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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L863	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus	Myotis emarginatus		[MSW2] Subgenus Myotis.; [MSW3] Reviewed in part by Harrison and Bates (1991); also see Gaucher (1995) and Horácek et al. (2000). Apparently closely related to welwitschii; see Ruedi and Mayer (2001).; [HMW] Vespertilio emarginatus E. Geof: froy Saint-Hilaire, 1806 , “les souterrains des fortifications de Charlemont [= tunnels in the Fort of Charlemont],” Givet, Ardennes, northern France . Subgenus Chrysopteron. See M. tricolor . Populations of Central Asia are sometimes separated as race turcomanicus . Two subspecies recognized.; [batnames2022] Reviewed in part by Harrison and Bates (1991); also see Gaucher (1995) and Horácek et al. (2000). Apparently closely related to welwitschii ; see Ruedi and Mayer (2001). Records from Iraq are unvouchered (Al-Sheikhly et al., 2015).; [IUCN] The population of the western part of the distributional range (NW Africa, Europe, Caucasus and Levantine regions) is considered as being the nominotypic subspecies (M. emarginatus emarginatus ); in the Asian part of the range, one or two subspecies are differentiated, namely M. e. sogdianus and M. e. desertorum .; [batnames2023] Reviewed in part by Harrison and Bates (1991); also see Gaucher (1995) and Horácek et al. (2000). Apparently closely related to welwitschii ; see Ruedi and Mayer (2001). Records from Iraq are unvouchered (Al-Sheikhly et al., 2015).; [batnames2025_1.7] Reviewed in part by Harrison and Bates (1991); also see Gaucher (1995) and Horácek et al. (2000). Apparently closely related to welwitschii; see Ruedi and Mayer (2001). Records from Iraq are unvouchered (Al-Sheikhly et al., 2015).						budapestiensis, ciliatus, desertorum, lanaceus, neglectus, saturatus, turcomanicus.	emarginatus, desertorum, turcomanicus, saturatus	emarginatus, desertorum, turcomanicus	budapestiensis, ciliatus, kuzyakini, neglectus, rufescens, saturatus, schrankii; desertorum - lanaceus	emarginatus, desertorum		emarginatus, desertorum, turcomanicus	emarginatus - budapestiensis, ciliatus, kuzyakini, saturatus, neglectus, rufescens, saturatus, schrankii; desertorum - lanaceus	emarginatus, rufescens, ciliatus, schrankii, desertorum, budapestiensis, neglectus, lanaceus, turcomanicus, saturatus, kuzyakini	The population of the western part of the distributional range (NW Africa, Europe, Caucasus and Levantine regions) is considered as being the nominotypic subspecies (M. emarginatus emarginatus ); in the Asian part of the range, one or two subspecies are differentiated, namely M. e. sogdianus and M. e. desertorum .	emarginatus, desertorum, turcomanicus	emarginatus - budapestiensis, ciliatus, kuzyakini, neglectus, rufescens, saturatus, schrankii; desertorum - lanaceus	emarginatus, rufescens, ciliatus, schrankii, desertorum, budapestiensis, neglectus, lanaceus, turcomanicus, saturatus, kuzyakini 	emarginatus, rufescens, ciliatus, schrankii, desertorum, budapestiensis, neglectus, lanaceus, lanceus, turcomanicus, saturatus, kuzyakini	desertorum, emarginatus, turcomanicus	desertorum - lanaceus; emarginatus - budapestiensis, ciliatus, kuzyakini, neglectus, rufescens, saturatus, schrankii 	emarginatus (É. Geoffroy Saint-Hilaire, 1806)|rufescens (Crespon, 1844) [preoccupied]|ciliatus (J. H. Blasius, 1853)|schrankii (Kolenati, 1856) [nomen nudum | preoccupied]|desertorum (Dobson in Blanford, 1875)|budapestiensis Margó, 1880|neglectus (Fatio, 1890)|lanaceus Wroughton, 1920 [unjustified emendation]|lanceus O. Thomas, 1920|turcomanicus Bobrinski, 1925|saturatus Kuzyakin, 1934 [preoccupied]|kuzyakini Rossolimo & Pavlinov, 1979 [nomen novum]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Geoffroy's bat	SW Europe – Russian Turkestan, E Iran, Morocco	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis emarginatus	France, Ardennes, Givet, Charlemont.	E. Geoffroy	1806	Ann. Mus. Hist. Nat. Paris, 8:198.	Distribution: Ranging from the Netherlands, Portugal, and Morocco across southern Eurasia to Uz- bekistan and Afghanistan.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Geoffroy's bat	SW Europe – Turkestan, Afghanistan, N Africa	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	E. Geoffroy	1806	Ann. Mus. Hist. Nat. Paris, 8:198.	Subgenus Myotis.	S Europe, north to Netherlands and S Poland, Crimea, Caucasus and Kopet Dag Mtns, east to Uzbekistan and E Iran; Israel; Morocco; Algeria; Tunisia; Lebanon; Afghanistan.	France, Ardennes, Givet, Charlemont.		GEOFFROY	1806	Size medium (forearm length, 37-43 mm; condylobasal length, 14-16 mm). Ear medium in length. Braincase relatively high but rostrum fairly long and narrow. Nasal emargination medium. Middle upper premolar in toothrow. No fringe of hair on margin of uropatagium. Posterior border of ear pinna with a conspicuous angular emargination.	Distribution: Ranging from the Netherlands, Portugal, and Morocco across southern Eurasia to Uzbekistan and Afghanistan.	Four subspecies are here recognized:	M. e. emarginatus (Europe, northwestern Africa, and southwestern Asia), M. e. desertorum (Oman to Afghanistan), M. e. turcomanicus (Turkmenia to Afghanistan), M. e. saturatus (Uzbekistan).	102	species	M. emarginatus	GEOFFROY	1806	Myotis	subgenus	Myotis emarginatus				Size medium (forearm length, 37-43 mm; condylobasal length, 14-16 mm). Ear medium in length. Braincase relatively high but rostrum fairly long and narrow. Nasal emargination medium. Middle upper premolar in toothrow. No fringe of hair on margin of uropatagium. Posterior border of ear pinna with a conspicuous angular emargination.	Four subspecies are here recognized:		13. M. emarginatus (GEOFFROY 1806) [emarginatus group].	13	_M. e. desertorum_ (Dobson, 1875) (synonyms: _kuzyakini_ Россолимо & Павлинов, 1979, _lanceus_ Thomas, 1920, _saturatus_ Кузякин, 1934, _turcomanicus_ Бобринский, 1925); _M. e. emarginatus_ (Geoffroy Saint-Hilaire, 1806) (synonyms: _budapestiensis_ Margó, 1880, _ciliatus_ (Blasius, 1853), _neglectus_ (Fatio, 1890), _rufescens_ (Crespon, 1844), _schrankii_ (Kolenati, 1856))			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis emarginatus	Myotis		emarginatus	E. Geoffroy	y	1806		Ann. Mus. Natn. Hist. Nat. Paris	8		198		Geoffroy's Myotis	France, Ardennes, Givet, Charlemont.	S Europe, north to Netherlands and S Poland, Crimea, Caucasus and Kopet Dag Mtns, east and south to Israel, Jordan, Syria, Lebanon, Saudi Arabia, Oman, E Iran, Kyrgyzstan, Tajikistan, Uzbekistan, and Afghanistan; Morocco, Algeria, and Tunisia.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.	budapestiensis Margo, 1880; ciliatus Blasius, 1853; kuzyakini Pavilnov, 1979 [replacement name for saturatus Kuzyakin, 1934]; neglectus Fatio, 1890; rufescens Crespon 1844 [not Brehm, 1829]; saturatus Kuzyakin, 1934 [not Miller, 1897]; schrankii Kolenati, 1856 [nomen nudum; not schranki Wagner, 1843]; desertorum Dobson, 1875; lanaceus Thomas, 1920; turcomanicus Bobrinskii, 1925.	Reviewed in part by Harrison and Bates (1991); also see Gaucher (1995) and Horácek et al. (2000). Apparently closely related to welwitschii; see Ruedi and Mayer (2001).	4C3D87E8FF476AF9FA51956F1B1FB870	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	950	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF476AF9FA51956F1B1FB870.xml	Myotis emarginatus	Vespertilionidae	Myotis	emarginatus		1806	Murin a oreilles échancrées @fr | \Wimperfledermaus @de | Ratonero pardo @es | Geoffroy’s Bat @en	Vespertilio emarginatus E. Geof: froy Saint-Hilaire, 1806 , “les souterrains des fortifications de Charlemont [= tunnels in the Fort of Charlemont],” Givet, Ardennes, northern France . Subgenus Chrysopteron. See M. tricolor . Populations of Central Asia are sometimes separated as race turcomanicus . Two subspecies recognized.	M.e. emarginatus E. Geoffroy Saint-Hilaire, 1806 — Europe and N Africa, including most larger Mediterranean Is, E to Caucasus and Middle East. M.e. desertorum Dobson, 1875 — Asian part of range E of Caucasus.	Head-body 41-54 mm , tail 38-46 mm , ear 15-17 mm , hindfoot 9-11- 5 mm , forearm 36-1-44- 7 mm ; weight 5-5-15- 5 g . Fur of Geoffroy’s Myotisis long and woolly. Dorsal pelage is grayish brown to bright rusty brown (hairs tricolored, being straw yellow with gray bases and tips matching dorsal color). Ventral pelage is slightly to much paler than dorsum, ranging from yellowish gray to pale rufous (hairs bicolored, dark gray or dark grayish brown on basal one-half). Juveniles are darker and grayer with less conspicuously tricolored dorsal hairs. Muzzle is hairy and reddish to medium brown. Ears are grayish brown and relatively long, outer margin with conspicuous notch at about two-thirds from base; tragusis lancet-shaped with bluntly pointed tip, beingjust over one-half the ear height. Membranes are uniformly grayish brown and wings attach to base of outer toe; hindfeet are relatively short and calcaris straight, extending about halfway to tail tip. Skull is moderately sized and fairly delicate; braincase is high and forehead region is strongly concave;sagittal crest is absent anteriorly and weak posteriorly. Upper incisors are subequal in crown area to each other; P? is reduced but within tooth row. Chromosomal complement has 2n = 44 and FNa = 50 ( Spain ) or FNa = 52 (former Czechoslovakia and Greece ).	Usually reported from hardwood forests, with clear preference for deciduous trees and shrublands. It favors complex, structurally rich habitats, and positively selects riparian forests and low-vegetation ecosystems. It prefers sloping ground rather than the open spaces used by other species. It is commonly found feeding in more humanized habitats such as cattle sheds, orchards, or parks. Ranges from sea level to 1800 m .	Geoffroy’s Myotis feeds mainly on flies and spiders, with smaller amounts of Coleoptera and Hymenoptera . The species captures most of its prey by gleaning off surfaces; it tends to fly very close to ground, scanning for potential prey. It typically forages in rich shrublands and grasslands; also around cowsheds or sheep pens, where flies are abundant around cattle or sheep droppings.	Maternity colonies occur in a wide range of places such as caves, roofs of churches or houses, cattle sheds, and other buildings, usually less than 900 m above sea level. Maternity colonies tend to be densely clustered, mostly with adult females and young, and frequently mixed with other species. During maternity period, single individuals, mostly males, occupy rock holes and tree holes. Young are commonly born in early summer (earlyJune to mid-July), one per female, rarely twins. Lactating period lasts 25-35 days. Maternity colonies usually number 20-500 females and a few adult males. Females do not often mate during their first year, but a few do in their first autumn. Nursery roosts are frequently abandoned in August.	Being a cave-dwelling species, Geoffroy’s Myotis always occurs in habitats close to underground sites. After maternity period, it mainly occupies caves and mines or other underground sites for swarming. Roosts are commonly in group; the species tends to emerge from roosts ¢.15-20 minutes after sunset, and will fly up to 12: 5 km from the roost each night to visit hunting grounds, which can be as large as 50-70 ha; it can fly at up to 55 km /h. Caves and mines used as winter roosts usually have temperature fairly constant and often up to 13°C, much higher than in other similar species. The species can hibernate for long periods, to mid-April or even later. In winter,it is normally free-hanging, especially males that tend to roost solitarily. However, small clusters have also been found hibernating together with other species. Echolocation is characterized by short, highly modulated pulses, reaching values of 140 kHz at start of call, and ending at 38 kHz (broad frequency range), rarely below 30 kHz. These calls are typical for all Myotis . Pulses are always very short (1-3 milliseconds).	Longest known distance of migration is only 105 km , but few hibernation sites are known and longer distancesare to be expected. In some regions, the species disappears during winter and is not seen again until spring. During the swarming period,it forms groups mainly in caves and mines, generally dominated by males. Typically found roosting with other species such as Rhinolophus spp. and Schreibers’s Long-fingered Bat ( Miniopterus schreibersii ).	Classified as Least Concern on The [UCN Red List. In previous decades numbers declined, but the species has now clearly recovered and is even spreading to new regions. Cave-dwelling habits make the species vulnerable to disturbance, especially from leisure activities, and recently from fires and vandalism; itis also hunted for traditional medicine.	Aulagnier (2013k) | Dekker et al. (2013) | Fenton & Bogdanowicz (2002) | Findley (1972) | Flaquer et al. (2008) | Garcia & Arbona (2009) | Goiti et al. (2011) | Hutterer et al. (2005) | Kervyn et al. (2012) | Krull et al. (1991) | Piraccini ( 2016g) | Schumm et al. (1991) | Steck & Brinkmann (2006) | Volleth & Heller (2012) | Zahn et al. (2010)	https://zenodo.org/record/6398829/files/figure.png	425. Geoftroy’s Myotis Myotis emarginatus French: Murin a oreilles échancrées / German: \Wimperfledermaus / Spanish: Ratonero pardo Other common names: Geoffroy’s Bat Taxonomy. Vespertilio emarginatus E. Geof: froy Saint-Hilaire, 1806 , “les souterrains des fortifications de Charlemont [= tunnels in the Fort of Charlemont],” Givet, Ardennes, northern France . Subgenus Chrysopteron. See M. tricolor . Populations of Central Asia are sometimes separated as race turcomanicus . Two subspecies recognized. Subspecies and Distribution. M.e. emarginatus E. Geoffroy Saint-Hilaire, 1806 — Europe and N Africa, including most larger Mediterranean Is, E to Caucasus and Middle East. M.e. desertorum Dobson, 1875 — Asian part of range E of Caucasus. Descriptive notes. Head-body 41-54 mm , tail 38-46 mm , ear 15-17 mm , hindfoot 9-11- 5 mm , forearm 36-1-44- 7 mm ; weight 5-5-15- 5 g . Fur of Geoffroy’s Myotisis long and woolly. Dorsal pelage is grayish brown to bright rusty brown (hairs tricolored, being straw yellow with gray bases and tips matching dorsal color). Ventral pelage is slightly to much paler than dorsum, ranging from yellowish gray to pale rufous (hairs bicolored, dark gray or dark grayish brown on basal one-half). Juveniles are darker and grayer with less conspicuously tricolored dorsal hairs. Muzzle is hairy and reddish to medium brown. Ears are grayish brown and relatively long, outer margin with conspicuous notch at about two-thirds from base; tragusis lancet-shaped with bluntly pointed tip, beingjust over one-half the ear height. Membranes are uniformly grayish brown and wings attach to base of outer toe; hindfeet are relatively short and calcaris straight, extending about halfway to tail tip. Skull is moderately sized and fairly delicate; braincase is high and forehead region is strongly concave;sagittal crest is absent anteriorly and weak posteriorly. Upper incisors are subequal in crown area to each other; P? is reduced but within tooth row. Chromosomal complement has 2n = 44 and FNa = 50 ( Spain ) or FNa = 52 (former Czechoslovakia and Greece ). Habitat. Usually reported from hardwood forests, with clear preference for deciduous trees and shrublands. It favors complex, structurally rich habitats, and positively selects riparian forests and low-vegetation ecosystems. It prefers sloping ground rather than the open spaces used by other species. It is commonly found feeding in more humanized habitats such as cattle sheds, orchards, or parks. Ranges from sea level to 1800 m . Food and Feeding. Geoffroy’s Myotis feeds mainly on flies and spiders, with smaller amounts of Coleoptera and Hymenoptera . The species captures most of its prey by gleaning off surfaces; it tends to fly very close to ground, scanning for potential prey. It typically forages in rich shrublands and grasslands; also around cowsheds or sheep pens, where flies are abundant around cattle or sheep droppings. Breeding. Maternity colonies occur in a wide range of places such as caves, roofs of churches or houses, cattle sheds, and other buildings, usually less than 900 m above sea level. Maternity colonies tend to be densely clustered, mostly with adult females and young, and frequently mixed with other species. During maternity period, single individuals, mostly males, occupy rock holes and tree holes. Young are commonly born in early summer (earlyJune to mid-July), one per female, rarely twins. Lactating period lasts 25-35 days. Maternity colonies usually number 20-500 females and a few adult males. Females do not often mate during their first year, but a few do in their first autumn. Nursery roosts are frequently abandoned in August. Activity patterns. Being a cave-dwelling species, Geoffroy’s Myotis always occurs in habitats close to underground sites. After maternity period, it mainly occupies caves and mines or other underground sites for swarming. Roosts are commonly in group; the species tends to emerge from roosts ¢.15-20 minutes after sunset, and will fly up to 12: 5 km from the roost each night to visit hunting grounds, which can be as large as 50-70 ha; it can fly at up to 55 km /h. Caves and mines used as winter roosts usually have temperature fairly constant and often up to 13°C, much higher than in other similar species. The species can hibernate for long periods, to mid-April or even later. In winter,it is normally free-hanging, especially males that tend to roost solitarily. However, small clusters have also been found hibernating together with other species. Echolocation is characterized by short, highly modulated pulses, reaching values of 140 kHz at start of call, and ending at 38 kHz (broad frequency range), rarely below 30 kHz. These calls are typical for all Myotis . Pulses are always very short (1-3 milliseconds). Movements, Home range and Social organization. Longest known distance of migration is only 105 km , but few hibernation sites are known and longer distancesare to be expected. In some regions, the species disappears during winter and is not seen again until spring. During the swarming period,it forms groups mainly in caves and mines, generally dominated by males. Typically found roosting with other species such as Rhinolophus spp. and Schreibers’s Long-fingered Bat ( Miniopterus schreibersii ). Status and Conservation. Classified as Least Concern on The [UCN Red List. In previous decades numbers declined, but the species has now clearly recovered and is even spreading to new regions. Cave-dwelling habits make the species vulnerable to disturbance, especially from leisure activities, and recently from fires and vandalism; itis also hunted for traditional medicine. Bibliography. Aulagnier (2013k), Dekker et al. (2013), Fenton & Bogdanowicz (2002), Findley (1972), Flaquer et al. (2008), Garcia & Arbona (2009), Goiti et al. (2011), Hutterer et al. (2005), Kervyn et al. (2012), Krull et al. (1991), Piraccini ( 2016g ), Schumm et al. (1991), Steck & Brinkmann (2006), Volleth & Heller (2012), Zahn et al. (2010).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis emarginatus	Myotis	Chrysopteron	emarginatus	E. Geoffroy	1806	1	Ann. Mus. Natn. Hist. Nat. Paris	0.4708	Geoffroy's Myotis	 budapestiensis Margo, 1880; ciliatus Blasius, 1853; kuzyakini Pavilnov, 1979 [replacement name for saturatus Kuzyakin, 1934]; neglectus Fatio, 1890; rufescens Crespon 1844 [not Brehm, 1829]; saturatus Kuzyakin, 1934 [not Miller, 1897]; schrankii Kolenati, 1856 [ nomen nudum ; not schranki Wagner, 1843]; <b>desertorum</b> Dobson, 1875; lanaceus Thomas, 1920; <b>turcomanicus</b> Bobrinskii, 1925.	France, Ardennes, Givet, Charlemont.	S Europe, north to Netherlands and S Poland, Crimea, Caucasus and Kopet Dag Mtns, east and south to Israel, Jordan, Syria, Lebanon, Saudi Arabia, Oman, E Iran, Iraq, Kyrgyzstan, Tajikistan, Uzbekistan, and Afghanistan; Morocco, Algeria, and Tunisia.	Not listed.	Least Concern	Reviewed in part by Harrison and Bates (1991); also see Gaucher (1995) and Horácek et al. (2000). Apparently closely related to welwitschii ; see Ruedi and Mayer (2001). Records from Iraq are unvouchered (Al-Sheikhly et al., 2015).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis emarginatus	23	Geoffroy's Myotis	Geoffroy's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Chrysopteron	emarginatus	É. Geoffroy Saint-Hilaire	1806	1	Vespertilio_emarginatus	Geoffroy Saint-Hilaire, É. (1806). Mémoire sur le genre et les espèces de Vespertilion, l'un des genres de la famille des chauve-souris. Annales du Muséum d'histoire naturelle, 8, 198.	https://www.biodiversitylibrary.org/item/93165#page/201/mode/1up	MNHN 1997-1819		"les souterrains des fortifications de Charlemont [= tunnels in the Fort of Charlemont]," Givet, Ardennes, northern France.			emarginatus (É. Geoffroy Saint-Hilaire, 1806)|rufescens (Crespon, 1844) [preoccupied]|ciliatus (Blasius, 1853)|schrankii (Kolenati, 1856) [nomen nudum]|desertorum (Dobson, 1875)|budapestiensis Margó, 1880|neglectus (Fatio, 1890)|lanaceus O. Thomas, 1920|turcomanicus Bobrinski, 1925|saturatus Kuzyakin, 1934 [preoccupied]|kuzyakini Pavlinov, 1979	NA	NA	Morocco|Algeria|Tunisia|Portugal|Spain|France|Belgium|Luxembourg|Netherlands|Germany|Switzerland|Austria|Italy|Czech Republic|Slovakia|Poland|Hungary|Slovenia|Croatia|Bosnia & Herzegovina|Serbia|Kosovo|Montenegro|North Macedonia|Albania|Greece|Bulgaria|Romania|Moldova|Ukraine|Russia|Georgia|Armenia|Azerbaijan|Turkey|Cyprus|Syria|Lebanon|Israel|Palestine|Jordan|Iraq|Iran|Turkmenistan|Uzbekistan|Kazakhstan|Tajikistan|Kyrgyzstan|Afghanistan	Africa|Asia|Europe	Palearctic	LC	0	0	0	Myotis_emarginatus	0	sciname match	Myotis_emarginatus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14129	Myotis emarginatus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	emarginatus	(É. Geoffroy, 1806)	The population of the western part of the distributional range (NW Africa, Europe, Caucasus and Levantine regions) is considered as being the nominotypic subspecies (M. emarginatus emarginatus ); in the Asian part of the range, one or two subspecies are differentiated, namely M. e. sogdianus and M. e. desertorum .	20000000	Myotis emarginatus	Least Concern		2016	2016-04-25 00:00:00 UTC	3.1	English	The species experienced a significant decline in at least parts of its range from the 1960s to the 1990s, but now it is expanding in central Europe and is stable or not significantly declining elsewhere. The range is still large and the species is not specialized to restricted habitat, although it has a very specialized diet. Assessed as Least Concern.	Geoffroy's Bat (Myotis emarginatus ) has an unusual diet in that feeds mainly on spiders and flies. It forages over scrub and grassland but also in cow sheds where flies are abundant (Zahn et al. 2010, ;Dekker et al. 2013). In the Mediterranean Basin it has been reported to use ;also ;olive groves as foraging sites (Flaquer et al. 2008). In summer this bat roosts in underground habitats and in buildings (in attics), generally together with Rhinolophus species. It winters in underground sites. In Iran and the Caucasus, it occurs in a variety of habitats, but in low numbers (M. Sharifi pers. comm. 2005). Reportedly a sedentary species with movements of up to 105 km recorded (Schunger et al. 2004 in Hutterer et al. 2005), but may in fact move longer distances as winter roosts are not known in parts of its range where it occurs in summer.	In Europe the species is mainly associated with agricultural landscapes, therefore all agricultural activities can affect populations of this species. Loss of and disturbance to roost sites in buildings (including remedial timber treatment in attics) and underground sites are also threats. In the African part of the range, cave habitat where the species roosts is being destroyed by fires and vandalism. The species is also collected for traditional medicine practices in North Africa.	Locally it can be rare or common. The species experienced a significant decline from the 1960s to the 1990s, but in recent time the numbers in several regions have increased and the species has spread into new areas. It lives in large colonies (up to 1,200 individuals in Austria in one maternity colony).	Stable	Myotis emarginatus occurs in southern Europe from Portugal in the west to the Balkans in the east and southern part of western and central Europe and non-arid parts of south-western Asia from Asia Minor, Caucasus region and Palestine to Oman, Uzbekistan and Tajikistan; also in north-west Africa (recorded from northern Maghreb (Algeria, Morocco and Tunisia)). In the eastern part of the Mediterranean it occurs in various parts of Turkey, Syria, Lebanon and Israel. Recently it has been recorded in northern Iraq (Al-Sheikhly et al. 2015). It occurs from sea level to 1,800 m, highest records in the Alps are 812 m (maternity colony) and 1,505 m (hibernaculum) (Spitzenberger 2002).		Terrestrial	It is protected by national legislation in most range states. There are also international legal obligations for its protection through the Bonn Convention (Eurobats) and Bern Convention in the range states where these apply. It is included in Annex IV of EU Habitats and Species Directive, and there is some habitat protection through Natura 2000. Protection of roosts and promotion of awareness about the lack of medicinal value of the species is required.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Chrysopteron	emarginatus	É. Geoffroy Saint-Hilaire	1806	1	Ann. Mus. Natn. Hist. Nat. Paris	0.470833	Geoffroy's Myotis	 budapestiensis Margo, 1880; ciliatus Blasius, 1853; kuzyakini Pavilnov, 1979 [replacement name for saturatus Kuzyakin, 1934]; neglectus Fatio, 1890; rufescens Crespon 1844 [not Brehm, 1829]; saturatus Kuzyakin, 1934 [not Miller, 1897]; schrankii Kolenati, 1856 [ nomen nudum ; not schranki Wagner, 1843]; <b>desertorum</b> Dobson, 1875; lanaceus Thomas, 1920; <b>turcomanicus</b> Bobrinskii, 1925.	France, Ardennes, Givet, Charlemont.	S Europe, north to Netherlands and S Poland, Crimea, Caucasus and Kopet Dag Mtns, east and south to Israel, Jordan, Syria, Lebanon, Saudi Arabia, Oman, E Iran, Iraq, Kyrgyzstan, Tajikistan, Uzbekistan, and Afghanistan; Morocco, Algeria, and Tunisia.	Not listed.	Least Concern	Reviewed in part by Harrison and Bates (1991); also see Gaucher (1995) and Horácek et al. (2000). Apparently closely related to welwitschii ; see Ruedi and Mayer (2001). Records from Iraq are unvouchered (Al-Sheikhly et al., 2015).	Myotis emarginatus	1005402	23	Geoffroy's Myotis	Geoffroy's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Chrysopteron	emarginatus	É. Geoffroy Saint-Hilaire	1806	1	Vespertilio_emarginatus	Geoffroy Saint-Hilaire, É. (1806). Mémoire sur le genre et les espèces de Vespertilion, l'un des genres de la famille des chauve-souris. Annales du Muséum d'histoire naturelle, 8, 198.	https://www.biodiversitylibrary.org/item/93165#page/201/mode/1up	MNHN 1997-1819		"les souterrains des fortifications de Charlemont [= tunnels in the Fort of Charlemont]," Givet, Ardennes, northern France.			emarginatus (É. Geoffroy Saint-Hilaire, 1806)|rufescens (Crespon, 1844) [preoccupied]|ciliatus (Blasius, 1853)|schrankii (Kolenati, 1856) [nomen nudum]|desertorum (Dobson, 1875)|budapestiensis Margó, 1880|neglectus (Fatio, 1890)|lanaceus O. Thomas, 1920|turcomanicus Bobrinski, 1925|saturatus Kuzyakin, 1934 [preoccupied]|kuzyakini Pavlinov, 1979	NA	NA				Morocco|Algeria|Tunisia|Portugal|Spain|France|Belgium|Luxembourg|Netherlands|Germany|Switzerland|Austria|Italy|Czech Republic|Slovakia|Poland|Hungary|Slovenia|Croatia|Bosnia & Herzegovina|Serbia|Kosovo|Montenegro|North Macedonia|Albania|Greece|Bulgaria|Romania|Moldova|Ukraine|Russia|Georgia|Armenia|Azerbaijan|Turkey|Cyprus|Syria|Lebanon|Israel|Palestine|Jordan|Iraq|Iran|Turkmenistan|Uzbekistan|Kazakhstan|Tajikistan|Kyrgyzstan|Afghanistan	Africa|Asia|Europe	Palearctic	LC	0	0	0	Myotis_emarginatus	0	sciname match	Myotis_emarginatus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_emarginatus	1005402	23	Geoffroy's Myotis	Geoffroy's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Chrysopteron	emarginatus	É. Geoffroy Saint-Hilaire	1	Vespertilio emarginatus	Geoffroy Saint-Hilaire, É. 1806. Mémoire sur le genre et les espèces de Vespertilion, l'un des genres de la famille des chauve-souris. Annales du Muséum d'histoire naturelle 8:187-205.	https://www.biodiversitylibrary.org/page/29407868	MNHN-ZM-MO-1997-1819	syntypes	http://coldb.mnhn.fr/catalognumber/mnhn/zm/mo-1997-1819	"les souterrains des fortifications de Charlemont [= tunnels in the Fort of Charlemont]," Givet, Ardennes, northern France.			NA	NA				Morocco|Algeria|Tunisia|Portugal|Spain|France|Belgium|Luxembourg|Netherlands|Germany|Switzerland|Austria|Italy|Czech Republic|Slovakia|Poland|Hungary|Slovenia|Croatia|Bosnia and Herzegovina|Serbia|Kosovo|Montenegro|North Macedonia|Albania|Greece|Bulgaria|Romania|Moldova|Ukraine|Russia|Georgia|Armenia|Azerbaijan|Turkey|Cyprus|Syria|Lebanon|Israel|Palestine|Jordan|Iraq|Iran|Turkmenistan|Uzbekistan|Kazakhstan|Tajikistan|Kyrgyzstan|Afghanistan	Africa|Asia|Europe	Palearctic	LC	0	0	0	Myotis_emarginatus	0	sciname match	Myotis_emarginatus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Chrysopteron	emarginatus	É. Geoffroy Saint-Hilaire	1806	1	Ann. Mus. Natn. Hist. Nat. Paris	0.470833	Geoffroy's Myotis	budapestiensis Margo, 1880; ciliatus Blasius, 1853; kuzyakini Pavilnov, 1979 [replacement name for saturatus Kuzyakin, 1934]; neglectus Fatio, 1890; rufescens Crespon 1844 [not Brehm, 1829]; saturatus Kuzyakin, 1934 [not Miller, 1897]; schrankii Kolenati, 1856 [nomen nudum; not schranki Wagner, 1843]; desertorum Dobson, 1875; lanaceus Thomas, 1920; turcomanicus Bobrinskii, 1925.	France, Ardennes, Givet, Charlemont.	S Europe, north to Netherlands and S Poland, Crimea, Caucasus and Kopet Dag Mtns, east and south to Israel, Jordan, Syria, Lebanon, Saudi Arabia, Oman, E Iran, Iraq, Kyrgyzstan, Tajikistan, Uzbekistan, and Afghanistan; Morocco, Algeria, and Tunisia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14129/22051191/' target='_blank'>Least Concern</a>	Reviewed in part by Harrison and Bates (1991); also see Gaucher (1995) and Horácek et al. (2000). Apparently closely related to welwitschii; see Ruedi and Mayer (2001). Records from Iraq are unvouchered (Al-Sheikhly et al., 2015).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis emarginatus; Myotis emarginatus; Myotis emarginatus; Myotis emarginatus; Myotis emarginatus; Myotis emarginatus; emarginatus; desertorum; turcomanicus; budapestiensis; ciliatus; kuzyakini; neglectus; rufescens; saturatus; schrankii; desertorum - lanaceus; emarginatus; desertorum; desertorum; turcomanicus; budapestiensis; ciliatus; kuzyakini; saturatus; neglectus; rufescens; saturatus; schrankii; desertorum - lanaceus; emarginatus; rufescens; ciliatus; schrankii; desertorum; budapestiensis; neglectus; lanaceus; turcomanicus; saturatus; kuzyakini; Murin a oreilles échancrées; \Wimperfledermaus; Ratonero pardo; Geoffroy’s Bat; Geoffroy's Myotis; Geoffroy's Bat; Geoffroy's Myotis; Geoffroy's Myotis; M. emarginatus