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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L869	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis fimbriatus	Myotis macrodactylus [synonym of]	Myotis fimbriatus	Myotis macrodactylus [synonym of]	Myotis macrodactylus fimbriatus	Myotis fimbriatus	Myotis fimbriatus	Myotis fimbriatus	Myotis fimbriatus	Myotis fimbriatus	Myotis fimbriatus	Myotis fimbriatus	Myotis fimbriatus	Myotis fimbriatus	Myotis fimbriatus		[MSW3] Not conspecific with macrodactylus or capaccinii; see Corbet (1978c) and Corbet and Hill (1992).; [HMW] Vespertilio fimbriatus Peters in Swinhoe, 1871, “Amoy,” Fujian , China . Subgenus Myotis ; macrodactylus species group. See M. petax . Specimens from Yunnan did not cluster with the rest of M. fimbriatus (being sister to M. pilosus and true M. fimbriatus ) but are similar in general morphology, except for their larger size. These specimens likely represent an undescribed species (M. cf. fimbriatus ), although further studies are needed, and these populations are tentatively included here. Myotis taiwanensis was recently recognized as a distinct species, but later studies found it to be a subspecies of M. fimbriatus . ‘Two subspecies recognized.; [batnames2022] Not conspecific with macrodactylus or capaccinii ; see Corbet (1978 c ) and Corbet and Hill (1992). Includes taiwanensis , see Ruedi et al. (2015).; [IUCN] Various authors have suggested relationships with ;M. pilosus , ;M. macrodactylus , and ;M. petax , but ;M. fimbriatus ;should be best considered a Chinese endemic species (Smith and Xie 2008, Ruedi et al. 2015).; [batnames2023] Not conspecific with macrodactylus or capaccinii ; see Corbet (1978 c ) and Corbet and Hill (1992). Includes taiwanensis , see Ruedi et al. (2015).; [batnames2025_1.7] Not conspecific with macrodactylus or capaccinii; see Corbet (1978c) and Corbet and Hill (1992). Includes taiwanensis, see Ruedi et al. (2015).									hirsutus	fimbriatus, taiwanensis		fimbriatus, taiwanensis	fimbriatus - hirsutus	fimbriatus, taiwanensis, hirsutus	Various authors have suggested relationships with ;M. pilosus , ;M. macrodactylus , and ;M. petax , but ;M. fimbriatus ;should be best considered a Chinese endemic species (Smith and Xie 2008, Ruedi et al. 2015).	fimbriatus, taiwanensis	fimbriatus - hirsutus	fimbriatus, taiwanensis, hirsutus 	fimbriatus, taiwanensis, hirsutus 	fimbriatus, taiwanensis	fimbriatus - hirsutus	fimbriatus (W. C. H. Peters in Swinhoe, 1871)|taiwanensis Ärnbäck-Christie-Linde, 1908|hirsutus A. B. Howell, 1926		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Fukien, SE China		N/A							Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Fujian, SE China																															NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis fimbriatus	Myotis		fimbriatus	Peters	y	1870	1871	In R. Swinhoe, Catalogue of Mammals of China, Proc. Zool. Soc. Lond.	1870		617		Fringed Long-footed Myotis	China, Fujian, Amoy.	SE China.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (nt).	hirsutus Howell, 1926.	Not conspecific with macrodactylus or capaccinii; see Corbet (1978c) and Corbet and Hill (1992).	4C3D87E8FF2F6A90FF51975E1FA4B225	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	974	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF2F6A90FF51975E1FA4B225.xml	Myotis fimbriatus	Vespertilionidae	Myotis	fimbriatus		1871	Murin de Swinhoe @fr | Haarige Wasserfledermaus @de | Ratonero de Swinhoe @es | Hairy-legged Myotis @en	Vespertilio fimbriatus Peters in Swinhoe, 1871, “Amoy,” Fujian , China . Subgenus Myotis ; macrodactylus species group. See M. petax . Specimens from Yunnan did not cluster with the rest of M. fimbriatus (being sister to M. pilosus and true M. fimbriatus ) but are similar in general morphology, except for their larger size. These specimens likely represent an undescribed species (M. cf. fimbriatus ), although further studies are needed, and these populations are tentatively included here. Myotis taiwanensis was recently recognized as a distinct species, but later studies found it to be a subspecies of M. fimbriatus . ‘Two subspecies recognized.	M.f.fimbriatusPeters,1871—E&SEChina(Shandong,Jiangsu,Anhui,Zhejiang,Jiangxi,Fujian,andGuangdong). M. f. taiwanensis Arnback-Christie-Linde, 1908 — Taiwan . Specimens from Yunnan , SC China and possibly specimens from Sichuan and Guizhou represent an undescribed species.	Head-body 35- 7-52 mm ,tail 33- 2-48 mm , hindfoot 8-12- 4 mm , ear 10:6-17- 2 mm , forearm 37-42- 7 mm . Dorsal pelage of the Fringed Long-footed Myotis is grayish brown; venteris taupe or off-white, becoming almost pure white toward anal region; hairs have dark slate-gray bases and much lighter tips. Ears are relatively long; tragus is nearly parallel, long, and pointed, reaching one-half the ear length. Wings attach to ankle or base of metatarsus; calcar is very long and unkeeled, extending four-fifths the rear margin of uropatagium. Feet are large, and uropatagium has fringe of comb-shaped hairs along its rear border. Skull has inflated, globose braincase (especially subspecies taiwanensis ); C' is robust and taller than P* P? is small and in tooth row orslightly leaning inward; C, is weaker but still taller than P; and lower molars are strong, with high cusps, and are all myotodont. Chromosomal complement has 2n = 44 and FNa = 50 ( Guizhou , Henan , and Taiwan) .	Lowland and montane forest habitats, generally near water.	The Fringed Long-footed Myotis forages by catching insects above the water surface and in open grasslands and farmlands. It flies slowly and is maneuverable. It feeds mostly on Diptera by aerial hawking but also occasionally gleans prey off surfaces.	Newborn Fringed Long-footed Myotis have been reported in November— June in Taiwan , suggesting a very long breeding season.	Fringed Long-footed Myotis primarily roost in caves. Calls are steep FM sweeps, with frequencies of 29-93 kHz on first harmonic and 70-122 kHz on second harmonic and mean duration of 10-6 milliseconds recorded in China .	Fringed Long-footed Myotis roost in large colonies; maternity colonies can be up to 1000 individuals in Taiwan .	Classified as Least Concern on The IUCN Red Lust.	Findley (1972) | Han Naijian et al. (2010) | Lin Liangkong, Motokawa & Harada (2002a) | Liu Ying et al. (2003) | Niu Hongxing, Yu Yan & Wang Yanmei (2007) | Ruedi, Csorba et al. (2015) | Ruedi, Stadelmann et al. (2013) | Smith & Xie Yan (2008) | Smith, Johnston, Jones & Rossiter ( 2008g) | Wang Hui et al. (2009) | Zhang Zhenzhen et al. (2009)	https://zenodo.org/record/6398973/files/figure.png	477. Fringed Long-footed Myotis Myotis fimbriatus French: Murin de Swinhoe / German: Haarige Wasserfledermaus / Spanish: Ratonero de Swinhoe Other common names: Hairy-legged Myotis Taxonomy. Vespertilio fimbriatus Peters in Swinhoe, 1871, “Amoy,” Fujian , China . Subgenus Myotis ; macrodactylus species group. See M. petax . Specimens from Yunnan did not cluster with the rest of M. fimbriatus (being sister to M. pilosus and true M. fimbriatus ) but are similar in general morphology, except for their larger size. These specimens likely represent an undescribed species (M. cf. fimbriatus ), although further studies are needed, and these populations are tentatively included here. Myotis taiwanensis was recently recognized as a distinct species, but later studies found it to be a subspecies of M. fimbriatus . ‘Two subspecies recognized. Subspecies and Distribution. M.f.fimbriatusPeters,1871—E&SEChina(Shandong,Jiangsu,Anhui,Zhejiang,Jiangxi,Fujian,andGuangdong). M. f. taiwanensis Arnback-Christie-Linde, 1908 — Taiwan . Specimens from Yunnan , SC China and possibly specimens from Sichuan and Guizhou represent an undescribed species. Descriptive notes. Head-body 35- 7-52 mm ,tail 33- 2-48 mm , hindfoot 8-12- 4 mm , ear 10:6-17- 2 mm , forearm 37-42- 7 mm . Dorsal pelage of the Fringed Long-footed Myotis is grayish brown; venteris taupe or off-white, becoming almost pure white toward anal region; hairs have dark slate-gray bases and much lighter tips. Ears are relatively long; tragus is nearly parallel, long, and pointed, reaching one-half the ear length. Wings attach to ankle or base of metatarsus; calcar is very long and unkeeled, extending four-fifths the rear margin of uropatagium. Feet are large, and uropatagium has fringe of comb-shaped hairs along its rear border. Skull has inflated, globose braincase (especially subspecies taiwanensis ); C' is robust and taller than P* P? is small and in tooth row orslightly leaning inward; C, is weaker but still taller than P; and lower molars are strong, with high cusps, and are all myotodont. Chromosomal complement has 2n = 44 and FNa = 50 ( Guizhou , Henan , and Taiwan) . Habitat. Lowland and montane forest habitats, generally near water. Food and Feeding. The Fringed Long-footed Myotis forages by catching insects above the water surface and in open grasslands and farmlands. It flies slowly and is maneuverable. It feeds mostly on Diptera by aerial hawking but also occasionally gleans prey off surfaces. Breeding. Newborn Fringed Long-footed Myotis have been reported in November— June in Taiwan , suggesting a very long breeding season. Activity patterns. Fringed Long-footed Myotis primarily roost in caves. Calls are steep FM sweeps, with frequencies of 29-93 kHz on first harmonic and 70-122 kHz on second harmonic and mean duration of 10-6 milliseconds recorded in China . Movements, Home range and Social organization. Fringed Long-footed Myotis roost in large colonies; maternity colonies can be up to 1000 individuals in Taiwan . Status and Conservation. Classified as Least Concern on The IUCN Red Lust. Bibliography. Findley (1972), Han Naijian et al. (2010), Lin Liangkong, Motokawa & Harada (2002a), Liu Ying et al. (2003), Niu Hongxing, Yu Yan & Wang Yanmei (2007), Ruedi, Csorba et al. (2015), Ruedi, Stadelmann et al. (2013), Smith & Xie Yan (2008), Smith, Johnston, Jones & Rossiter ( 2008g ), Wang Hui et al. (2009), Zhang Zhenzhen et al. (2009).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis fimbriatus	Myotis	Unassigned-Myotis	fimbriatus	Peters	1871	1	In R. Swinhoe, Catalogue of Mammals of China, Proc. Zool. Soc. Lond.	1880:17:00	Fringed Long-footed Myotis	 hirsutus Howell, 1926;<b> taiwanensis </b>?rnb?ck-Christie-Linde, 1908.	China, Fujian, Amoy.	SE China, Taiwan.	Not listed.	Least Concern	Not conspecific with macrodactylus or capaccinii ; see Corbet (1978 c ) and Corbet and Hill (1992). Includes taiwanensis , see Ruedi et al. (2015).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis fimbriatus	23	Fringed Long-footed Myotis	Hairy-legged Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	fimbriatus	W. Peters	1871	1	Vespertilio_fimbriatus	Peters, W. C. H. (1871). In Swinhoe, R. Catalogue of the mammals of China (south of the River Yangtse) and of the Island of Formosa. Proceedings of the zoological Society of London, 1871 [for 1870], 617.	https://www.biodiversitylibrary.org/item/90543#page/707/mode/1up	ZMB 4148		"Amoy," Fujian, China.			fimbriatus (W. Peters, 1871)|taiwanensis Ärnbäck-Christie-Linde, 1908|hirsutus A. B. Howell, 1926	NA	NA	China|Taiwan	Asia	Palearctic	LC	0	0	0	Myotis_fimbriatus	0	sciname match	Myotis_fimbriatus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Myotis fimbriatus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	fimbriatus	(Peters, 1871)	Various authors have suggested relationships with ;M. pilosus , ;M. macrodactylus , and ;M. petax , but ;M. fimbriatus ;should be best considered a Chinese endemic species (Smith and Xie 2008, Ruedi et al. 2015).	20000000	Myotis fimbriatus	Least Concern		2019	2018-08-21 00:00:00 UTC	3.1	English	Although there are few data regarding this species, it is listed as Least Concern in view of its wide distribution, relatively large population, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	It is a cave-dwelling species of bat (Smith and Xie 2008). It presumably forages in the surrounding habitat. Its tolerance of disturbed habitats is not known.	At present there are no known major threats to this species.	The largest known population is of close to two hundred individuals in Shandong and Yunnan province, China (T. Jiang pers. comm.).	Unknown	Since the type of taiwanensis was considered as a valid subspecies of M. fimbriatus. The species is widespread and common. This Chinese endemic species has a wide distribution occurring in Hong Kong and the provinces of Sichuan, Yunnan, Guizhou, Fujian, Zhejiang, Anhui, Jiangsu, Jiangxi (Smith and Xie 2008), Taiwan, Shandong and Anhui (Han et al. 2010, Jiang et al. 2015).		Terrestrial	This species occurs in Qingliangfeng Nature Reserve (CSIS 2008), and is also likely to occur in Yangzie, Tianmushan, Laoshan, and An'jilongwangshan Nature Reserves and may be present in other protected areas. Research is needed in the areas of population status, biology and ecology, habitat status, and threats. In China, this species has been regionally Red Listed as Near Threatened, nearly meeting the listing for Vulnerable under criteria A4cd; B1ab(i,ii,iii) (Wang and Xie 2004, Jiang et al. 2016).	Indomalayan|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	fimbriatus	Peters	1871	1	In R. Swinhoe, Catalogue of Mammals of China, Proc. Zool. Soc. Lond.	1880:17:00	Fringed Long-footed Myotis	 hirsutus Howell, 1926;<b> taiwanensis </b>?rnb?ck-Christie-Linde, 1908.	China, Fujian, Amoy.	SE China, Taiwan.	Not listed.	Least Concern	Not conspecific with macrodactylus or capaccinii ; see Corbet (1978 c ) and Corbet and Hill (1992). Includes taiwanensis , see Ruedi et al. (2015).	Myotis fimbriatus	1005406	23	Fringed Long-footed Myotis	Hairy-legged Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	fimbriatus	W. Peters	1871	1	Vespertilio_fimbriatus	Peters, W. C. H. (1871). In Swinhoe, R. Catalogue of the mammals of China (south of the River Yangtse) and of the Island of Formosa. Proceedings of the zoological Society of London, 1871 [for 1870], 617.	https://www.biodiversitylibrary.org/item/90543#page/707/mode/1up	ZMB 4148		"Amoy," Fujian, China.			fimbriatus (W. Peters, 1871)|taiwanensis Ärnbäck-Christie-Linde, 1908|hirsutus A. B. Howell, 1926	NA	NA				China|Taiwan	Asia	Palearctic	LC	0	0	0	Myotis_fimbriatus	0	sciname match	Myotis_fimbriatus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_fimbriatus	1005406	23	Fringed Long-footed Myotis	Hairy-legged Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	fimbriatus	W. C. H. Peters in Swinhoe	1	Vespertilio fimbriatus	Swinhoe, R. 1871-04. Catalogue of the mammals of China (south of the river Yangtsze) and of the island of Formosa. Proceedings of the Zoological Society of London 1870(3):615-653.	https://www.biodiversitylibrary.org/page/28555479	ZMB 4148	holotype		"Amoy," Fujian, China.			NA	NA				China|Taiwan	Asia	Palearctic	LC	0	0	0	Myotis_fimbriatus	0	sciname match	Myotis_fimbriatus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	fimbriatus	Peters	1871	1	In R. Swinhoe, Catalogue of Mammals of China, Proc. Zool. Soc. Lond.	1880:17:00	Fringed Long-footed Myotis	hirsutus Howell, 1926; taiwanensis ?rnb?ck-Christie-Linde, 1908.	China, Fujian, Amoy.	SE China, Taiwan.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/85735587/22058886/' target='_blank'>Least Concern</a>	Not conspecific with macrodactylus or capaccinii; see Corbet (1978c) and Corbet and Hill (1992). Includes taiwanensis, see Ruedi et al. (2015).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis fimbriatus; Myotis fimbriatus; Myotis fimbriatus; Myotis fimbriatus; Myotis fimbriatus; Myotis fimbriatus; hirsutus; fimbriatus; taiwanensis; taiwanensis; hirsutus; fimbriatus; taiwanensis; hirsutus; Murin de Swinhoe; Haarige Wasserfledermaus; Ratonero de Swinhoe; Hairy-legged Myotis; Fringed Long-footed Myotis; Hairy-legged Myotis; Fringed Long-footed Myotis; Fringed Long-footed Myotis; M. fimbriatus