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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L854	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis daubentoni	Myotis daubentoni	Myotis daubentonii	Myotis daubentoni	Myotis daubentoni	Myotis daubentonii	Myotis daubentonii	Myotis daubentonii	Myotis daubentonii	Myotis daubentonii	Myotis daubentonii	Myotis daubentonii	Myotis daubentonii	Myotis daubentonii	Myotis daubentonii		[MSW2] Subgenus Leuconoe. Includes laniger; see Ellerman and Morrison-Scott (1951:147). Includes nathalinae; see Hor4£ek and Hanak (1984).; [MSW3] Includes nathalinae; see Horácek and Hanák (1984), Fairon (1985), Mayer and von Helversen (2001a), and Ruedi and Mayer (2001). Reviewed in part by Yoshiyuki (1989), Yoon (1990), Bates and Harrison (1997), Bates et al. (1999), and Horácek et al. (2000). Does not appear to include laniger; see Topál (1997) and Bates et al. (1999), though also see Corbet and Hill (1992). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Bogdanowicz (1994), but note that laniger was included in daubentonii in that publication. Apparently closely related to bechsteinii; see Ruedi and Mayer (2001). Subspecies limits are problematic, see Bogdanowicz (1994), Horácek et al. (2000), and Kruskop (2002). Genetic studies suggest that this complex includes more than one species, with at least some Russian and Japanese specimens representing a taxon distinct from the European form (Kawai et al., 2003).; [HMW] Vespertilio daubentonii Kuhl, 1817 , Hanau, Hessen , Germany . Subgenus Myotis ; daubentonii species group. Phylogenetic studies in 2013 and 2015 recovered M. daubentonii in a clade with M. sicarius , M. bechsteini , M. longicaudatus , M. soror, and M. frater . Monotypic.; [batnames2022] Does not include abei or petax ( petax is the senior synonym of abei ); see Matveev et al. (2005), Kruskop et al. (2012), and Katzmann et al. (2012). Does not include nathalinae ; see Horácek and Hanák (1984), Fairon (1985), Mayer and von Helversen (2001 a ), and Ruedi and Mayer (2001). Reviewed in part by Yoshiyuki (1989), Yoon (1990), Bates and Harrison (1997), Bates et al. (1999), andHorácek et al. (2000). Does not appear to include laniger ; see Topál (1997) and Bates et al. (1999), though also see Corbet and Hill(1992). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Bogdanowicz (1994), but note that laniger was includedin daubentonii in that publication. Apparently closely related to bechsteinii ; see Ruedi and Mayer (2001). Subspecies limitsare problematic, see Bogdanowicz (1994), Horácek et al. (2000), and Kruskop (2002). Genetic studies suggest that this complex includes morethan one species, with at least some Russian and Japanese specimens representing a taxon distinct from the European form (Kawai et al., 2003).; [MDD2022] previously included M. petax (and abei); [IUCN] <span>This is the new taxonomic concept for Myotis daubentonii . Previously the assessment referred to M. daubentonii sensu lato, thus including M. petax from Altai and Siberia which is now considered a distinct species (Matveev et al. 2005).; [batnames2023] Does not include abei or petax ( petax is the senior synonym of abei ); see Matveev et al. (2005), Kruskop et al. (2012), and Katzmann et al. (2012). Does not include nathalinae ; see Horácek and Hanák (1984), Fairon (1985), Mayer and von Helversen (2001 a ), and Ruedi and Mayer (2001). Reviewed in part by Yoshiyuki (1989), Yoon (1990), Bates and Harrison (1997), Bates et al. (1999), andHorácek et al. (2000). Does not appear to include laniger ; see Topál (1997) and Bates et al. (1999), though also see Corbet and Hill(1992). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Bogdanowicz (1994), but note that laniger was includedin daubentonii in that publication. Apparently closely related to bechsteinii ; see Ruedi and Mayer (2001). Subspecies limitsare problematic, see Bogdanowicz (1994), Horácek et al. (2000), and Kruskop (2002). Genetic studies suggest that this complex includes morethan one species, with at least some Russian and Japanese specimens representing a taxon distinct from the European form (Kawai et al., 2003).; [MDD2023] previously included M. petax (and abei); [MDD2025_2.0] previously included M. petax (and abei); [batnames2025_1.7] Does not include abei or petax (petax is the senior synonym of abei); see Matveev et al. (2005), Kruskop et al. (2012), and Katzmann et al. (2012). Does not include nathalinae; see Horácek and Hanák (1984), Fairon (1985), Mayer and von Helversen (2001 a), and Ruedi and Mayer (2001). Reviewed in part by Yoshiyuki (1989), Yoon (1990), Bates and Harrison (1997), Bates et al. (1999), andHorácek et al. (2000). Does not appear to include laniger; see Topál (1997) and Bates et al. (1999), though also see Corbet and Hill(1992). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Bogdanowicz (1994), but note that laniger was includedin daubentonii in that publication. Apparently closely related to bechsteinii; see Ruedi and Mayer (2001). Subspecies limitsare problematic, see Bogdanowicz (1994), Horácek et al. (2000), and Kruskop (2002). Genetic studies suggest that this complex includes morethan one species, with at least some Russian and Japanese specimens representing a taxon distinct from the European form (Kawai et al., 2003).; [MDD2025_2.2] previously included M. petax (and abei)				laniger	(petax) (nathalinae)	aedilus, albus, capucinellus, lanatus, laniger, loukashkini, minutellus, nathalinae, petax, staufferi, ussuriensis, volgensis.	nathalinae, daubentoni, volgensis, ussuriensis, petax, laniger	daubentonii, chasanensis, loukashkini, nathalinae, petax, ussuriensis, volgensis	aedilus, albus, capucinellus, lanatus, minutellus, staufferi			daubentonii, chasanensis, loukashkini, nathalinae, ussuriensis, volgensis	daubentonii - aedilus, albus, capucinellus, lanatus, minutellus, staufferi	daubentonii, aedilus, volgensis, lanatus, albus, capucinellus, minutellus, staufferi, nathalinae	<span>This is the new taxonomic concept for Myotis daubentonii . Previously the assessment referred to M. daubentonii sensu lato, thus including M. petax from Altai and Siberia which is now considered a distinct species (Matveev et al. 2005).	daubentonii, chasanensis, loukashkini, nathalinae, ussuriensis, volgensis	daubentonii - aedilus, albus, capucinellus, lanatus, minutellus, staufferi	daubentonii, aedilus, volgensis, lanatus, albus, capucinellus, minutellus, staufferi, nathalinae	daubentonii, aedilis, volgensis, lanatus, albus, capucinellus, minutellus, staufferi, daubentoni, nathalinae, aedilus	chasanensis, daubentonii, loukashkini, nathalinae, ussuriensis, volgensis	daubentonii - aedilus, albus, capucinellus, lanatus, minutellus, staufferi 	daubentonii (Kuhl, 1817)|aedilis (Jenyns, 1839)|daubentoni (von Nordmann, 1839) [incorrect subsequent spelling]|volgensis (Eversmann, 1840)|lanatus (Crespon, 1844)|albus (Fitzinger, 1871) [nomen novum]|capucinellus (Fitzinger, 1871) [not used as valid]|minutellus (Fitzinger, 1871) [not used as valid]|staufferi (Fatio, 1890)|nathalinae Tupinier, 1977|aedilus Simmons, 2005 [incorrect subsequent spelling | not used as valid]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Daubenton's bat	Spain, Britain – E Siberia Manchuria, Sakhalin, Hokkaido	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis daubentoni	Germany, Hessen, Hanau.	Kuhl	1819	Ann. Wetterau Ges. Naturk., 4(2):195.	Distribution: Ranging from western Europe to eastern Siberia and south to southern China and northeastern India, including Sakhalin, Kuriles, and Hokkaido in Japan.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Daubenton's bat	Portugal, Britain – E Siberia, Manchuria, Sakhalin, Japan; ref. 4.64	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Kuhl	1817	Die Deutschen Fledermause. Hanau, p. 14.	Subgenus Leuconoe. Includes laniger; see Ellerman and Morrison-Scott (1951:147). Includes nathalinae; see Hor4£ek and Hanak (1984).	Europe east to Kamtschatka, Vladivostok, Sakhalin and Kurile Isis (Russia); Japan; Korea; Manchuria, E and S China; Britain and Ireland; Scandinavia; Assam (India).	Germany, Hessen, Hanau.		KUHL	1819	Size medium to fairly small (forearm length, 35-39 mm; condylobasal length, 12-15 mm). Margin of plagiopatagium attached to side of foot near the ankle. Margin of uropatagium without a fringe of hair. Braincase relatively low without a sagittal crest. Rostrum relatively slender. Middle upper premolar in toothrow.	Distribution: Ranging from western Europe to eastern Siberia and south to southern China and northeastern India, including Sakhalin, Kuriles, and Hokkaido in Japan.	Six subspecies are here recognized:	M. d. nathalinae (southwestern Europe), M. d. daubentoni (northwestern Europe), M. d. volgensis (eastern Europe to central Siberia), M. d. ussuriensis (eastern Siberia and northeastern China to Japan and the Kuriles), M. d. petax (Altai region of Siberia and Mongolia), M. d. laniger (southern China and northeastern India).	107	species	M. daubentoni	KUHL	1819	Leuconoe	subgenus	Myotis daubentoni				Size medium to fairly small (forearm length, 35-39 mm; condylobasal length, 12- 15 mm). Margin of plagiopatagium attached to side of foot near the ankle. Margin of uropatagium without a fringe of hair. Braincase relatively low without a sagittal crest. Rostrum relatively slender. Middle upper premolar in toothrow.	Six subspecies are here recognized:		67. M. daubentoni (KUHL 1819) [daubentoni group],	67	_M. d. daubentonii_ (Kuhl, 1817) (synonyms: _aedilis_ (Jenyns, 1839), _albus_ (Fitzinger, 1871), _capucinellus_ (Fitzinger, 1871), _lanatus_ (Crespon, 1844), _minutellus_ (Fitzinger, 1871), _staufferi_ (Fatio, 1890)); _M. d. nathalinae_ Tupinier, 1977; _M. d. volgensis_ (Eversmann, 1840)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis daubentonii	Myotis		daubentonii	Kuhl	y	1817		Die Deutschen Fledermäuse. Hanau			14		Daubenton's Myotis	Germany, Hessen, Hanau.	Europe (including Britain and Ireland; Scandinavia) east to Kamtschatka, Vladivostok, Sakhalin and Kurile Isls (Russia), Japan, Korea, Manchuria, N and E China (including Tibet), Vietnam.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	aedilus Jenyns, 1839; albus Fitzinger, 1871; capucinellus Fitzinger, 1871; lanatus Crespon, 1844; minutellus Fitzinger, 1871; staufferi Fatio, 1890; chasanensis Tiunov, 1997; loukashkini Shamel, 1942; nathalinae Tupinier, 1977; petax Hollister, 1912; ussuriensis Ognev, 1927; volgensis Eversmann, 1840.	Includes nathalinae; see Horácek and Hanák (1984), Fairon (1985), Mayer and von Helversen (2001a), and Ruedi and Mayer (2001). Reviewed in part by Yoshiyuki (1989), Yoon (1990), Bates and Harrison (1997), Bates et al. (1999), and Horácek et al. (2000). Does not appear to include laniger; see Topál (1997) and Bates et al. (1999), though also see Corbet and Hill (1992). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Bogdanowicz (1994), but note that laniger was included in daubentonii in that publication. Apparently closely related to bechsteinii; see Ruedi and Mayer (2001). Subspecies limits are problematic, see Bogdanowicz (1994), Horácek et al. (2000), and Kruskop (2002). Genetic studies suggest that this complex includes more than one species, with at least some Russian and Japanese specimens representing a taxon distinct from the European form (Kawai et al., 2003).	4C3D87E8FF2D6A92FF4F978916A7B9F2	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	976	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF2D6A92FF4F978916A7B9F2.xml	Myotis daubentonii	Vespertilionidae	Myotis	daubentonii		1817	Murin de Daubenton @fr | \Wasserfledermaus @de | Ratonero ribereno @es | Daubenton’s Bat @en	Vespertilio daubentonii Kuhl, 1817 , Hanau, Hessen , Germany . Subgenus Myotis ; daubentonii species group. Phylogenetic studies in 2013 and 2015 recovered M. daubentonii in a clade with M. sicarius , M. bechsteini , M. longicaudatus , M. soror, and M. frater . Monotypic.	Most of Europe and W Asia, from S Fennoscandia, British Is, and Iberian Peninsula E to W Russia (E to Omsk region ), the Caucasus, and N Turkey ; disrupted distribution in Mediterranean areas.	Head-body 45-55 mm , tail 27-50 mm , ear 9-14 mm , hindfoot 9-11 mm , forearm 33-42 mm ; weight 6-10 g . Daubenton’s Myotis has grayish or brownish dorsal and ventral hairs, with dark bases and more brighttips, sometimes reddish. Wings are generally dark brown or gray. Young are generally darker than adults. Plagiopatagium starts at base of fingers, Uropatagium is naked; venter is slightly lighter but not white. Earsare relatively small and triangular, with 4-5 folds. Tragusis straight, short, and sharp and barely reaches one-half of ear length;it is whitish, with blackish tip at top and bends forward. Feet are relatively large (equally as long as tibia), covered by long fur or bristles. Atfirst sight, Daubenton’s Myotis might be confused with the Common Whiskered Myotis ( M. mystacinus ) or Brandt's Myotis ( M. brandtii ), but it has shorter feet. Skull is relatively flat and broad with weak sagittal and somewhat strong lambdoidal crest. P° lies within tooth row; upper molars have developed protoconules; and lower molars are myotodont. Chromosomal complement has 2n = 44 and FNa = 52 ( Greece , Germany , and Spain ).	Mostly riparian habitats and water ecosystems (natural and artificial water), urban areas, forests, parks and traditional orchards from sea level up to elevations of ¢. 1400 m . Daubenton’s Myotis usually commute through woodlands (where they roost) to reach water where they feed. They can also be abundant in wetlands, rocky areas, inland cliffs, mountain peaks, and deciduous and mixed forests.	Daubenton’s Myotis is specialized to hunt just above calm water surfaces of streams, rivers, ponds, or lakes, being remarkably more abundant along open banks. They hunt ¢. 5-40 cm above the water surface, which makes trawling and capturing insects from the water surface easier. They move to hunting areas in groups of 2-6 individuals, but hunting is mostly done alone. They also hunt in meadows, open spaces, grasslands, and forest edges. It has been considered a specialist and generalist because it hunts Diptera opportunistically in some regions but also some Hymenoptera , Hemiptera , and Lepidoptera , very specifically in others. It has been shown experimentally that it can successfully hunt fish from the water.	Maternity colonies of Daubenton’s Myotis are formed in late spring (May or early June), generally in tree holes (hollow trees, rotten branches, abandoned woodpecker nests, etc.) and also bat boxes, bridges, and buildings. Maternity colony range from a few tens to a few hundred females and their young; some locations have up to 600 individuals. Females reach sexual maturity after the first year of their life. Small maternity colonies tend to change location after 3-5 days to avoid predation pressure, which means that a single nursery colony can use up to 40 tree holes/year. After parturition (with generally one young), females tend to forage in less than a 2-5-km radius around the roost. Young are commonly born from mid-June to July, depending on weather and region. In general, they need c.3—4 weeks to start flying. During the maternity period, solitary males or small groups of males will roost in the maternity colony or elsewhere in cellars, mines, or tunnels. Breeding season usually ends in August. During mating in September—November (particularly intense in October-November), they are most commonly found at swarming sites (e.g. caves, mines, or buildings) where large numbers of individuals converge to mate after traveling up to 30 km . Maximum reported longevity is c.4 years, butit is assumed it can live longer similar to some congeners.	In winter, most roosts are caves, mines, and other underground sites with temperatures of 3-8°C, but some individuals also roost in tree holes and deep cavities in hollow trees, caves, bunkers, and mines. Echolocation calls have short, highly modulated pulses, starting at more than 60 kHz and ending below 30 kHz (broad frequency range), with frequency of maximum energy of c.40 kHz. When they emerge at night, usually after sunset, they move along linearstructuresin the landscape such as forest corridors, ditches, edges,trails, etc. During the non-breeding period, individuals can fly up to 10 km /night in straight line to reach hunting areas.	Daubenton’s Myotis is considered non-migratory but will move ¢. 150-304 km from maternity roosts to hibernation sites. Individuals from lowland regions are assumed to move longer distances than those roosting in high mountains. Especially in the north-eastern part ofits distribution, they congregate in winter in large numbers (e.g. 20,000 in Nietoperek, Poland or up to 5000 individuals in Berlin , Germany ).	Classified as Least Concern on The IUCN Red List. Daubenton’s Myotis is one of the most abundant species in many parts of its distribution, especially in Central and Eastern Europe. Some monitoring programs suggest that its distribution and population are increasing, but fluctuations in numbers can be extreme depending on the area studied.	Akasaka et al. (2009) | Bogdanowicz (1994) | Boonman, A.M. & Jones (2002) | Boonman, A.M. et al. (1998) | Boonman, M. (2000) | Britton & Jones (1999) | Dietz & Kalko (2006) | Dietz et al. (2006) | Encarnacao & Dietz (2006) | Encarnacao, Becker & Ekschmitt (2010) | Encarnagéo, Dietz & Kierdorf (2004) | Encarnagao, Dietz, Kierdorf & Wolters (2004) | Encarnacao, Kierdorf et al. (2005) | Flavin et al. (2001) | Jones & Kokurewicz (1994) | Kokurewicz (2004) | Kriiger et al. (2012) | Lépez-Baucells, Casanova et al. (2017) | Luan & Radil (2010) | Parsons & Jones (2003) | Russo (2002) | Shirley et al. (2001) | Siemers, Dietz, Nill & Schnitzler (2001) | Siemers, Stilz & Schnitzler (2001) | Siivonen & Wermundsen (2008) | Simmons (2005) | Stubbe, Ariunbold, Buuveibaatar, Dorjderem, Monkhzul, Otgonbaatar, Tsogbadrakh, Hutson et al. (2008) | Vesterinen, Lilley et al. (2013) | Vesterinen, Ruokolainen et al. (2016) | Volleth & Heller (2012) | Warren et al. (2000)	https://zenodo.org/record/6398985/files/figure.png	482. Daubenton’s Myotis Myotis daubentonii French: Murin de Daubenton / German: \Wasserfledermaus / Spanish: Ratonero ribereno Other common names: Daubenton’s Bat Taxonomy. Vespertilio daubentonii Kuhl, 1817 , Hanau, Hessen , Germany . Subgenus Myotis ; daubentonii species group. Phylogenetic studies in 2013 and 2015 recovered M. daubentonii in a clade with M. sicarius , M. bechsteini , M. longicaudatus , M. soror, and M. frater . Monotypic. Distribution. Most of Europe and W Asia, from S Fennoscandia, British Is, and Iberian Peninsula E to W Russia (E to Omsk region ), the Caucasus, and N Turkey ; disrupted distribution in Mediterranean areas. Descriptive notes. Head-body 45-55 mm , tail 27-50 mm , ear 9-14 mm , hindfoot 9-11 mm , forearm 33-42 mm ; weight 6-10 g . Daubenton’s Myotis has grayish or brownish dorsal and ventral hairs, with dark bases and more brighttips, sometimes reddish. Wings are generally dark brown or gray. Young are generally darker than adults. Plagiopatagium starts at base of fingers, Uropatagium is naked; venter is slightly lighter but not white. Earsare relatively small and triangular, with 4-5 folds. Tragusis straight, short, and sharp and barely reaches one-half of ear length;it is whitish, with blackish tip at top and bends forward. Feet are relatively large (equally as long as tibia), covered by long fur or bristles. Atfirst sight, Daubenton’s Myotis might be confused with the Common Whiskered Myotis ( M. mystacinus ) or Brandt's Myotis ( M. brandtii ), but it has shorter feet. Skull is relatively flat and broad with weak sagittal and somewhat strong lambdoidal crest. P° lies within tooth row; upper molars have developed protoconules; and lower molars are myotodont. Chromosomal complement has 2n = 44 and FNa = 52 ( Greece , Germany , and Spain ). Habitat. Mostly riparian habitats and water ecosystems (natural and artificial water), urban areas, forests, parks and traditional orchards from sea level up to elevations of ¢. 1400 m . Daubenton’s Myotis usually commute through woodlands (where they roost) to reach water where they feed. They can also be abundant in wetlands, rocky areas, inland cliffs, mountain peaks, and deciduous and mixed forests. Food and Feeding. Daubenton’s Myotis is specialized to hunt just above calm water surfaces of streams, rivers, ponds, or lakes, being remarkably more abundant along open banks. They hunt ¢. 5-40 cm above the water surface, which makes trawling and capturing insects from the water surface easier. They move to hunting areas in groups of 2-6 individuals, but hunting is mostly done alone. They also hunt in meadows, open spaces, grasslands, and forest edges. It has been considered a specialist and generalist because it hunts Diptera opportunistically in some regions but also some Hymenoptera , Hemiptera , and Lepidoptera , very specifically in others. It has been shown experimentally that it can successfully hunt fish from the water. Breeding. Maternity colonies of Daubenton’s Myotis are formed in late spring (May or early June), generally in tree holes (hollow trees, rotten branches, abandoned woodpecker nests, etc.) and also bat boxes, bridges, and buildings. Maternity colony range from a few tens to a few hundred females and their young; some locations have up to 600 individuals. Females reach sexual maturity after the first year of their life. Small maternity colonies tend to change location after 3-5 days to avoid predation pressure, which means that a single nursery colony can use up to 40 tree holes/year. After parturition (with generally one young), females tend to forage in less than a 2-5-km radius around the roost. Young are commonly born from mid-June to July, depending on weather and region. In general, they need c.3—4 weeks to start flying. During the maternity period, solitary males or small groups of males will roost in the maternity colony or elsewhere in cellars, mines, or tunnels. Breeding season usually ends in August. During mating in September—November (particularly intense in October-November), they are most commonly found at swarming sites (e.g. caves, mines, or buildings) where large numbers of individuals converge to mate after traveling up to 30 km . Maximum reported longevity is c.4 years, butit is assumed it can live longer similar to some congeners. Activity patterns. In winter, most roosts are caves, mines, and other underground sites with temperatures of 3-8°C, but some individuals also roost in tree holes and deep cavities in hollow trees, caves, bunkers, and mines. Echolocation calls have short, highly modulated pulses, starting at more than 60 kHz and ending below 30 kHz (broad frequency range), with frequency of maximum energy of c.40 kHz. When they emerge at night, usually after sunset, they move along linearstructuresin the landscape such as forest corridors, ditches, edges,trails, etc. During the non-breeding period, individuals can fly up to 10 km /night in straight line to reach hunting areas. Movements, Home range and Social organization. Daubenton’s Myotis is considered non-migratory but will move ¢. 150-304 km from maternity roosts to hibernation sites. Individuals from lowland regions are assumed to move longer distances than those roosting in high mountains. Especially in the north-eastern part ofits distribution, they congregate in winter in large numbers (e.g. 20,000 in Nietoperek, Poland or up to 5000 individuals in Berlin , Germany ). Status and Conservation. Classified as Least Concern on The IUCN Red List. Daubenton’s Myotis is one of the most abundant species in many parts of its distribution, especially in Central and Eastern Europe. Some monitoring programs suggest that its distribution and population are increasing, but fluctuations in numbers can be extreme depending on the area studied. Bibliography. Akasaka et al. (2009), Bogdanowicz (1994), Boonman, A.M. & Jones (2002), Boonman, A.M. et al. (1998), Boonman, M. (2000), Britton & Jones (1999), Dietz & Kalko (2006), Dietz et al. (2006), Encarnacao & Dietz (2006), Encarnacao, Becker & Ekschmitt (2010), Encarnagéo, Dietz & Kierdorf (2004), Encarnagao, Dietz, Kierdorf & Wolters (2004), Encarnacao, Kierdorf et al. (2005), Flavin et al. (2001), Jones & Kokurewicz (1994), Kokurewicz (2004), Kriiger et al. (2012), Lépez-Baucells, Casanova et al. (2017), Luan & Radil (2010), Parsons & Jones (2003), Russo (2002), Shirley et al. (2001), Siemers, Dietz, Nill & Schnitzler (2001), Siemers, Stilz & Schnitzler (2001), Siivonen & Wermundsen (2008), Simmons (2005), Stubbe, Ariunbold, Buuveibaatar, Dorjderem, Monkhzul, Otgonbaatar, Tsogbadrakh, Hutson et al. (2008), Vesterinen, Lilley et al. (2013), Vesterinen, Ruokolainen et al. (2016), Volleth & Heller (2012), Warren et al. (2000).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis daubentonii	Myotis	Unassigned-Myotis	daubentonii	Kuhl	1817	1	Die Deutschen Flederm&auml;use. Hanau	p. 14	Daubenton's Myotis	 aedilus Jenyns, 1839; albus Fitzinger, 1871; capucinellus Fitzinger, 1871; lanatus Crespon, 1844; minutellus Fitzinger, 1871; staufferi Fatio, 1890; <b> chasanensis </b> Tiunov, 1997; <b>loukashkini</b> Shamel, 1942; <b>nathalinae</b> Tupinier, 1977; <b> ussuriensis </b> Ognev, 1927; <b> volgensis </b> Eversmann, 1840.	Germany, Hessen, Hanau.	Europe (including Britain and Ireland; Scandinavia) through Kazakhstan.	Not listed.	Least Concern	Does not include abei or petax ( petax is the senior synonym of abei ); see Matveev et al. (2005), Kruskop et al. (2012), and Katzmann et al. (2012). Does not include nathalinae ; see Horácek and Hanák (1984), Fairon (1985), Mayer and von Helversen (2001 a ), and Ruedi and Mayer (2001). Reviewed in part by Yoshiyuki (1989), Yoon (1990), Bates and Harrison (1997), Bates et al. (1999), andHorácek et al. (2000). Does not appear to include laniger ; see Topál (1997) and Bates et al. (1999), though also see Corbet and Hill(1992). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Bogdanowicz (1994), but note that laniger was includedin daubentonii in that publication. Apparently closely related to bechsteinii ; see Ruedi and Mayer (2001). Subspecies limitsare problematic, see Bogdanowicz (1994), Horácek et al. (2000), and Kruskop (2002). Genetic studies suggest that this complex includes morethan one species, with at least some Russian and Japanese specimens representing a taxon distinct from the European form (Kawai et al., 2003).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2027	Myotis daubentonii	23	Daubenton's Myotis	Daubenton's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	daubentonii	Kuhl	1817	1						Hanau, Hessen, Germany.			daubentonii (Kuhl, 1817)|aedilus (Jenyns, 1839)|volgensis (Eversmann, 1840)|lanatus (Crespon, 1844)|albus (Fitzinger, 1871)|capucinellus (Fitzinger, 1871)|minutellus (Fitzinger, 1871)|staufferi (Fatio, 1890)|nathalinae Tupinier, 1977	previously included M. petax (and abei)	Tsytsulina, K. (2004). On the taxonomical status of Myotis abei Yoshikura, 1944 (Chiroptera, Vespertilionidae). Zoological science, 21(9), 963-966.|Matveev, V. A., Kruskop, S. V., & Kramerov, D. A. (2005). Revalidation of Myotis petax Hollister, 1912 and its new status in connection with M. daubentonii (Kuhl, 1817)(Vespertilionidae, Chiroptera). Acta Chiropterologica, 7(1), 23-37.	Ireland|United Kingdom|Spain|Portugal|France|Belgium|Netherlands|Luxembourg|Germany|Denmark|Switzerland|Liechtenstein?|Austria|Italy|Czech Republic|Slovakia|Hungary|Slovenia|Croatia|Bosnia & Herzegovina|Albania?|Montenegro?|Serbia|Kosovo|North Macedonia|Greece|Bulgaria|Romania|Ukraine|Belarus|Poland|Lithuania|Latvia|Estonia|Norway|Sweden|Finland|Russia|Georgia|Turkey|Kazakhstan	Asia|Europe	Palearctic	LC	0	0	0	Myotis_daubentonii	0	sciname match	Myotis_daubentonii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Myotis daubentonii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	daubentonii	(Kuhl, 1817)	<span>This is the new taxonomic concept for Myotis daubentonii . Previously the assessment referred to M. daubentonii sensu lato, thus including M. petax from Altai and Siberia which is now considered a distinct species (Matveev et al. 2005).	200000000	Myotis daubentonii	Least Concern		2020	2019-05-14 00:00:00 UTC	3.1	English	Listed as Least Concern because it is widespread and abundant, there are no major threats and there are indications that its population is currently increasing.	Maternity colonies may comprise up to 600 females and roost in tree cavities, buildings or other artificial structures (e.g. bridges), as well as in bat boxes. The common colony size is about 40 females which frequently change tree roosts, while roosts in buildings are occupied for a longer time. Males may also form summer colonies of up to 200 individuals roosting in various underground and overground shelters (Simon et al. 2013). It winters in a wide range of underground habitats, sometimes forming large clusters on cave walls and roof. Seasonal movements between winter and summer roosts are mostly within a distance of 100-150 km (Hutterer et al. 2005). The longest distance covered is 304 km for males and 261 km for females, the maximum age 30 and 22 years respectively (Steffens et al. 2007). M. daubentonii is associated with aquatic habitats, where it preys either on the wing or trawls the water surface with its feet and/or its wing membrane. Feeding areas are usually at a maximum distance of 2–5 km from the roosts, but may occasionally be as far as 10 km away from the roost. Females tend to forage closer to their roost than males. Altitudinal habitat segregation between sexes was described for this species across the European and Caucasian range: males tend to occupy upstream areas and females exploit more productive habitats at lower elevations Feeding areas of pregnant and lactating females are typically small. After the young are weaned females also use larger areas. Females show a high fidelity to quality feeding areas even though they might change the roost quite often. It preys on Chironomidae/ Ceratopogonidae, Diptera and Trichoptera. Terrestrial insects like Brachycera or Coleoptera were also detected in Germany and Finland (see the literature review in Kyheröinen et al. 2019).	Although it is a major vector of European Bat Lyssavirus 2 (Racey et al. 2012), there appears to be no major threats to this species overall. Changes in water quality may reduce food supply, and loss of or damage and disturbance to roost sites in trees, buildings, other artificial structures, and underground habitats may cause temporary localized losses. However, these are not thought to be serious threats to the survival of this abundant and expanding species.	M. daubentonii ; is widespread and the population appears to be stable.	Stable	The species is rather rare in Mediterranean, but was recorded in all Balkan countries, including Montenegro (Karapandža et al. 2014, Presetnik et al. 2014). It wasn't found in a steppe belt of Ukraine and Russia, any records of Daubenton's bat are missing for Crimea despite special efforts for its detecting. M. daubentonii is common in North Caucasus and woodlands of Georgia including those at the border with Turkey, whereas validated records from Armenia and Azerbaijan are still lacking (Gazaryan et al. 2008). The range in Turkey may surround the southern Black Sea coast, although the species hasn't yet been revealed between Samsun province and Georgia (Çoraman, pers. comm. 2018). The taxon formerly included the Asian species Myotis petax which is distinct both genetically and in terms of morphology. Distributional limits for both species or areas of their possible sympatry are still to be ascertained, but are likely to occur in the Omsk region (Matveev et al. 2005).		Terrestrial	It is protected by national legislation in most of the main ; range states, except Russia. There are also international legal obligations for its protection through the Convention on Migratory Species (CMS) and EUROBATS Agreement. It is included in Appendix II ; of the Council of Europe’s Convention on the Conservation of European Wildlife and Natural Habitats (Bern Convention). Also included in Annex IV of EU Habitats and Species Directive. Important habitats partially protected by Natura 2000 sites and national networks of protected areas. ;Real population trends outside the EU are barely known due to the lack of research.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	daubentonii	Kuhl	1817	1	Die Deutschen Flederm&auml;use. Hanau	p. 14	Daubenton's Myotis	 aedilus Jenyns, 1839; albus Fitzinger, 1871; capucinellus Fitzinger, 1871; lanatus Crespon, 1844; minutellus Fitzinger, 1871; staufferi Fatio, 1890; <b> chasanensis </b> Tiunov, 1997; <b>loukashkini</b> Shamel, 1942; <b>nathalinae</b> Tupinier, 1977; <b> ussuriensis </b> Ognev, 1927; <b> volgensis </b> Eversmann, 1840.	Germany, Hessen, Hanau.	Europe (including Britain and Ireland; Scandinavia) through Kazakhstan.	Not listed.	Least Concern	Does not include abei or petax ( petax is the senior synonym of abei ); see Matveev et al. (2005), Kruskop et al. (2012), and Katzmann et al. (2012). Does not include nathalinae ; see Horácek and Hanák (1984), Fairon (1985), Mayer and von Helversen (2001 a ), and Ruedi and Mayer (2001). Reviewed in part by Yoshiyuki (1989), Yoon (1990), Bates and Harrison (1997), Bates et al. (1999), andHorácek et al. (2000). Does not appear to include laniger ; see Topál (1997) and Bates et al. (1999), though also see Corbet and Hill(1992). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Bogdanowicz (1994), but note that laniger was includedin daubentonii in that publication. Apparently closely related to bechsteinii ; see Ruedi and Mayer (2001). Subspecies limitsare problematic, see Bogdanowicz (1994), Horácek et al. (2000), and Kruskop (2002). Genetic studies suggest that this complex includes morethan one species, with at least some Russian and Japanese specimens representing a taxon distinct from the European form (Kawai et al., 2003).	Myotis daubentonii	1005395	23	Daubenton's Myotis	Daubenton's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	daubentonii	Kuhl	1817	1						Hanau, Hessen, Germany.			daubentonii (Kuhl, 1817)|aedilus (Jenyns, 1839)|volgensis (Eversmann, 1840)|lanatus (Crespon, 1844)|albus (Fitzinger, 1871)|capucinellus (Fitzinger, 1871)|minutellus (Fitzinger, 1871)|staufferi (Fatio, 1890)|nathalinae Tupinier, 1977	previously included M. petax (and abei)	Tsytsulina, K. (2004). On the taxonomical status of Myotis abei Yoshikura, 1944 (Chiroptera, Vespertilionidae). Zoological science, 21(9), 963-966.|Matveev, V. A., Kruskop, S. V., & Kramerov, D. A. (2005). Revalidation of Myotis petax Hollister, 1912 and its new status in connection with M. daubentonii (Kuhl, 1817)(Vespertilionidae, Chiroptera). Acta Chiropterologica, 7(1), 23-37.				Ireland|United Kingdom|Spain|Portugal|France|Belgium|Netherlands|Luxembourg|Germany|Denmark|Switzerland|Liechtenstein?|Austria|Italy|Czech Republic|Slovakia|Hungary|Slovenia|Croatia|Bosnia & Herzegovina|Albania?|Montenegro?|Serbia|Kosovo|North Macedonia|Greece|Bulgaria|Romania|Ukraine|Belarus|Poland|Lithuania|Latvia|Estonia|Norway|Sweden|Finland|Russia|Georgia|Turkey|Kazakhstan	Asia|Europe	Palearctic	LC	0	0	0	Myotis_daubentonii	0	sciname match	Myotis_daubentonii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_daubentonii	1005395	23	Daubenton's Myotis	Daubenton's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	daubentonii	Kuhl	1	Vespertilio daubentonii	Kuhl, H. 1817. Die deutschen Fledermäuse. None, Hanau, 67 pp.	https://nbn-resolving.org/urn:nbn:de:hebis:30-91692				Hanau, Hessen, Germany.			previously included M. petax (and abei)	Tsytsulina, K. (2004). On the taxonomical status of Myotis abei Yoshikura, 1944 (Chiroptera, Vespertilionidae). Zoological science, 21(9), 963-966.|Matveev, V. A., Kruskop, S. V., & Kramerov, D. A. (2005). Revalidation of Myotis petax Hollister, 1912 and its new status in connection with M. daubentonii (Kuhl, 1817)(Vespertilionidae, Chiroptera). Acta Chiropterologica, 7(1), 23-37.				Ireland|United Kingdom|Spain|Portugal|France|Belgium|Netherlands|Luxembourg|Germany|Denmark|Switzerland|Liechtenstein?|Austria|Italy|Czech Republic|Slovakia|Hungary|Slovenia|Croatia|Bosnia and Herzegovina|Albania?|Montenegro?|Serbia|Kosovo|North Macedonia|Greece|Bulgaria|Romania|Ukraine|Belarus|Poland|Lithuania|Latvia|Estonia|Norway|Sweden|Finland|Russia|Georgia|Turkey|Kazakhstan	Asia|Europe	Palearctic	LC	0	0	0	Myotis_daubentonii	0	sciname match	Myotis_daubentonii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	daubentonii	Kuhl	1817	1	Die Deutschen Flederm&auml;use. Hanau	p. 6, 51	Daubenton's Myotis	aedilus Jenyns, 1839; albus Fitzinger, 1871; capucinellus Fitzinger, 1871; lanatus Crespon, 1844; minutellus Fitzinger, 1871; staufferi Fatio, 1890; chasanensis Tiunov, 1997; loukashkini Shamel, 1942; nathalinae Tupinier, 1977; ussuriensis Ognev, 1927; volgensis Eversmann, 1840.	Germany, Hessen, Hanau.	Europe (including Britain and Ireland; Scandinavia) through Kazakhstan.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/85342710/195858793/' target='_blank'>Least Concern</a>	Does not include abei or petax (petax is the senior synonym of abei); see Matveev et al. (2005), Kruskop et al. (2012), and Katzmann et al. (2012). Does not include nathalinae; see Horácek and Hanák (1984), Fairon (1985), Mayer and von Helversen (2001 a), and Ruedi and Mayer (2001). Reviewed in part by Yoshiyuki (1989), Yoon (1990), Bates and Harrison (1997), Bates et al. (1999), andHorácek et al. (2000). Does not appear to include laniger; see Topál (1997) and Bates et al. (1999), though also see Corbet and Hill(1992). For discussion of correct spelling see Bogdanowicz and Kock (1998). See Bogdanowicz (1994), but note that laniger was includedin daubentonii in that publication. Apparently closely related to bechsteinii; see Ruedi and Mayer (2001). Subspecies limitsare problematic, see Bogdanowicz (1994), Horácek et al. (2000), and Kruskop (2002). Genetic studies suggest that this complex includes morethan one species, with at least some Russian and Japanese specimens representing a taxon distinct from the European form (Kawai et al., 2003).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis daubentonii; Myotis daubentonii; Myotis daubentonii; Myotis daubentonii; Myotis daubentonii; Myotis daubentonii; daubentonii; chasanensis; loukashkini; nathalinae; petax; ussuriensis; volgensis; aedilus; albus; capucinellus; lanatus; minutellus; staufferi; chasanensis; loukashkini; nathalinae; ussuriensis; volgensis; aedilus; albus; capucinellus; lanatus; minutellus; staufferi; daubentonii; aedilus; volgensis; lanatus; albus; capucinellus; minutellus; staufferi; nathalinae; Murin de Daubenton; \Wasserfledermaus; Ratonero ribereno; Daubenton’s Bat; Daubenton's Myotis; Daubenton's Bat; Daubenton's Myotis; Daubenton's Myotis; M. daubentonii