough to qualify for listing in a more threatened category. In South Asia,its habitat is being deforested for timber, firewood, and agriculture. It is also threatened by disturbance of roosting sites. Bibliography. Abramov et al. (2010), Amador et al. (2018), Bates & Harrison (1997), Bates, Hendrichsen et al. (1999), Bates, Nwe Tin et al. (2005), Boitani et al. (2006), Borisenko & Kruskop (2003), Boro et al. (2018), Corbet & Hill (1992), Ellerman & Morrison-Scott (1951), Francis (2008a), Francis, Borisenko et al. (2010), Francis, Guillén-Servent & Robinson (1999), Harada & Kobayashi (1980), Heaney, Balete, Dolar et al. (1998), Heaney, Balete & Rickart (2016), Hendrichsen, Bates, Hayes & Walston (2001), Hill (1983), Honacki etal. (1982), Huang, J.C.C. etal. (2014), Hughes et al. (2011), Keng etal. (2014), Khajuria (1984), Kingston et al. (2006), Kock & Dobat (2000), Koopman (1994), Kruskop (2013a, 2013b), Lim etal. (2017), Medway (1983), Molur et al. (2002), Payne et al. (1985), Phillipps & Phillipps (2016), Robinson et al. (1995), Rosell-Ambal, Tabaranza, Heaney, Gonzales et al. (2008), Ruedi & Mayer (2001), Ruedi et al. (2013), Sedlock (2001), Sedlock et al. (2008), Shamel (1942), Simmons (2005), Smith & Xie Yan (2008), Srinivasulu & Srinivasulu (2012), Srinivasulu et al. (2017), Stadelmann, Jacobs et al. (2004), Stadelmann, Lin Liangkong et al. (2007), Supanuam etal. (2013), Vanitharani (2006), Volleth & Heller (2012), Wordley et al. (2014), Wu Yi, Motokawa et al. (2009).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis horsfieldii	Myotis	Unassigned-Myotis	horsfieldii	Temminck	1840	1	Monogr. Mamm.	0.2403	Horsfield's Myotis	 lepidus Thomas, 1915; <b> deignani </b> Shamel, 1942; <b>dryas</b> K. Andersen, 1907; <b>jeannei</b> Taylor, 1934; <b> peshwa </b> Thomas, 1915.	Indonesia, Java, Mount Gede.	India (including Andaman Isls), SE China, Thailand, Burma, Laos, Vietnam, W Malaysia, Java, Bali, Sulawesi, Borneo, Philippines.	Not listed.	Least Concern	Reviewed by Hill (1983), Bates and Harrison (1997), Bates et al. (1999), and Hendrichsen et al. (2001). Apparently closely related to macrotarsus and hasseltii ; see Ruedi and Mayer (2001).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis horsfieldii	23	Horsfield's Myotis	Common Asiatic Myotis|Horsfield's Bat|Lesser Large-tooth Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	horsfieldii	Temminck	1840	1	Vespertilio_Horsfieldii	Temminck, C. J. (1840). Monographies de Mammalogie, ou description de quelques genres de mammiféres, dont les espéces ont été observées dens les différens musées de l'Europe. G. Dufour and E. d'Ocagne, Paris, Vol. 2, 226.	https://books.google.com/books?hl=en&lr=&id=Lw9jAAAAcAAJ&oi=fnd&pg=PA1&dq=monographies+de+mammalogie+temminck+vol.+2&ots=o25ZP2qQxx&sig=jNZE7B2jrj-W7sB7y_Bz5_qjud8#v=onepage&q=monographies%20de%20mammalogie%20temminck%20vol.%202&f=false	ZMB 2558 [syntype]		Mount Gede, Java, Indonesia.			horsfieldii (Temminck, 1840)|dryas K. Andersen, 1907|lepidus (O. Thomas, 1915)|peshwa (O. Thomas, 1915)|jeannei E. H. Taylor, 1934|deignani Shamel, 1942	NA	NA	India|China|Vietnam|Laos|Cambodia|Myanmar|Thailand|Malaysia|Singapore|Indonesia|Brunei|Philippines	Asia	Indomalaya|Oceania	LC	0	0	0	Myotis_horsfieldii	0	sciname match	Myotis_horsfieldii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14166	Myotis horsfieldii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	horsfieldii	(Temminck, 1840)	This ;species ;represents a ;complex ;that ;requires ;revision, ;with ;some closely related species ;indistinguishable (G. ;Csorba ;pers. comm.). Earlier included under ;Myotis ;adversus ; Horsfield, 1824 (Ellerman and Morrison-Scott 1951), the taxa ;dryas ; Andersen, 1907 and ;peshwa ; Thomas, 1915, are considered subspecies of this taxon (Hill 1983, Corbet and Hill 1992, Bates and Harrison 1997, Simmons 2005).	20000000	Myotis horsfieldii	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern ;due to the species’ ;widespread ;distribution ;across South and Southeast Asia, presumed large population, ;occurrence ;in a number of protected areas, ;tolerance ;to ;a degree of habitat modification ;and roost disturbance, and because it is unlikely to be declining ;at a rate ;to qualify for listing in a more threatened category.	In South Asia, this species is found in prime forests and tea estates near to water source. It roosts in tunnels, caves, bridges, palm fronds, crevices in old buildings, cracks and hollows between wooden beams (Aul ;and Vijaykumar 2003) either singly or in small groups of a few individuals (Bates and Harrison 1997). In Myanmar the species has been recorded in lowland forests and agricultural areas adjacent to limestone karst, it has also been collected in a limestone cave in disturbed forest (P. ;Bates pers. comm. 2006). In Viet Nam (Borissenko and ;Kruskop ;2003), Thailand (S. ;Bumrungsri ;pers. comm.) and the Philippines, it has been recorded near to streams in lowland forest, disturbed forest and agricultural areas. In the Philippines, it ;roosts in caves ;(Tanalgo ;et al. 2011, ;Phelps et al. , 2016) ;and in tunnels and has been reported roosting beneath a large rock over a stream (Taylor, 1934). ;In Indonesia ;(Sumatra), ;several individuals were ;captured over ;a ;slow-moving ;river (Huang et al. 2014).	Throughout the range of this species, ;suitable habitat ;is being deforested for timber, firewood and conversion for agricultural use ;and urbanization. It is also threatened through disturbance of roosting sites (Molur ;et al. 2002), including caves (Phelps et al. 2016).	This is a moderately common species over much of its range. The abundance, population size and trends for this species ;across its entire range ;are not ;well ;known.	Stable	This ;wide-ranging ;species has been recorded from South Asia, southern China and ;Southeast Asia. In South Asia, this species is presently known from India (Andaman and Nicobar Islands [Great Nicobar, Little Nicobar and Car Nicobar], Goa, Karnataka, Kerala, Madhya Pradesh, Maharashtra and Tamil Nadu) (Aul ;and Vijaykumar 2003, ;Molur ;et al. ;2002, ;Vanitharani ;2006, ;Wordley ;et al. 2014). It has been recorded from sea level to around 800 m ;asl ;(Molur ;et al. 2002). In China, it is found in Guangdong, Hainan and Hong Kong (Smith and ;Xie ;2008, Hu et al. 2014) ;as well as Yunnan Province ;in ;XishuangbannaTropical Botanical Garden (A. Hughes pers. comm. 2018). In mainland Southeast Asia, it ranges from Myanmar in the west, through Thailand, Lao PDR, Viet Nam, Cambodia, Peninsular Malaysia and ;Singapore. Within insular Southeast Asia, it has been recorded from Indonesia (the islands of Java, Lombok, Sumatra ;and Sulawesi), Borneo (Brunei, Indonesia and Malaysia) and the Philippines, where it has been recorded from sea level to 800 m ;asl ;(Heaney ;et al. ;1998) although there is a record from 1,450 m ;asl ;on ;Mount ;Isarog ;in the southeastern portion of Luzon Island. There are records from the Philippine islands of Bohol, ;Catanduanes, Luzon (Camarines ;Sur, Cagayan, Laguna, Pampanga, Quezon, and Rizal provinces), Mindanao (Lanao del Norte Province, Misamis Occidental), Negros, and Palawan (Heaney ;et al. ;1998) and ;Polillo ;(P. ;Alviola ;pers. comm. ;2018).		Terrestrial	The species has been recorded from many protected areas. Within India, the species has been recorded from the Silent Valley National Park in Kerala, Agasthiyamalai Biosphere Reserve in Tamil Nadu (Vanitharani 2006) and Kanha National Park in Madhya Pradesh. Further studies on distribution, abundance, breeding biology, general ecology and population monitoring are recommended (Molur et al. 2002).	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	horsfieldii	Temminck	1840	1	Monogr. Mamm.	0.240278	Horsfield's Myotis	 lepidus Thomas, 1915; <b> deignani </b> Shamel, 1942; <b>dryas</b> K. Andersen, 1907; <b>jeannei</b> Taylor, 1934; <b> peshwa </b> Thomas, 1915.	Indonesia, Java, Mount Gede.	India (including Andaman Isls), SE China, Thailand, Burma, Laos, Vietnam, W Malaysia, Java, Bali, Sulawesi, Borneo, Philippines.	Not listed.	Least Concern	Reviewed by Hill (1983), Bates and Harrison (1997), Bates et al. (1999), and Hendrichsen et al. (2001). Apparently closely related to macrotarsus and hasseltii ; see Ruedi and Mayer (2001).	Myotis horsfieldii	1005417	23	Horsfield's Myotis	Common Asiatic Myotis|Horsfield's Bat|Lesser Large-tooth Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	horsfieldii	Temminck	1840	1	Vespertilio_Horsfieldii	Temminck, C. J. (1840). Monographies de Mammalogie, ou description de quelques genres de mammiféres, dont les espéces ont été observées dens les différens musées de l'Europe. G. Dufour and E. d'Ocagne, Paris, Vol. 2, 226.	https://books.google.com/books?hl=en&lr=&id=Lw9jAAAAcAAJ&oi=fnd&pg=PA1&dq=monographies+de+mammalogie+temminck+vol.+2&ots=o25ZP2qQxx&sig=jNZE7B2jrj-W7sB7y_Bz5_qjud8#v=onepage&q=monographies%20de%20mammalogie%20temminck%20vol.%202&f=false	ZMB 2558 [syntype]		Mount Gede, Java, Indonesia.			horsfieldii (Temminck, 1840)|dryas K. Andersen, 1907|lepidus (O. Thomas, 1915)|peshwa (O. Thomas, 1915)|jeannei E. H. Taylor, 1934|deignani Shamel, 1942	NA	NA				India|China|Vietnam|Laos|Cambodia|Myanmar|Thailand|Malaysia|Singapore|Indonesia|Brunei|Philippines	Asia	Indomalaya|Oceania	LC	0	0	0	Myotis_horsfieldii	0	sciname match	Myotis_horsfieldii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_horsfieldii	1005417	23	Horsfield's Myotis	Common Asiatic Myotis|Horsfield's Bat|Lesser Large-tooth Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	horsfieldii	Temminck	1	Vespertilio horsfieldii	Temminck, C.J. 1840. Livraison 3. Pp. 141–272 in Temminck, C.J. 1835-1841. Monographies de Mammalogie. Tome second. C. C. van der Hoek, Leiden, 392 pp.	https://archive.org/details/monographiedema00temmgoog/page/140/mode/2up	RMNH.MAM.35551, RMNH.MAM.35552, RMNH.MAM.35553, RMNH.MAM.35554, RMNH.MAM.35555, RMNH.MAM.35556, RMNH.MAM.35557, RMNH.MAM.35558, RMNH.MAM.35559, RMNH.MAM.35560, ZMB 2558	syntypes	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35551.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35551.b | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35552 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35553 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35554 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35555 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35556 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35557 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35558 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35559 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35560	Mount Gede, Java, Indonesia.			NA	NA				India|China|Vietnam|Laos|Cambodia|Myanmar|Thailand|Malaysia|Singapore|Indonesia|Brunei|Philippines	Asia	Indomalaya|Australasia	LC	0	0	0	Myotis_horsfieldii	0	sciname match	Myotis_horsfieldii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	horsfieldii	Temminck	1840	1	Monogr. Mamm.	0.240278	Horsfield's Myotis	lepidus Thomas, 1915; deignani Shamel, 1942; dryas K. Andersen, 1907; jeannei Taylor, 1934; peshwa Thomas, 1915.	Indonesia, Java, Mount Gede.	India (including Andaman Isls), SE China, Thailand, Burma, Laos, Vietnam, W Malaysia, Java, Bali, Sulawesi, Borneo, Philippines.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14166/22057415/' target='_blank'>Least Concern</a>	Reviewed by Hill (1983), Bates and Harrison (1997), Bates et al. (1999), and Hendrichsen et al. (2001). Apparently closely related to macrotarsus and hasseltii; see Ruedi and Mayer (2001).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis horsfieldii; Myotis horsfieldii; Myotis horsfieldii; Myotis horsfieldii; Myotis horsfieldii; Myotis horsfieldii; horsfieldii; deignani; dryas; jeannei; peshwa; lepidus; horsfieldii; deignani; dryas; jeannei; peshwa; deignani; dryas; jeannei; peshwa; lepidus; horsfieldii; dryas; lepidus; peshwa; jeannei; deignani; Murin de Horsfield; Horsfield-Langfu Rfledermaus; Ratonero de Horsfield; Common Asiatic Myotis; Horsfield's Bat; Lesser Large-tooth Bat; Horsfield's Myotis; Common Asiatic Myotis; Horsfield's Bat; Lesser Large-tooth Bat; Horsfield's Myotis; Horsfield's Myotis; M. horsfieldii