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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L874	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	Myotis gomantongensis	Myotis gomantongensis	Myotis gomantongensis	Myotis gomantongensis	Myotis gomantongensis	Myotis gomantongensis	Myotis gomantongensis	Myotis gomantongensis	Myotis gomantongensis	Myotis gomantongensis		[MSW3] Based on specimens previously referred to ater by Hill and Francis (1984) and Payne et al. (1985).; [HMW] Myotis gomantongensis Hill & Francis, 1998 , “Gomantong Caves, 5°31°N , 118°04’E ,” Sabah , Borneo, Malaysia . Subgenus Myotis ; muricola species group. Previously treated within M. ater nugax. Genetic studies place this species as sister to M. annectans ; these two may be most closely related to M. muricola , or to M. pilosus and M. chinensis , and perhaps M. montivagus . Relationship with M. ater and M. munricola 1s unclear; all three occur in Borneo, where respective species limits require further study. Monotypic.; [batnames2022] Based on specimens previously referred to ater by Hill and Francis (1984) and Payne et al. (1985).; [IUCN] Myotis gomantongensis was previously listed in the Red List prior to 2004 as M. gomatongensis - an orthographic error in the Chiroptera Action Plan.; [batnames2023] Based on specimens previously referred to ater by Hill and Francis (1984) and Payne et al. (1985).; [batnames2025_1.7] Based on specimens previously referred to ater by Hill and Francis (1984) and Payne et al. (1985).														gomantongensis	Myotis gomantongensis was previously listed in the Red List prior to 2004 as M. gomatongensis - an orthographic error in the Chiroptera Action Plan.			gomantongensis	gomantongensis			gomantongensis C. M. Francis & J. Edwards Hill, 1998						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis gomantongensis	Myotis		gomantongensis	Francis and Hill		1998		Mammalia	62	2	248		Gomantong Myotis	Malaysia, Borneo, Sabah, Gomantong Caves, 5°31'N, 118°04'E.	Sabah (Borneo, Malaysia).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Data Deficient.		Based on specimens previously referred to ater by Hill and Francis (1984) and Payne et al. (1985).	4C3D87E8FF386A80FA869CFE1AABBED0	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	957	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF386A80FA869CFE1AABBED0.xml	Myotis gomantongensis	Vespertilionidae	Myotis	gomantongensis	Hill & Francis	1998	Murin de Gomantong @fr | Gomantong-Mausohr @de | Ratonero de Gomantong @es | Gomantong Bat @en	Myotis gomantongensis Hill & Francis, 1998 , “Gomantong Caves, 5°31°N , 118°04’E ,” Sabah , Borneo, Malaysia . Subgenus Myotis ; muricola species group. Previously treated within M. ater nugax. Genetic studies place this species as sister to M. annectans ; these two may be most closely related to M. muricola , or to M. pilosus and M. chinensis , and perhaps M. montivagus . Relationship with M. ater and M. munricola 1s unclear; all three occur in Borneo, where respective species limits require further study. Monotypic.	N & W Borneo; only known from three localities in Sabah (Gomantong Caves, Baturong Caves, and Sinoa) and recently found in Sarawak (Bukit Sarang, Bintulu division); may be more widespread.	Tail 36-45 mm , ear 12:6—15- 5 mm , hindfoot 8-9 mm , forearm 40- 43 mm ; weight 6-4-8- 8 g . Differs from Peters’s Myotis ( M. ater ) in longer forearm and skull; feet small. Dorsal fur has blackish bases and dark brown tips; underparts are dark brownish gray with black hair bases; there is a golden-brown patch in center of belly, as in its sister species, the Hairy-faced Myotis ( M. annectans ). Ears are medium-sized, with anterior margin convex and a rounded tip; tragus is narrow, c.40% ear length, with slightly concave anterior margin and more strongly convex posterior margin, with a marked triangular lobe at its base. Wing membrane is attached at base of toe; calcar is long and narrow toward tip, with a narrow keel. Skull is long and not especially broad, with inflated braincase and shallow frontal depression; no conspicuous sagittal crest. Skull is much narrower and less massive than in the Bornean Whiskered Myotis ( M. browni ). C' is short, roughly equal to or slightly shorter than posterior upper premolar; P? is small, equal to or smaller than I’, and displaced slightly inward; P* is greatly reduced, less than quarter the surface area of P* and displaced inward, so that P* and P* are in contact; C, is small, ¢.20% shorter than P, and ¢.40% smaller than that tooth in surface area; P, is not particularly small, about half the height of canine, and about one-and-a-half times the surface area of I,, completely in line in tooth row; P, is tiny, completely displaced inward so that P, and P, are in contact or nearly so; P, is about the same height as M,. Differs from Peters’s Myotis in greater reduction of middle premolars, and in the small canines which are similar in length to P*, and distinctly shorter than P,. Condylo-basal length 14-3-15- 3 mm ; maxillary tooth row 5:6- 6-9 mm .	No information.	No information.	No information.	Roosts in limestone caves, and lives in small colonies.	No relevant information.	Classified as Least Concern on The IUCN Red Lis . No threats have been identified. Found in Gomantong Forest Reserve.	Abdullah et al. (2007) | Francis & Hill (1998) | Francis & Hutson (2008) | Francis et al. (2010) | Hill & Francis (1984) | Koopman (1993, 1994) | Kruskop & Borisenko (2013) | Medway (1963, 1977) | Payne et al. (1985) | Phillipps & Phillipps (2016) | Simmons (2005) | Wang Xiaoyun et al. (2017) | Wilson et al. (2006)	https://zenodo.org/record/6398876/files/figure.png	444. Gomantong Myotis Myotis gomantongensis French: Murin de Gomantong / German: Gomantong-Mausohr / Spanish: Ratonero de Gomantong Other common names: Gomantong Bat Taxonomy. Myotis gomantongensis Hill & Francis, 1998 , “Gomantong Caves, 5°31°N , 118°04’E ,” Sabah , Borneo, Malaysia . Subgenus Myotis ; muricola species group. Previously treated within M. ater nugax. Genetic studies place this species as sister to M. annectans ; these two may be most closely related to M. muricola , or to M. pilosus and M. chinensis , and perhaps M. montivagus . Relationship with M. ater and M. munricola 1s unclear; all three occur in Borneo, where respective species limits require further study. Monotypic. Distribution. N & W Borneo; only known from three localities in Sabah (Gomantong Caves, Baturong Caves, and Sinoa) and recently found in Sarawak (Bukit Sarang, Bintulu division); may be more widespread. Descriptive notes. Tail 36-45 mm , ear 12:6—15- 5 mm , hindfoot 8-9 mm , forearm 40- 43 mm ; weight 6-4-8- 8 g . Differs from Peters’s Myotis ( M. ater ) in longer forearm and skull; feet small. Dorsal fur has blackish bases and dark brown tips; underparts are dark brownish gray with black hair bases; there is a golden-brown patch in center of belly, as in its sister species, the Hairy-faced Myotis ( M. annectans ). Ears are medium-sized, with anterior margin convex and a rounded tip; tragus is narrow, c.40% ear length, with slightly concave anterior margin and more strongly convex posterior margin, with a marked triangular lobe at its base. Wing membrane is attached at base of toe; calcar is long and narrow toward tip, with a narrow keel. Skull is long and not especially broad, with inflated braincase and shallow frontal depression; no conspicuous sagittal crest. Skull is much narrower and less massive than in the Bornean Whiskered Myotis ( M. browni ). C' is short, roughly equal to or slightly shorter than posterior upper premolar; P? is small, equal to or smaller than I’, and displaced slightly inward; P* is greatly reduced, less than quarter the surface area of P* and displaced inward, so that P* and P* are in contact; C, is small, ¢.20% shorter than P, and ¢.40% smaller than that tooth in surface area; P, is not particularly small, about half the height of canine, and about one-and-a-half times the surface area of I,, completely in line in tooth row; P, is tiny, completely displaced inward so that P, and P, are in contact or nearly so; P, is about the same height as M,. Differs from Peters’s Myotis in greater reduction of middle premolars, and in the small canines which are similar in length to P*, and distinctly shorter than P,. Condylo-basal length 14-3-15- 3 mm ; maxillary tooth row 5:6- 6-9 mm . Habitat. No information. Food and Feeding. No information. Breeding. No information. Activity patterns. Roosts in limestone caves, and lives in small colonies. Movements, Home range and Social organization. No relevant information. Status and Conservation. Classified as Least Concern on The IUCN Red Lis . No threats have been identified. Found in Gomantong Forest Reserve. Bibliography. Abdullah et al. (2007), Francis & Hill (1998), Francis & Hutson (2008), Francis et al. (2010), Hill & Francis (1984), Koopman (1993, 1994), Kruskop & Borisenko (2013), Medway (1963, 1977), Payne et al. (1985), Phillipps & Phillipps (2016), Simmons (2005), Wang Xiaoyun et al. (2017), Wilson et al. (2006).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis gomantongensis	Myotis	Unassigned-Myotis	gomantongensis	Francis and Hill	1998	0	Mammalia	62(2): 248	Gomantong Myotis	None.	Malaysia, Borneo, Sabah, Gomantong Caves, 5°31'N, 118°04'E.	Sabah (Borneo, Malaysia).	Not listed.	Least Concern	Based on specimens previously referred to ater by Hill and Francis (1984) and Payne et al. (1985).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis gomantongensis	23	Gomantong Myotis	Gomantong Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	gomantongensis	J. Edwards Hill & Francis	1998	0	Myotis_gomantongensis	Francis, C. M. & Hill, J. E. (1998). New records and a new species of Myotis (Chiroptera, Vespertilionidae) from Malaysia. Mammalia, 62(2), 248.	https://www.degruyter.com/document/doi/10.1515/mamm.1998.62.2.241/html	ROM 107811		"Gomantong Caves, 5°31'N, 118°04'E," Sabah, Borneo, Malaysia.	5.5167	118.07	gomantongensis J. Edwards Hill & Francis, 1998	NA	NA	Malaysia|Indonesia?	Asia	Indomalaya	LC	0	0	0	Myotis_gomantongensis	0	sciname match	Myotis_gomantongensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	40035	Myotis gomantongensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	gomantongensis	Francis &; Hill, 1998	Myotis gomantongensis was previously listed in the Red List prior to 2004 as M. gomatongensis - an orthographic error in the Chiroptera Action Plan.	20000000	Myotis gomantongensis	Least Concern		2021	2021-08-29 00:00:00 UTC	3.1	English	Myotis gomantongensis is assessed as Least Concern given its assumed distribution throughout Borneo. While data are very limited, the number of mature individuals of this cave-dependent bat is inferred to be declining, but not at a rate that qualifies it for a threatened category, due to cave disturbance and loss, and the degradation and loss of low elevation rainforests where the species has been document. The species has not been reported from more recent field surveys.	There is limited ecological data available for this insectivorous bat species. Myotis gomantongensis is known to roost in small colonies in large limestone cave systems (Francis and Hill 1998). These large cave systems have several other species of bats, some of which may exist in very large numbers (e.g., Chaerephon plicata ; Abdullah et al. 2007) and effectively limit opportunities for detecting the less common species. Detailed understanding of the species foraging habitats is unknown but is presumably linked to lowland tropical rainforests where its roosts sites have been documented.	Comprehensive understanding of threats to M. gomantongensis is lacking. Given the species is assumed to be dependent on caves for its roosts, continued cave disturbance and loss from recreation and mining are presumed to be significant threats. The loss and degradation of lowland tropical rainforest due to logging and conversion to agriculture is extensive in many parts of Borneo (Meijaard et al. 2005) and are also inferred to pose serious threats to the species.	Population number and trend data are not available for this species. It has only been encountered in small numbers in Gomantong Cave (type locality; 5°31'N, 118°04'E) and Baturong Cave Systems in Sabah, Borneo. The number of mature individuals is inferred to be declining due to cave disturbance and destruction, as well as ongoing loss of lowland tropical rainforests. More recent surveys, including at the species historic roost, have not reported the species (Struebig et al. 2006a,b; Abdullah et al. 2007; Benda 2010; Struebig et al. 2010; Wasti et al. 2020).	Unknown	This species is endemic to Borneo and has only been recorded from the Gomantong Cave (type locality, 5°31'N, 118°04'E) and Baturong Cave Systems in Sabah, Borneo (Lundberg and McFarlane 2011, Francis and Hill 1998). While the species is hypothesized to be more broadly distributed throughout Borneo, including in most of the major cave systems, more recent field surveys have either failed to document and expand the species distribution or failed to report the species (e.g., Struebig et al. 2006a,b; Abdullah et al. 2007; Benda 2010; Struebig et al. 2010; Wasti et al. 2020). Myotis gomantongensis has only been documented in lowland forests below 100 m asl.	The species is not known to be hunted, used, or traded.	Terrestrial	Myotis gomantongensis is known from the Gomantong Forest Reserve (Francis and Hill 1998) and is presumed to be present in other protected areas across Borneo. Research designed to capture this species using appropriate techniques is needed to verify if the species is broadly distributed across Borneo and determine the species' population size and trend, ecology, threats, and priority conservation actions.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	gomantongensis	Francis and Hill	1998	0	Mammalia	62(2): 248	Gomantong Myotis	None.	Malaysia, Borneo, Sabah, Gomantong Caves, 5°31'N, 118°04'E.	Sabah (Borneo, Malaysia).	Not listed.	Least Concern	Based on specimens previously referred to ater by Hill and Francis (1984) and Payne et al. (1985).	Myotis gomantongensis	1005411	23	Gomantong Myotis	Gomantong Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	gomantongensis	Francis & J. Edwards Hill	1998	0	Myotis_gomantongensis	Francis, C. M. & Hill, J. E. (1998). New records and a new species of Myotis (Chiroptera, Vespertilionidae) from Malaysia. Mammalia, 62(2), 248.	https://www.degruyter.com/document/doi/10.1515/mamm.1998.62.2.241/html	ROM 107811		"Gomantong Caves, 5°31'N, 118°04'E," Sabah, Borneo, Malaysia.	5.516667	118.0667	gomantongensis Francis & J. Edwards Hill, 1998	NA	NA				Malaysia|Indonesia?	Asia	Indomalaya	LC	0	0	0	Myotis_gomantongensis	0	sciname match	Myotis_gomantongensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_gomantongensis	1005411	23	Gomantong Myotis	Gomantong Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	gomantongensis	C. M. Francis & J. Edwards Hill	0	Myotis gomantongensis	Francis, C.M. and Hill, J.E. 1998. New records and a new species of _Myotis_ (Chiroptera, Vespertilionidae) from Malaysia. Mammalia 62(2):241-252.	https://doi.org/10.1515/mamm.1998.62.2.241	ROM:MAM:107811	holotype	http://portal.vertnet.org/o/rom/mammals?id=uri-catalog-rom-mammals-107811	"Gomantong Caves, 5°31'N, 118°04'E," Sabah, Borneo, Malaysia.	5.516667	118.0667	NA	NA				Malaysia|Indonesia?	Asia	Indomalaya	LC	0	0	0	Myotis_gomantongensis	0	sciname match	Myotis_gomantongensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	gomantongensis	Francis & Hill	1998	0	Mammalia	62(2): 248	Gomantong Myotis	None.	Malaysia, Borneo, Sabah, Gomantong Caves, 5°31'N, 118°04'E.	Sabah (Borneo, Malaysia).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/40035/22060096/' target='_blank'>Least Concern</a>	Based on specimens previously referred to ater by Hill and Francis (1984) and Payne et al. (1985).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis gomantongensis; Myotis gomantongensis; Myotis gomantongensis; Myotis gomantongensis; Myotis gomantongensis; Myotis gomantongensis; gomantongensis; Murin de Gomantong; Gomantong-Mausohr; Ratonero de Gomantong; Gomantong Bat; Gomantong Myotis; Gomantong Bat; Gomantong Myotis; Gomantong Myotis; M. gomantongensis