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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L881	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis hermani	N/A	Myotis hermani	Myotis formosus [synonym of]	Myotis formosus hermani	Myotis hermani	Myotis hermani	Myotis hermani	Myotis hermani	Myotis hermani	Myotis hermani	Myotis hermani	Myotis hermani	Myotis hermani	Myotis hermani		[MSW3] Included in formosus by Findley (1972), but see Corbet and Hill (1992).; [HMW] Myotis hermani Thomas, 1923 , “Sabang [Island], N.W. of Sumatra ,” Indonesia . Subgenus Chrysopteron (15 species). See M. formosus . Myotis herman : seems to be closely related to M. bartelsii , especially in craniodental traits; they may be conspecific, in which case the name bartelsi: has priority. Monotypic.; [batnames2022] Included in formosus by Findley (1972), but see Corbet and Hill (1992) and Csorba et al. (2014).; [IUCN] <p><span lang="HU">Koopman (1994) treated Myotis hermani as a synonym of M. formosus , but Corbet and Hill (1992) and Csorba et al . (2014) evidenced its specific distinctness.</p>; [batnames2023] Included in formosus by Findley (1972), but see Corbet and Hill (1992) and Csorba et al. (2014).; [batnames2025_1.7] Included in formosus by Findley (1972), but see Corbet and Hill (1992) and Csorba et al. (2014).														hermani	<p><span lang="HU">Koopman (1994) treated Myotis hermani as a synonym of M. formosus , but Corbet and Hill (1992) and Csorba et al . (2014) evidenced its specific distinctness.</p>			hermani 	hermani 			hermani O. Thomas, 1923		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		NW Sumatra		N/A							Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Herman's bat	NW Sumatra; R																															NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis hermani	Myotis		hermani	Thomas		1923		Ann. Mag. Nat. Hist., ser. 9	11		252		Herman's Myotis	Indonesia, NW Sumatra, Sabang.	Sumatra (Indonesia).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Data Deficient.		Included in formosus by Findley (1972), but see Corbet and Hill (1992).	4C3D87E8FF406AFFFF7E97551A17B2BF	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	949	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF406AFFFF7E97551A17B2BF.xml	Myotis hermani	Vespertilionidae	Myotis	hermani	Thomas	1923	Murin de Herman @fr | Herman-Mausohr @de | Ratonero de Herman @es | Herman's Bat @en | Herman's Mouse-eared Bat @en	Myotis hermani Thomas, 1923 , “Sabang [Island], N.W. of Sumatra ,” Indonesia . Subgenus Chrysopteron (15 species). See M. formosus . Myotis herman : seems to be closely related to M. bartelsii , especially in craniodental traits; they may be conspecific, in which case the name bartelsi: has priority. Monotypic.	Known from three localities: Songkhla (peninsular Thailand ); Belum-Temenggor Forest Reserve (Peninsular Malaysia ); and Sabang I (oft NW Sumatra).	Head—body 64-68 mm , tail 54-58 mm , ear 19- 8-21 mm , hindfoot 12-19- 8 mm , forearm 56- 1-60 mm ; weight 16-24 g . Largest member of subgenus Chrysopteron. Overall coloration is “rufoniger-type.” Fur on upperparts is bright orange brown, with paler bases to hairs; underparts are similar and only slightly paler; hairs have buffcolored roots and orange tips, the latter frosted with black (not present in the Blackand-orange Myotis , M. formosus ). Wing and interfemoral membranes have orange and black patches, with those parts near the body, forearms and fingers pinkish orange. Ears are dark orange with black rims, and have concave posterior border; tragus is slender,fairly long, and pointed. Feet not greatly enlarged. Larger and redder than the Black-and-orange Myotis . The Painted Woolly Bat (Keriwwoula picta ), with similar wing pattern, is smaller, with funnel-shaped ears, more pointed tragus, and no black on ears and legs. Skull is as in congeners, but significantly larger, with shallow but distinct frontal depression, posteriorly elongated supraoccipital, and well-developed sagittal and lambdoid crests. Dental formula for all members of genusis12/3,C1/1,P 3/3, M 3/3 (x2) = 38. Dental characters are similar to other members of subgenus Chrysopteron. Basal dimensions of C' exceed those of P*, whereas P° is minute, fully displaced lingually, and obscured in lateral view of skull. P, is half the size of P, and partly out of the tooth row. Condylo-basal length 19- 5-20 mm ; maxillary tooth row 8-6-8- 8 mm .	Found in mature lowland forest in Malaysia . In Thailand , it has been found in disturbed old-growth forest at altitudes of 100-200 m . All known records are from below 500 m . It seems to be associated with small streams and understory of rainforest.	Herman's Myotis probably feeds outside fragmented forests.	No information.	Nocturnal. The specimen from Thailand was captured over a seasonal small stream in Khuan Kaowang Forest Park, on the edge of secondary tropical lowland rainforest, characterized by many large standing trees and dense shrub layer. It was covered in a 3-mm layer offat, which may have been energy reserve for breeding period. Nothing is known of roost ecology, but distinctive color pattern may serve as camouflage, as in the Painted Woolly Bat, which roosts in dry black and orange leaves.	No information.	Classified as Data Deficient on The IUCN Red List. Herman's Myotis is only known from only three individuals. It may be declining due to loss of habitat, and is a priority for survey work. The known records are all from protected areas (Khuan Kaowang Forest Park in Thailand , Belum-Temenggor Forest Reserve in Peninsular Malaysia , and Pulau Weh Nature Recreation Park on Sabang).	Amador et al. (2018) | Boitani et al. (2006) | Bumrungsri et al. (2006) | Corbet & Hill (1992) | Csorba, Chou Cheng-Han et al. (2014) | Csorba, Hutson et al. (2016) | Findley (1972) | Francis (1995a, 2008a) | Heaney et al. (1998) | Honacki et al. (1982) | Koopman (1994) | Lim et al. (2017) | Ruedi et al. (2013) | Simmons (2005) | Stadelmann, Jacobs et al. (2004) | Stadelmann, Lin Liangkong et al. (2007) | Thomas (1923)	https://zenodo.org/record/6398817/files/figure.png	420. Herman's Myotis Myotis hermani French: Murin de Herman / German: Herman-Mausohr / Spanish: Ratonero de Herman Other common names: Herman's Bat , Herman's Mouse-eared Bat Taxonomy. Myotis hermani Thomas, 1923 , “Sabang [Island], N.W. of Sumatra ,” Indonesia . Subgenus Chrysopteron (15 species). See M. formosus . Myotis herman : seems to be closely related to M. bartelsii , especially in craniodental traits; they may be conspecific, in which case the name bartelsi: has priority. Monotypic. Distribution. Known from three localities: Songkhla (peninsular Thailand ); Belum-Temenggor Forest Reserve (Peninsular Malaysia ); and Sabang I (oft NW Sumatra). Descriptive notes. Head—body 64-68 mm , tail 54-58 mm , ear 19- 8-21 mm , hindfoot 12-19- 8 mm , forearm 56- 1-60 mm ; weight 16-24 g . Largest member of subgenus Chrysopteron. Overall coloration is “rufoniger-type.” Fur on upperparts is bright orange brown, with paler bases to hairs; underparts are similar and only slightly paler; hairs have buffcolored roots and orange tips, the latter frosted with black (not present in the Blackand-orange Myotis , M. formosus ). Wing and interfemoral membranes have orange and black patches, with those parts near the body, forearms and fingers pinkish orange. Ears are dark orange with black rims, and have concave posterior border; tragus is slender,fairly long, and pointed. Feet not greatly enlarged. Larger and redder than the Black-and-orange Myotis . The Painted Woolly Bat (Keriwwoula picta ), with similar wing pattern, is smaller, with funnel-shaped ears, more pointed tragus, and no black on ears and legs. Skull is as in congeners, but significantly larger, with shallow but distinct frontal depression, posteriorly elongated supraoccipital, and well-developed sagittal and lambdoid crests. Dental formula for all members of genusis12/3,C1/1,P 3/3, M 3/3 (x2) = 38. Dental characters are similar to other members of subgenus Chrysopteron. Basal dimensions of C' exceed those of P*, whereas P° is minute, fully displaced lingually, and obscured in lateral view of skull. P, is half the size of P, and partly out of the tooth row. Condylo-basal length 19- 5-20 mm ; maxillary tooth row 8-6-8- 8 mm . Habitat. Found in mature lowland forest in Malaysia . In Thailand , it has been found in disturbed old-growth forest at altitudes of 100-200 m . All known records are from below 500 m . It seems to be associated with small streams and understory of rainforest. Food and Feeding. Herman's Myotis probably feeds outside fragmented forests. Breeding. No information. Activity patterns. Nocturnal. The specimen from Thailand was captured over a seasonal small stream in Khuan Kaowang Forest Park, on the edge of secondary tropical lowland rainforest, characterized by many large standing trees and dense shrub layer. It was covered in a 3-mm layer offat, which may have been energy reserve for breeding period. Nothing is known of roost ecology, but distinctive color pattern may serve as camouflage, as in the Painted Woolly Bat, which roosts in dry black and orange leaves. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Data Deficient on The IUCN Red List. Herman's Myotis is only known from only three individuals. It may be declining due to loss of habitat, and is a priority for survey work. The known records are all from protected areas (Khuan Kaowang Forest Park in Thailand , Belum-Temenggor Forest Reserve in Peninsular Malaysia , and Pulau Weh Nature Recreation Park on Sabang). Bibliography. Amador et al. (2018), Boitani et al. (2006), Bumrungsri et al. (2006), Corbet & Hill (1992), Csorba, Chou Cheng-Han et al. (2014), Csorba, Hutson et al. (2016), Findley (1972), Francis (1995a, 2008a), Heaney et al. (1998), Honacki et al. (1982), Koopman (1994), Lim et al. (2017), Ruedi et al. (2013), Simmons (2005), Stadelmann, Jacobs et al. (2004), Stadelmann, Lin Liangkong et al. (2007), Thomas (1923).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis hermani	Myotis	Chrysopteron	hermani	Thomas	1923	0	Ann. Mag. Nat. Hist.	ser. 9, 11: 252	Herman's Myotis	None.	Indonesia, NW Sumatra, Sabang.	Sumatra (Indonesia), Thailand, and Malaysia.	Not listed.	Data Deficient	Included in formosus by Findley (1972), but see Corbet and Hill (1992) and Csorba et al. (2014).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis hermani	23	Herman's Myotis	Herman's Bat|Herman's Mouse-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Chrysopteron	hermani	O. Thomas	1923	0	Myotis_hermani	Thomas, O. (1923). On some small mammals, chiefly bats, from the East Indian Archipelago. Annals and Magazine of Natural History, ser. 9, 11, 252.	https://www.biodiversitylibrary.org/item/107006#page/272/mode/1up	BM 1923.1.2.13		"Sabang [Island], N.W. of Sumatra," Indonesia.			hermani O. Thomas, 1923	NA	NA	Thailand|Malaysia|Indonesia	Asia	Indomalaya	DD	0	0	0	Myotis_hermani	0	sciname match	Myotis_hermani	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14165	Myotis hermani	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	hermani	Thomas, 1923	<p><span lang="HU">Koopman (1994) treated Myotis hermani as a synonym of M. formosus , but Corbet and Hill (1992) and Csorba et al . (2014) evidenced its specific distinctness.</p>	20000000	Myotis hermani	Data Deficient		2016	2015-12-30 00:00:00 UTC	3.1	English	Herman's Myotis is assessed as Data Deficient as there is little information available on range, population status and threats for this species. It may be potentially declining due to loss of habitat, and is therefore a priority for survey work. The taxonomic status of this species also needs to be confirmed.	This is a poorly known species. Resistance to human disturbance is unknown but it has been found in disturbed old growth forest in Thailand at an altitude of 100-200 m asl (Bumrungsri et al. , 2006). It probably feeds outside of fragmented forests (S. Bumrungsri pers. comm.). All of the known records are from below 500 m asl.	Major threats to this species are not known, but habitat loss would be a likely threat to the species.	The abundance and population size of this species are not known. The species has a distinct morphology, and is unlikely to have been misidentified.	Unknown	<p><span lang="HU">This species is known by three individuals only, from Sabang Island, Sumatera, Indonesia; Sungai Singgoh in Temenggor Forest Reserve, Perak State, Peninsular Malaysia; (Francis 1995); and Songkhla, Thailand (Bumrungsri et al. 2006).</p>		Terrestrial	<p><span lang="HU">The known records are all from protected areas. However, Pulau Weh Recreation Park (Indonesia) and Temenggor Forest Reserve (Malaysia) offer limited protection, whereas Khuan Kaowang Forest Park (Thailand) is a very small park. There is a need to determine the distributional limits and population status of the species, as well as potential threats and tolerance to human disturbance.</p>	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Chrysopteron	hermani	Thomas	1923	0	Ann. Mag. Nat. Hist.	ser. 9, 11: 252	Herman's Myotis	None.	Indonesia, NW Sumatra, Sabang.	Sumatra (Indonesia), Thailand, and Malaysia.	Not listed.	Data Deficient	Included in formosus by Findley (1972), but see Corbet and Hill (1992) and Csorba et al. (2014).	Myotis hermani	1005416	23	Herman's Myotis	Herman's Bat|Herman's Mouse-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Chrysopteron	hermani	O. Thomas	1923	0	Myotis_hermani	Thomas, O. (1923). On some small mammals, chiefly bats, from the East Indian Archipelago. Annals and Magazine of Natural History, ser. 9, 11, 252.	https://www.biodiversitylibrary.org/item/107006#page/272/mode/1up	BM 1923.1.2.13		"Sabang [Island], N.W. of Sumatra," Indonesia.			hermani O. Thomas, 1923	NA	NA				Thailand|Malaysia|Indonesia	Asia	Indomalaya	DD	0	0	0	Myotis_hermani	0	sciname match	Myotis_hermani	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_hermani	1005416	23	Herman's Myotis	Herman's Bat|Herman's Mouse-eared Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Chrysopteron	hermani	O. Thomas	0	Myotis hermani	Thomas, O. 1923-01-31. On some small mammals, chiefly bats, from the East Indian Archipelago. Annals and Magazine of Natural History (9)11(62):250-255.	https://www.biodiversitylibrary.org/page/34355498	BMNH:Mamm:1923.1.2.13	holotype	https://data.nhm.ac.uk/object/b4435e7f-a56d-4a33-915e-ee5ca28ab418	"Sabang [Island], N.W. of Sumatra," Indonesia.			NA	NA				Thailand|Malaysia|Indonesia	Asia	Indomalaya	DD	0	0	0	Myotis_hermani	0	sciname match	Myotis_hermani	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Chrysopteron	hermani	Thomas	1923	0	Ann. Mag. Nat. Hist.	ser. 9, 11: 252	Herman's Myotis	None.	Indonesia, NW Sumatra, Sabang.	Sumatra (Indonesia), Thailand, and Malaysia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14165/22057251/' target='_blank'>Data Deficient</a>	Included in formosus by Findley (1972), but see Corbet and Hill (1992) and Csorba et al. (2014).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis hermani; Myotis hermani; Myotis hermani; Myotis hermani; Myotis hermani; Myotis hermani; hermani; Murin de Herman; Herman-Mausohr; Ratonero de Herman; Herman's Bat; Herman's Mouse-eared Bat; Herman's Myotis; Herman's Bat; Herman's Mouse-eared Bat; Herman's Myotis; Herman's Myotis; M. hermani
