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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L879	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis hasselti	Myotis hasseltii	Myotis hasseltii	Myotis hasseltii	Myotis hasseltii	Myotis hasseltii	Myotis hasseltii	Myotis hasseltii	Myotis hasseltii	Myotis hasseltii	Myotis hasseltii	Myotis hasseltii	Myotis hasseltii	Myotis hasseltii	Myotis hasseltii		[MSW2] Subgenus Leuconoe.; [MSW3] Reviewed by Hill (1983), Bates and Harrison (1997), and Bates et al. (1999). Apparently closely related to macrotarsus and horsfieldii; see Ruedi and Mayer (2001).; [HMW] Vespertilio hasseltii Temminck, 1840 , “ Bantam ,” Java , Indonesia . Subgenus Myotis ; horsfieldii species group. See M. ridley: and M. macropus . In the past some taxa (abbotti and macellus), now included under M. hasseltii , have been treated as distinct species, treated as subspecies (continentis) of M. adversus , or listed as subspecies (abbotti) under M. adversus . A comprehensive study (including all subspecies across its distribution) is needed to resolve its limits within the current subspecies concept and relationship to M. adversus . Four subspecies recognized.; [batnames2022] Reviewed by Hill (1983), Bates and Harrison (1997), and Bates et al. (1999). Apparently closely related to macrotarsus and horsfieldii ; see Ruedi and Mayer (2001).; [IUCN] Earlier included under Myotis adversus Horsfield, 1824 (Ellerman and Morrison-Scott 1951), now considered distinct (Hill 1983, Corbet and Hill 1992, Koopman 1993, Bates and Harrison 1997, Simmons 2005) (Srinivasulu et al. in press).; [batnames2023] Reviewed by Hill (1983), Bates and Harrison (1997), and Bates et al. (1999). Apparently closely related to macrotarsus and horsfieldii ; see Ruedi and Mayer (2001).; [batnames2025_1.7] Reviewed by Hill (1983), Bates and Harrison (1997), and Bates et al. (1999). Apparently closely related to macrotarsus and horsfieldii; see Ruedi and Mayer (2001).				continentis		abboti, continentis, macellus (see Hill, 1983).	continentis, hasseltii, macellus, abbotti	hasseltii, abboti, continentis, macellus	continentis - berdmorei	hasseltii, abbotti, continentis, macellus		hasseltii, abboti, continentis, macellus	continentis - berdmorei	hasseltii, macellus, berdmorei, abbotii, continentis	Earlier included under Myotis adversus Horsfield, 1824 (Ellerman and Morrison-Scott 1951), now considered distinct (Hill 1983, Corbet and Hill 1992, Koopman 1993, Bates and Harrison 1997, Simmons 2005) (Srinivasulu et al. in press).	hasseltii, abboti, continentis, macellus	continentis - berdmorei	hasseltii, macellus, berdmorei, abbotii, continentis	hasseltii, macellus, macelius, berdmorei, abbotti, continentis, hasselti, abboti	abboti, continentis, hasseltii, macellus	continentis - berdmorei 	hasseltii (Temminck, 1840)|macellus (Temminck, 1840)|macelius (Lesson, 1842) [incorrect subsequent spelling]|berdmorei E. Blyth, 1863|abbotti Lyon, 1916|hasselti (W. W. A. Phillips, 1924) [incorrect subsequent spelling]|continentis Shamel, 1942|abboti Simmons, 2005 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Lesser large-footed bat	Sri Lanka; Thailand – Malaya, Java, Borneo; ref. 4.61	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis hasseltii	Indonesia, Java.	Temminck	1840	Monogr. Mamm., 2:225.	Distribution: Ranging from Sri Lanka and Burma to Java and Borneo.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Sri Lanka; Burma – Malaya, Java, Borneo	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Temminck	1840	Monogr. Mamm., 2:225.	Subgenus Leuconoe.	Burma, Thailand, Cambodia, Vietnam, W Malaysia, Sumatra, Mentawai Isis, Riau Arch., Java, Borneo, Sri Lanka.	Indonesia, Java.		TEMMINCK	1840	Size medium (forearm length, 35-42 mm; condylobasal length, 14-16 mm). Margin of uropatagium attached to ankle. Rostrum fairly broad. Middle upper premolar greatly reduced and displaced medially from the toothrow. Postpalatal extension of skull shortened.	Distribution: Ranging from Sri Lanka and Burma to Java and Borneo.	Four subspecies are currently recognized:	M. h. continentis (Burma to Cambodia), M. h. hasseltii (Malaya to Java), M. h. macellus (Borneo), M. h. abbotti (Mentawei islands west of Sumatra). The Sri Lankan population has not been allocated subpecifically.	106	species	M. hasseltii	TEMMINCK	1840	Leuconoe	subgenus	Myotis hasseltii				Size medium (forearm length, 35-42 mm; condylobasal length, 14-16 mm). Margin of uropatagium attached to ankle. Rostrum fairly broad. Middle upper premolar greatly reduced and displaced medially from the toothrow. Postpalatal extension of skull shortened.	Four subspecies are currently recognized:		51. M. hasseltii (TEMMINCK 1840) [adversus group].	51	_M. h. abbotti_ Lyon, 1916; _M. h. continentis_ Shamel, 1942 (synonyms: _berdmorei_ Blyth, 1863); _M. h. hasseltii_ (Temminck, 1840); _M. h. macellus_ (Temminck, 1840)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis hasseltii	Myotis		hasseltii	Temminck	y	1840		Monogr. Mamm.	2		225		Lesser Large-footed Myotis	Indonesia, Java, Bantam.	E India, Sri Lanka, Burma, Thailand, Cambodia, Vietnam, W Malaysia, Sumatra, Mentawai Isls, Riau Arch., Java, Borneo.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	abboti Lyon, 1916; continentis Shamel, 1942; berdmorei Blyth, 1863; macellus Temminck, 1840.	Reviewed by Hill (1983), Bates and Harrison (1997), and Bates et al. (1999). Apparently closely related to macrotarsus and horsfieldii; see Ruedi and Mayer (2001).	4C3D87E8FF2B6A94FF519A191646B622	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	970	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF2B6A94FF519A191646B622.xml	Myotis hasseltii	Vespertilionidae	Myotis	hasseltii		1840	Murin de Hasselt @fr | Van-Hasselt-Mausohr @de | Ratonero de Hasselt @es | Brown Bat @en | Hasselt's Large-footed Myotis @en | Lesser Large-toothed Bat @en | Van Hasselt's Bat @en	Vespertilio hasseltii Temminck, 1840 , “ Bantam ,” Java , Indonesia . Subgenus Myotis ; horsfieldii species group. See M. ridley: and M. macropus . In the past some taxa (abbotti and macellus), now included under M. hasseltii , have been treated as distinct species, treated as subspecies (continentis) of M. adversus , or listed as subspecies (abbotti) under M. adversus . A comprehensive study (including all subspecies across its distribution) is needed to resolve its limits within the current subspecies concept and relationship to M. adversus . Four subspecies recognized.	M.h.hasselti:Temminck,1840—MalayPeninsula(includingoffshoreLangkawiI),RiauArchipelago,SWSumatra(BukitBarisanSelatanNationalPark),Java,andSumbawaI. M.h.abbottiLyon,1916—MentawaiIs,offWSumatra. M.h.continentisShamel,1942—patchilydistributedinSEAsia. M. h. macellus Temminck, 1840 — Borneo, also E India ( West Bengal ) and Sri Lanka ; but S Asian populations are assigned to the nominate subspecies by some authors. One record from S China ( Yunnan ), subspecies not known.	Head-body 44-60 mm , tail 35-43 mm , ear 19- 8-21 mm , hindfoot 9-11 mm , forearm 37-43 mm ; weight 7-125 g . Upperparts of the Lesser Large-footed Myotis are dark brown to dark gray, with grizzled gray to buffy brown hair tips and dark brown to black hair bases. Underparts are tipped grayish white, with dark bases. Fur short and velvety, it appears silvery when illuminated in flight at night. Face and muzzle are thinly haired, exposing pinkish skin. Ears are large and dark brown, with bluntly pointed tips. Feet are large and attached at ankles or tibiae, more or less opposite bases of calcars. Wing and uropatagium are pale brown. Skull is larger and robust. Braincase is bulbous, broad (8-8- 4 mm ), and almost spherical in outline. Sagittal crest is visible posteriorly; lambdoid crests are present. Zygomata are moderately robust. Rostrum is robust and broad. Coronoid process of each mandible is very well developed and greatly exceeds canine in height. C' is robust and broad and greatly exceeds P* in height. P? is very small and displaced inward so that P* and P* are in contact or nearly so. C, exceeds P, in height and crown area. P, is very small; it is one-third to one-half the crown area of P, and is usually situated in tooth row but can be displaced internally. Condylo-basal lengths are 14—15- 5 mm ; maxillary tooth row length are 5-7— 6- 2 mm . Chromosomal complement has 2n = 44 and FN = 52 ( Malaysia ).	The Lesser Large-footed Myotis prefers dry forests ( Sri Lanka ) but also coastal and mangrove forests, associated with open water (e.g. lakes, rivers, and coastal areas), from sea level up to elevations of ¢. 1000 m . The Lesser Large-footed Myotis has been reported in large cities (e.g. Rangoon and Bangkok).	The Lesser Large-footed Myotisis a low flier, hunting over water surfaces by dipping its large feet into the water. It feeds on small insects (e.g. mosquitoes, gnats, flies, and moths). In Malaysia , it frequented mangrove forests and was seen feeding below treetop level, often flying low over surface of the water to the edge of the sea. Diet included Lepidopteran wing scales. In Rangoon and Bangkok, it has been seen hawking for insects and fish over small ponds and lakes.	In Malaysia (Langkawi Island), adult females have been lactating or in advanced pregnancy in earlyJanuary. Females carry one fetus. In Vietnam , sexually active males with enlarged testes were found roosting with pregnant and lactating females in April; there are apparently no regular nursing colonies. On Sri Lanka , two young and a subadult were recorded in September, suggesting breeding in August-September.	Lesser Large-footed Myotis are nocturnal and roost in crevices of rocks. In Pulau Langkawi, it roosted in limestone caves. In Thailand ,it roosted inside cut bamboos used as the floor of forest dwellings. Calls are steeply FM without any CF component. Calls (durations 2-5-5-5 milliseconds) start at c.82-104 kHz and sweep down to 23-30 kHz.	The Lesser Large-footed Myotis roosts in large numbers. A colony of 25 individuals of both sexes was found in a crack in a large boulder on the foreshore, isolated at high tide. In Sri Lanka , the Lesser Large-footed Myotis roosts alone or in small groups in bamboo clumps, cracks in tree trunks, and abandoned buildings. In Myanmar , a colony of ten Lesser Large-footed Myotis roosted in a meter box located in a single-story, wooden office building on the bank of the Kispanadi River at the foot of a low, tree-covered hill.	Classified as Least Concern on The IUCN Red List. The Lesser Large-footed Myotisis widespread, presumably has a large population, occurs in a number of protected areas, and is unlikely to be declining fast enough to qualify for listing in a threatened category. In South Asia,its habitat is being deforested for timber, firewood, agricultural use, and human settlements, but it seemsto tolerate some habitat modification.	Amador et al. (2018) | Bates & Harrison (1997) | Bates, Hendrichsen et al. (1999) | Bates, Hutson, Carino et al. (2008) | Bates, Nwe Tin et al. (2005) | Boitani et al. (2006) | Borisenko & Kruskop (2003) | Corbet & Hill (1992) | Dobson (1878) | Ellerman & Morrison-Scott (1951) | Francis (2008a) | Heller (1989) | Heller & Volleth (1989) | Hill (1983) | Honacki et al. (1982) | Koopman (1993, 1994) | Kruskop (2013a) | Lekagul & McNeely (1988) | Lim et al. (2017) | Medway (1983) | MOE (2012) | Molur et al. (2002) | Payne et al. (1985) | Pearch & Writer (2009) | Pearch et al. (2003) | Phillips (1935, 1980) | Ruedi & Mayer (2001) | Ruedi et al. (2013) | Shamel (1942) | Simmons (2005) | Srinivasulu & Srinivasulu (2012) | Stadelmann, Jacobs etal. (2004) | Stadelmann, Lin Liangkong et al. (2007) | Tate (1941d) | Thompson & Fenton (1982) | Topal (1974) | Volleth & Heller (2012) | Wellappulli-Arachchi et al. (2014) | Yapa & Ratnavira (2013) | Zhang Libiao, Zhang Jinshuo et al. (2004)	https://zenodo.org/record/6398964/files/figure.png	474. Lesser Large-footed Myotis Myotis hasseltii French: Murin de Hasselt / German: Van-Hasselt-Mausohr / Spanish: Ratonero de Hasselt Other common names: Brown Bat , Hasselt's Large-footed Myotis , Lesser Large-toothed Bat , Van Hasselt's Bat Taxonomy. Vespertilio hasseltii Temminck, 1840 , “ Bantam ,” Java , Indonesia . Subgenus Myotis ; horsfieldii species group. See M. ridley: and M. macropus . In the past some taxa (abbotti and macellus), now included under M. hasseltii , have been treated as distinct species, treated as subspecies (continentis) of M. adversus , or listed as subspecies (abbotti) under M. adversus . A comprehensive study (including all subspecies across its distribution) is needed to resolve its limits within the current subspecies concept and relationship to M. adversus . Four subspecies recognized. Subspecies and Distribution. M.h.hasselti:Temminck,1840—MalayPeninsula(includingoffshoreLangkawiI),RiauArchipelago,SWSumatra(BukitBarisanSelatanNationalPark),Java,andSumbawaI. M.h.abbottiLyon,1916—MentawaiIs,offWSumatra. M.h.continentisShamel,1942—patchilydistributedinSEAsia. M. h. macellus Temminck, 1840 — Borneo, also E India ( West Bengal ) and Sri Lanka ; but S Asian populations are assigned to the nominate subspecies by some authors. One record from S China ( Yunnan ), subspecies not known. Descriptive notes. Head-body 44-60 mm , tail 35-43 mm , ear 19- 8-21 mm , hindfoot 9-11 mm , forearm 37-43 mm ; weight 7-125 g . Upperparts of the Lesser Large-footed Myotis are dark brown to dark gray, with grizzled gray to buffy brown hair tips and dark brown to black hair bases. Underparts are tipped grayish white, with dark bases. Fur short and velvety, it appears silvery when illuminated in flight at night. Face and muzzle are thinly haired, exposing pinkish skin. Ears are large and dark brown, with bluntly pointed tips. Feet are large and attached at ankles or tibiae, more or less opposite bases of calcars. Wing and uropatagium are pale brown. Skull is larger and robust. Braincase is bulbous, broad (8-8- 4 mm ), and almost spherical in outline. Sagittal crest is visible posteriorly; lambdoid crests are present. Zygomata are moderately robust. Rostrum is robust and broad. Coronoid process of each mandible is very well developed and greatly exceeds canine in height. C' is robust and broad and greatly exceeds P* in height. P? is very small and displaced inward so that P* and P* are in contact or nearly so. C, exceeds P, in height and crown area. P, is very small; it is one-third to one-half the crown area of P, and is usually situated in tooth row but can be displaced internally. Condylo-basal lengths are 14—15- 5 mm ; maxillary tooth row length are 5-7— 6- 2 mm . Chromosomal complement has 2n = 44 and FN = 52 ( Malaysia ). Habitat. The Lesser Large-footed Myotis prefers dry forests ( Sri Lanka ) but also coastal and mangrove forests, associated with open water (e.g. lakes, rivers, and coastal areas), from sea level up to elevations of ¢. 1000 m . The Lesser Large-footed Myotis has been reported in large cities (e.g. Rangoon and Bangkok). Food and Feeding. The Lesser Large-footed Myotisis a low flier, hunting over water surfaces by dipping its large feet into the water. It feeds on small insects (e.g. mosquitoes, gnats, flies, and moths). In Malaysia , it frequented mangrove forests and was seen feeding below treetop level, often flying low over surface of the water to the edge of the sea. Diet included Lepidopteran wing scales. In Rangoon and Bangkok, it has been seen hawking for insects and fish over small ponds and lakes. Breeding. In Malaysia (Langkawi Island), adult females have been lactating or in advanced pregnancy in earlyJanuary. Females carry one fetus. In Vietnam , sexually active males with enlarged testes were found roosting with pregnant and lactating females in April; there are apparently no regular nursing colonies. On Sri Lanka , two young and a subadult were recorded in September, suggesting breeding in August-September. Activity patterns. Lesser Large-footed Myotis are nocturnal and roost in crevices of rocks. In Pulau Langkawi, it roosted in limestone caves. In Thailand ,it roosted inside cut bamboos used as the floor of forest dwellings. Calls are steeply FM without any CF component. Calls (durations 2-5-5-5 milliseconds) start at c.82-104 kHz and sweep down to 23-30 kHz. Movements, Home range and Social organization. The Lesser Large-footed Myotis roosts in large numbers. A colony of 25 individuals of both sexes was found in a crack in a large boulder on the foreshore, isolated at high tide. In Sri Lanka , the Lesser Large-footed Myotis roosts alone or in small groups in bamboo clumps, cracks in tree trunks, and abandoned buildings. In Myanmar , a colony of ten Lesser Large-footed Myotis roosted in a meter box located in a single-story, wooden office building on the bank of the Kispanadi River at the foot of a low, tree-covered hill. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Lesser Large-footed Myotisis widespread, presumably has a large population, occurs in a number of protected areas, and is unlikely to be declining fast enough to qualify for listing in a threatened category. In South Asia,its habitat is being deforested for timber, firewood, agricultural use, and human settlements, but it seemsto tolerate some habitat modification. Bibliography. Amador et al. (2018), Bates & Harrison (1997), Bates, Hendrichsen et al. (1999), Bates, Hutson, Carino et al. (2008), Bates, Nwe Tin et al. (2005), Boitani et al. (2006), Borisenko & Kruskop (2003), Corbet & Hill (1992), Dobson (1878), Ellerman & Morrison-Scott (1951), Francis (2008a), Heller (1989), Heller & Volleth (1989), Hill (1983), Honacki et al. (1982), Koopman (1993, 1994), Kruskop (2013a), Lekagul & McNeely (1988), Lim et al. (2017), Medway (1983), MOE (2012), Molur et al. (2002), Payne et al. (1985), Pearch & Writer (2009), Pearch et al. (2003), Phillips (1935, 1980), Ruedi & Mayer (2001), Ruedi et al. (2013), Shamel (1942), Simmons (2005), Srinivasulu & Srinivasulu (2012), Stadelmann, Jacobs etal. (2004), Stadelmann, Lin Liangkong et al. (2007), Tate (1941d), Thompson & Fenton (1982), Topal (1974), Volleth & Heller (2012), Wellappulli-Arachchi et al. (2014), Yapa & Ratnavira (2013), Zhang Libiao, Zhang Jinshuo et al. (2004).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis hasseltii	Myotis	Unassigned-Myotis	hasseltii	Temminck	1840	1	Monogr. Mamm.	0.2396	Lesser Large-footed Myotis	<b> abboti </b>Lyon, 1916; <b> continentis </b>Shamel, 1942; berdmorei Blyth, 1863; <b> macellus </b> Temminck, 1840.	Indonesia, Java, Bantam.	E India, Sri Lanka, Burma, Thailand, Cambodia, Vietnam, W Malaysia, Sumatra, Mentawai Isls, Riau Arch., Java, Borneo.	Not listed.	Least Concern	Reviewed by Hill (1983), Bates and Harrison (1997), and Bates et al. (1999). Apparently closely related to macrotarsus and horsfieldii ; see Ruedi and Mayer (2001).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis hasseltii	23	Lesser Large-footed Myotis	Brown Bat|Hasselt's Large-footed Myotis|Lesser Large-toothed Bat|Van Hasselt's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	hasseltii	Temminck	1840	1						"Bantam," Java, Indonesia.			hasseltii (Temminck, 1840)|macellus (Temminck, 1840)|berdmorei Blyth, 1863|abbotii Lyon, 1916|continentis Shamel, 1942	NA	NA	Sri Lanka|India|China|Myanmar|Thailand|Vietnam|Cambodia|Malaysia|Singapore|Indonesia|Brunei	Asia	Indomalaya|Oceania	LC	0	0	0	Myotis_hasseltii	0	sciname match	Myotis_hasseltii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14164	Myotis hasseltii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	hasseltii	(Temminck, 1840)	Earlier included under Myotis adversus Horsfield, 1824 (Ellerman and Morrison-Scott 1951), now considered distinct (Hill 1983, Corbet and Hill 1992, Koopman 1993, Bates and Harrison 1997, Simmons 2005) (Srinivasulu et al. in press).	20000000	Myotis hasseltii	Least Concern		2020	2018-09-01 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, it occurs in a number of protected areas, has a tolerance of a degree of habitat modification, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	In South Asia, this species prefers dry forests but is also seen in mangrove forests. It roosts either solitary or in groups of few individuals among bamboo, cracks of tree trunks, and in old and ruined buildings. It is a low flyer, hunting over water surfaces even the sea. It feeds on small insects like the mosquitoes, gnats, flies and moths (Bates and Harrison 1997). In Southeast Asia, this species is known to feed over open seas, and probably roosts in mangrove forests; it has also been found more inland, where its roosting and feeding habits are poorly known. In view of the species possible association with coastal habitats, it may be the species might be more widespread on Borneo around the islands coasts. Payne et al. (1985) suggested that bats feeding over open water near Sandakan were probably this species. Borissenko and Kruskop (2003) recorded the species from the Bassac River in Cambodia and from Phnom-Penh, always near large water surfaces. The species has also been recorded from large cities, such as Rangoon and Bangkok, and there is also a locality from near Hanoi (Bates et al. 1999). In Rangoon and Bangkok, it is seen hawking for insects and fish over small ponds and lakes within the city. The species was found to roost in caves at Bukit Barisan Selatan (Opo pers. comm. 2006), and has been found roosting in limestone caves in Langkawi Island (Lim Boo Liat pers. comm. 2006).	There appear to be no major threats to this species as a whole. In South Asia, the habitat of this species is being deforested for timber, firewood and conversion for agricultural use and human settlements (Molur et al. 2002).	This is a reasonably common, but locally distributed species. In South Asia, although it is a fairly common bat a declining trend in the population has been observed (Molur et al. 2002).	Unknown	This species has a patchily recorded distribution in South Asia, Southeast Asia and China. In South Asia, this species is presently known from India (West Bengal) and Sri Lanka (Eastern, Northern, North Central and Southern provinces) and has been recorded up to an elevation of 1,000 m asl (Molur et al. 2002, Yapa 2017)). In Southeast Asia, the species is known from Myanmar, Thailand, Cambodia, Viet Nam (with a record from Co-Loa [Bates et al. 1999]), Peninsular Malaysia, Indonesia (the Mentawai Islands [Pagai Islands], Riau Archipelago, Sumatra [Bukit Barisan Selatan protected area (Huang et al. 2014)] Java and Sumbawa [Maryanto, pers comm. 2006]) and the island of Borneo (records from Kalimantan [Indonesia] and Sarawak [Malaysia]). In China, this species was collected in Bailong Dong, Mile County, Yunnan Province (Zhang et al. 2004)		Terrestrial	This species has been recorded from protected areas in Southeast Asia (eg. Co-Loa, Viet Nam). In South Asia, there are no conservation measures in place and it has not been recorded from any protected areas. Ecological studies and population monitoring recommended (Molur et al. 2002).	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	hasseltii	Temminck	1840	1	Monogr. Mamm.	0.239583	Lesser Large-footed Myotis	<b> abboti </b>Lyon, 1916; <b> continentis </b>Shamel, 1942; berdmorei Blyth, 1863; <b> macellus </b> Temminck, 1840.	Indonesia, Java, Bantam.	E India, Sri Lanka, Burma, Thailand, Cambodia, Vietnam, W Malaysia, Sumatra, Mentawai Isls, Riau Arch., Java, Borneo.	Not listed.	Least Concern	Reviewed by Hill (1983), Bates and Harrison (1997), and Bates et al. (1999). Apparently closely related to macrotarsus and horsfieldii ; see Ruedi and Mayer (2001).	Myotis hasseltii	1005415	23	Lesser Large-footed Myotis	Brown Bat|Hasselt's Large-footed Myotis|Lesser Large-toothed Bat|Van Hasselt's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	hasseltii	Temminck	1840	1						"Bantam," Java, Indonesia.			hasseltii (Temminck, 1840)|macellus (Temminck, 1840)|berdmorei Blyth, 1863|abbotii Lyon, 1916|continentis Shamel, 1942	NA	NA				Sri Lanka|India|China|Myanmar|Thailand|Vietnam|Cambodia|Malaysia|Singapore|Indonesia|Brunei	Asia	Indomalaya|Oceania	LC	0	0	0	Myotis_hasseltii	0	sciname match	Myotis_hasseltii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_hasseltii	1005415	23	Lesser Large-footed Myotis	Brown Bat|Hasselt's Large-footed Myotis|Lesser Large-toothed Bat|Van Hasselt's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	hasseltii	Temminck	1	Vespertilio hasseltii	Temminck, C.J. 1840. Livraison 3. Pp. 141–272 in Temminck, C.J. 1835-1841. Monographies de Mammalogie. Tome second. C. C. van der Hoek, Leiden, 392 pp.	https://archive.org/details/monographiedema00temmgoog/page/140/mode/2up	RMNH.MAM.35147, RMNH.MAM.35148	syntypes	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35147 | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35148.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.35148.b	"Bantam," Java, Indonesia.			NA	NA				Sri Lanka|India|China|Myanmar|Thailand|Vietnam|Cambodia|Malaysia|Singapore|Indonesia|Brunei	Asia	Indomalaya|Australasia	LC	0	0	0	Myotis_hasseltii	0	sciname match	Myotis_hasseltii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	hasseltii	Temminck	1840	1	Monogr. Mamm.	0.239583	Lesser Large-footed Myotis	abboti Lyon, 1916; continentis Shamel, 1942; berdmorei Blyth, 1863; macellus Temminck, 1840.	Indonesia, Java, Bantam.	E India, Sri Lanka, Burma, Thailand, Cambodia, Vietnam, W Malaysia, Sumatra, Mentawai Isls, Riau Arch., Java, Borneo.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14164/22056644/' target='_blank'>Least Concern</a>	Reviewed by Hill (1983), Bates and Harrison (1997), and Bates et al. (1999). Apparently closely related to macrotarsus and horsfieldii; see Ruedi and Mayer (2001).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis hasseltii; Myotis hasseltii; Myotis hasseltii; Myotis hasseltii; Myotis hasseltii; Myotis hasseltii; hasseltii; abboti; continentis; macellus; continentis - berdmorei; hasseltii; abbotti; continentis; macellus; abboti; continentis; macellus; continentis - berdmorei; hasseltii; macellus; berdmorei; abbotii; continentis; Murin de Hasselt; Van-Hasselt-Mausohr; Ratonero de Hasselt; Brown Bat; Hasselt's Large-footed Myotis; Lesser Large-toothed Bat; Van Hasselt's Bat; Lesser Large-footed Myotis; Brown Bat; Hasselt's Large-footed Myotis; Lesser Large-toothed Bat; Van Hasselt's Bat; Lesser Large-footed Myotis; Lesser Large-footed Myotis; M. hasseltii