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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L87	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus	Aselliscus stoliczkanus		[MSW3] See Sanborn (1952b).; [HMW] Asellia stoliczkana Dobson, 1871 , Penang Island, Malaysia . It is suspected that this could be a species complex: further genetic and morphological research is needed. Taxa trifidus and wheeleri are here treated as synonyms. Monotypic.; [batnames2022] Tu et al. (2015) describe molecular support for recognition of both wheeler and trifidus at the species level, pending further investigation. See Sanborn (1952 b ).; [IUCN] This could represent a species complex, further morphological and genetic studies are required (C. Francis pers. comm.).; [batnames2023] Tu et al. (2015) describe molecular support for recognition of both wheeler and trifidus at the species level, pending further investigation. See Sanborn (1952 b ).; [batnames2025_1.7] Tu et al. (2015) describe molecular support for recognition of both wheeler and trifidus at the species level, pending further investigation. See Sanborn (1952b).				trifidus, wheeleri		trifidus, wheeleri (see Sanborn, 1952b:3).			trifidus, wheeleri		trifidus, wheeleri	stoliczkanus	stoliczkanus - trifidus, wheeleri	stoliczkanus, trifidus, wheeleri	This could represent a species complex, further morphological and genetic studies are required (C. Francis pers. comm.).	stoliczkanus	stoliczkanus - trifidus, wheeleri	stoliczkanus, trifidus, wheeleri 	stoliczkanus, trifidus, wheeleri 	stoliczkanus	stoliczkanus - trifidus, wheeleri	stoliczkanus (Dobson, 1871)|trifidus (W. C. H. Peters, 1871)|wheeleri (Osgood, 1932)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Stoliczka's trident bat	Burma, S China – Indochina, Penang I	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Aselliscus stoliczkanus	Malaysia, Penang.	Dobson	1871	Proc. Asiat. Soc. Bengal, p. 106.	Distribution: Confined to southeastern Asia from Burma and southern China south to Malaya.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Stoliczka's trident bat	Burma, S China – Indochina, Penang I	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Dobson	1871	Proc. Asiat. Soc. Bengal, p. 106.		Burma, S China, Thailand, Laos, Vietnam, W Malaysia.	Malaysia, West, Penang.		DOBSON	1871	Three lateral leaflets. Tail relatively long. More sloping ros trum. Anterior lower premolar compressed by ca nine and posterior premolar. Size small (forearm length, 39-44 mm).	Distribution: Confined to southeastern Asia from Burma and southern China south to Malaya.	No subspecies (SAN BORN 1952).		67	species	A. stoliczkanus	DOBSON	1871	Aselliscus	genus	Aselliscus stoliczkanus				Three lateral leaflets. Tail relatively long. More sloping ros trum. Anterior lower premolar compressed by ca nine and posterior premolar. Size small (forearm length, 39-44 mm).	No subspecies (SAN BORN 1952).		2. A. stoliczkanus (DOBSON 1871) ( = trifida PETERS, 1871; wheeleriOSGOOD 1932).	2	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Aselliscus stoliczkanus	Aselliscus		stoliczkanus	Dobson	y	1871		Proc. Asiat. Soc. Bengal			106		Stoliczka's Asian Trident Bat	Malaysia, West, Penang Isl.	Burma, S China, Thailand, Laos, Vietnam, W Malaysia.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	trifidus Peters, 1871; wheeleri Osgood, 1932.	See Sanborn (1952b).	03BD87A2C671A203FF38F106F4624B76	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	234	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C671A203FF38F106F4624B76.xml	Aselliscus stoliczkanus	Hipposideridae	Aselliscus	stoliczkanus		1871	Phyllorhine de Stoliczka @fr | Stoliczka-Dreizackblattnase @de | Aseliseo de Stoliczka @es | Stoliczka's Asian Trident Bat @en	Asellia stoliczkana Dobson, 1871 , Penang Island, Malaysia . It is suspected that this could be a species complex: further genetic and morphological research is needed. Taxa trifidus and wheeleri are here treated as synonyms. Monotypic.	From NE Myanmar and S China S to S Thailand , C Laos , and C Vietnam ; also on Penang and Tioman Is.	Head-body 40-50 mm, tail 33-44 mm, ear 10—13 mm , hindfoot 9—10 mm , forearm 39-45-6 mm; weight 6-9 g. As in its sister species, Temminck’s Trident Bat (A. tricuspidatus ), Stoliczka’s Trident Bat has a tridentate noseleaf margin in posterior leaf. Intermediate point of trident is separated from lateral ones by large grooves. Stoliczka’s Trident Bat has slightly shorter forearm and tail than Temminck’s Trident Bat. Pelage is reddish brown on dorsum and grayish or white-gray in ventral part.Juveniles can be darker than adults. Chromosomal complement has 2n = 30 and FN = 56.	Stoliczka’s Trident Bat occurs at low densities in limestone areas, often roosting in smaller caves than other bat species, with high levels of humidity to maintain water balance. It has been reported not only in primary forest, but also in agricultural lands and other human-degraded habitats.	Stoliczka’s Trident Bat forages both in cluttered microhabitats and strongly disturbed areas. Its diet is based predominantly on insects, especially lepidopterans, hemipterans, beetles, and odonates, varying depending on the seasonal availability of prey. Its flight is slow, allowing a high capacity of maneuver, especially when hunting in closed environments. Due to its small size, predation by other bat species might occur.	Reproductive females have been captured in May-June in Laos and Vietnam .	During winter, Stoliczka’s Trident Bat hibernates in longer caves, which provide more stable microclimates with relatively low temperatures. Echolocation call frequency is c.120-125 kHz.	Stoliczka’s Trident Bats have been observed roosting in groups of 10—20 individuals, and sometimes separated from each other by few centimeters. They can share roosts with other bat species in larger caves as compared with single-species caves.	Classified as Least Concern on The IUCN Red List. Population size and trends of Stoliczka’s Trident Bat are still unknown, but population is probably not greatly fragmented and it is not declining markedly. Despite its generally low densities, this species can be locally common; it is present in several protected areas. There are no major threats for this species, but habitat destruction due to limestone mining and disturbance of roosts by tourism are increasingly affecting these animals.	Bates, Bumrungsri, Francis, Csorba & Furey (2008) | Bogdanowicz & Owen (1998) | Li Gang et al. (2007) | Tu Vuong Tan, Csorba et al. (2015) | Zhang Zongxiao et al. (2016)	https://zenodo.org/record/3749075/files/figure.png	19. Stoliczka’s Trident Bat Aselliscus stoliczkanus French: Phyllorhine de Stoliczka / German: Stoliczka-Dreizackblattnase / Spanish: Aseliseo de Stoliczka Other common names: Stoliczka's Asian Trident Bat Taxonomy. Asellia stoliczkana Dobson, 1871 , Penang Island, Malaysia . It is suspected that this could be a species complex: further genetic and morphological research is needed. Taxa trifidus and wheeleri are here treated as synonyms. Monotypic. Distribution. From NE Myanmar and S China S to S Thailand , C Laos , and C Vietnam ; also on Penang and Tioman Is. Descriptive notes. Head-body 40-50 mm, tail 33-44 mm, ear 10—13 mm , hindfoot 9—10 mm , forearm 39-45-6 mm; weight 6-9 g. As in its sister species, Temminck’s Trident Bat (A. tricuspidatus ), Stoliczka’s Trident Bat has a tridentate noseleaf margin in posterior leaf. Intermediate point of trident is separated from lateral ones by large grooves. Stoliczka’s Trident Bat has slightly shorter forearm and tail than Temminck’s Trident Bat. Pelage is reddish brown on dorsum and grayish or white-gray in ventral part.Juveniles can be darker than adults. Chromosomal complement has 2n = 30 and FN = 56. Habitat. Stoliczka’s Trident Bat occurs at low densities in limestone areas, often roosting in smaller caves than other bat species, with high levels of humidity to maintain water balance. It has been reported not only in primary forest, but also in agricultural lands and other human-degraded habitats. Food and Feeding. Stoliczka’s Trident Bat forages both in cluttered microhabitats and strongly disturbed areas. Its diet is based predominantly on insects, especially lepidopterans, hemipterans, beetles, and odonates, varying depending on the seasonal availability of prey. Its flight is slow, allowing a high capacity of maneuver, especially when hunting in closed environments. Due to its small size, predation by other bat species might occur. Breeding. Reproductive females have been captured in May-June in Laos and Vietnam . Activity patterns. During winter, Stoliczka’s Trident Bat hibernates in longer caves, which provide more stable microclimates with relatively low temperatures. Echolocation call frequency is c.120-125 kHz. Movements, Home range and Social organization. Stoliczka’s Trident Bats have been observed roosting in groups of 10—20 individuals, and sometimes separated from each other by few centimeters. They can share roosts with other bat species in larger caves as compared with single-species caves. Status and Conservation. Classified as Least Concern on The IUCN Red List. Population size and trends of Stoliczka’s Trident Bat are still unknown, but population is probably not greatly fragmented and it is not declining markedly. Despite its generally low densities, this species can be locally common; it is present in several protected areas. There are no major threats for this species, but habitat destruction due to limestone mining and disturbance of roosts by tourism are increasingly affecting these animals. Bibliography. Bates, Bumrungsri, Francis, Csorba & Furey (2008), Bogdanowicz & Owen (1998), Li Gang et al. (2007), Tu Vuong Tan, Csorba et al. (2015), Zhang Zongxiao et al. (2016).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Aselliscus stoliczkanus	Aselliscus		stoliczkanus	Dobson	1871	1	Proc. Asiat. Soc. Bengal	1872:46:00	Stoliczka's Asian Trident Bat	 trifidus Peters, 1871; wheeleri Osgood, 1932.	Malaysia, West, Penang Isl.	Burma, S China, Thailand, Laos, Vietnam, W Malaysia.	Not listed.	Least Concern	Tu et al. (2015) describe molecular support for recognition of both wheeler and trifidus at the species level, pending further investigation. See Sanborn (1952 b ).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Aselliscus stoliczkanus	23	Stoliczka's Trident Bat	Stoliczka's Asian Trident Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Aselliscus	NA	stoliczkanus	Dobson	1871	1						Penang Island, Malaysia.			stoliczkanus (Dobson, 1871)|trifidus (W. Peters, 1871)|wheeleri (Osgood, 1932)	NA	NA	Myanmar|China|Thailand|Laos|Vietnam|Malaysia	Asia	Indomalaya|Palearctic	LC	0	0	0	Aselliscus_stoliczkanus	0	sciname match	Aselliscus_stoliczkanus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	2155	Aselliscus stoliczkanus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Aselliscus	stoliczkanus	(Dobson, 1871)	This could represent a species complex, further morphological and genetic studies are required (C. Francis pers. comm.).	9000000	Aselliscus stoliczkanus	Least Concern		2008	2008-06-30 00:00:00 UTC	3.1	English	Listed as Least Concern as the species is locally common where it occurs and is widespread. It occurs in a number of protected areas.	The species occurs in limestone areas and is known to roost in caves (C. Francis and P. Bates pers. comm.). The species is not dependent on primary habitat and can survive in degraded areas and extends into agricultural zones and shares roosts with other species (P. Bates pers. comm.). A colony in Thailand roosted in loose association in limestone caves, with individuals at least 30 cm from one another. Reproductively active females have been collected in May and June in Lao PDR and Viet Nam.	The species is vulnerable to disturbance of cave roosts by tourism and also the destruction of caves through limestone extraction (P. Bates pers. comm.).	It is locally common in limestone areas.	Unknown	This species occurs from Myanmar (Struebig et al. 2005) and southern China (Yunnan, Guangxi and Guizhou) (Smith et al. 2008), south to the Malay Peninsula.		Terrestrial	The species occurs in a number of protected areas in Thailand, Lao PDR and Viet Nam.	Indomalayan|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Aselliscus		stoliczkanus	Dobson	1871	1	Proc. Asiat. Soc. Bengal	1872:46:00	Stoliczka's Asian Trident Bat	 trifidus Peters, 1871; wheeleri Osgood, 1932.	Malaysia, West, Penang Isl.	Burma, S China, Thailand, Laos, Vietnam, W Malaysia.	Not listed.	Least Concern	Tu et al. (2015) describe molecular support for recognition of both wheeler and trifidus at the species level, pending further investigation. See Sanborn (1952 b ).	Aselliscus stoliczkanus	1004561	23	Stoliczka's Trident Bat	Stoliczka's Asian Trident Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Aselliscus	NA	stoliczkanus	Dobson	1871	1						Penang Island, Malaysia.			stoliczkanus (Dobson, 1871)|trifidus (W. Peters, 1871)|wheeleri (Osgood, 1932)	NA	NA				Myanmar|China|Thailand|Laos|Vietnam|Malaysia	Asia	Indomalaya|Palearctic	LC	0	0	0	Aselliscus_stoliczkanus	0	sciname match	Aselliscus_stoliczkanus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Aselliscus_stoliczkanus	1004561	23	Stoliczka's Trident Bat	Stoliczka's Asian Trident Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Aselliscus	NA	stoliczkanus	Dobson	1	Asellia stoliczkana	Dobson, G.E. 1871. Description of four new species of Malayan bats, from the collection of Dr. Stoliczka. Journal of the Asiatic Society of Bengal 40(3):260-267.	https://www.biodiversitylibrary.org/page/35631127	ZSI 8946	holotype		Penang Island, Malaysia.			NA	NA				Myanmar|China|Thailand|Laos|Vietnam|Malaysia	Asia	Indomalaya|Palearctic	LC	0	0	0	Aselliscus_stoliczkanus	0	sciname match	Aselliscus_stoliczkanus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Aselliscus		stoliczkanus	Dobson	1871	1	Proc. Asiat. Soc. Bengal	1872:46:00	Stoliczka's Asian Trident Bat	trifidus Peters, 1871; wheeleri Osgood, 1932.	Malaysia, West, Penang Isl.	Burma, S China, Thailand, Laos, Vietnam, W Malaysia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/214518902/21976509/' target='_blank'>Least Concern</a>	Tu et al. (2015) describe molecular support for recognition of both wheeler and trifidus at the species level, pending further investigation. See Sanborn (1952b).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Aselliscus stoliczkanus; Aselliscus stoliczkanus; Aselliscus stoliczkanus; Aselliscus stoliczkanus; Aselliscus stoliczkanus; Aselliscus stoliczkanus; trifidus; wheeleri; trifidus; wheeleri; trifidus; wheeleri; stoliczkanus; trifidus; wheeleri; Phyllorhine de Stoliczka; Stoliczka-Dreizackblattnase; Aseliseo de Stoliczka; Stoliczka's Asian Trident Bat; Stoliczka's Trident Bat; Stoliczka's Asian Trident Bat; Stoliczka's Asian Trident Bat; Stoliczka's Asian Trident Bat; A. stoliczkanus