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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L847	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	Myotis ciliolabrum	Myotis leibii [synonym of]	Myotis leibii ciliolabrum	Myotis ciliolabrum	Myotis ciliolabrum	Myotis ciliolabrum	Myotis ciliolabrum	Myotis ciliolabrum	Myotis ciliolabrum	Myotis ciliolabrum	Myotis ciliolabrum	Myotis ciliolabrum	Myotis ciliolabrum		[MSW3] Formerly included in leibii (for which Hall [1981] used the name subulatus), but see van Zyll de Jong (1984). Does not include melanorhinus; see van Zyll de Jong (1984). Reviewed by Holloway and Barclay (2001), but note that they included melanorhinus as a subspecies of ciliolabrum.; [HMW] Vespertilio ciliolabrum Merriam, 1886 , “near the town of Banner,” Trego County, Kansas , USA . Subgenus Pyzonux; lucifugus species group. Populations of M. ciliolabrum were previously called M. keenii , M. leibir, M. subulatus , and M. melanorhinus . Monotypic.; [batnames2022] Formerly included in leibii (for which Hall [1981] used the name subulatus ), but see van Zyll de Jong (1984). Includes melanorhinus ; see Ammerman et al. (2016).; [MDD2022] includes melanorhinus; [IUCN] Formerly included in leibii . Reviewed by Holloway and Barclay (2001), but note that they included melanorhinus as a subspecies of ciliolabrum .; [batnames2023] Formerly included in leibii (for which Hall [1981] used the name subulatus ), but see van Zyll de Jong (1984). Includes melanorhinus ; see Ammerman et al. (2016).; [MDD2023] includes melanorhinus; [MDD2025_2.0] includes melanorhinus; [batnames2025_1.7] Formerly included in leibii (for which Hall [1981] used the name subulatus), but see van Zyll de Jong (1984). Includes melanorhinus; see Ammerman et al. (2016).; [MDD2025_2.2] includes melanorhinus													melanorhinus	subulatus, ciliolabrum, yumanensis, ciliolabrum, melanorhinus, henshawii, orinomus	Formerly included in leibii . Reviewed by Holloway and Barclay (2001), but note that they included melanorhinus as a subspecies of ciliolabrum .	ciliolabrum 	ciliolabrum - melanorhinus	subulatus, ciliolabrum, melanorhinus, henshawii, orinomus 	ciliolabrum, melanorhinus, henshawii, orinomus	ciliolabrum	ciliolabrum - melanorhinus 	ciliolabrum (C. H. Merriam, 1886)|melanorhinus (C. H. Merriam, 1890)|henshawii (H. Allen, 1894)|orinomus D. G. Elliot, 1903						N/A							Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		SW Canada – Nebraska, Oklahoma, W Mexico; ref. 4.107																															_M. c. ciliolabrum_ (Merriam, 1886) (synonyms: _henshawii_ (Allen, 1894)); _M. c. melanorhinus_ (Merriam, 1890) (synonyms: _orinomus_ Elliot, 1903)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis ciliolabrum	Myotis		ciliolabrum	Merriam		1886		Proc. Biol. Soc. Wash.	4		2		Western Small-footed Myotis	United States, Kansas, Trego Co., near Banner, about 1 mi. (1.5 km) from Castle Rock, bluff on Hackberry Creek.	S Alberta and Saskatchewan (Canada) south through E Colorado and W Kansas (USA).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Formerly included in leibii (for which Hall [1981] used the name subulatus), but see van Zyll de Jong (1984). Does not include melanorhinus; see van Zyll de Jong (1984). Reviewed by Holloway and Barclay (2001), but note that they included melanorhinus as a subspecies of ciliolabrum.	4C3D87E8FF586AE0FA869DA11A47B283	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	925	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF586AE0FA869DA11A47B283.xml	Myotis ciliolabrum	Vespertilionidae	Myotis	ciliolabrum		1886	Murin pygmée @fr | Westliches Kleinful3-Mausohr @de | Ratonero pigmeo @es	Vespertilio ciliolabrum Merriam, 1886 , “near the town of Banner,” Trego County, Kansas , USA . Subgenus Pyzonux; lucifugus species group. Populations of M. ciliolabrum were previously called M. keenii , M. leibir, M. subulatus , and M. melanorhinus . Monotypic.	From SE Alberta and SW Saskatchewan ( Canada ) S to SC USA , including E Idaho , Montana , SW North Dakota , Wyoming , W & S South Dakota , NE Utah , E Colorado , Nebraska , W Kansas , and extreme NE New Mexico .	Head-body ¢.39 41 mm , tail 37-49 mm , ear 12-16 mm , hindfoot 6-9 mm , forearm 31:2-35- 7 mm ; weight 3-6 g . Females are slightly larger than males. The Western Small-footed Myotis is morphologically similar to the Eastern Smallfooted Myotis (M. leibir) and the Dark-nosed Small-footed Myotis ( M. melanorhinus ). Fur is long and silky. Pelage is generally paler than in other North American Myotis . Dorsal hairs are bicolored, with blackish bases and flaxen to pale yellow tips; ventral hairs are is slightly paler, with dark brown bases and light yellow or nearly white tips. Ears are moderately long and extend slightly beyond nose when laid forward; tragus is long and narrow, being ¢.50% of ear length. Membranes, ears, and skin are blackish. Plagiopatagium is attached to feet by a broad band of membrane; calcar is distinctly keeled. Skull is moderate in size (greatest skull lengths 13-5-14- 7 mm ), and compared with the Eastern SmallHfooted Myotis , the Dark-nosed Small-footed Myotis , and the California Myotis ( M. californicus ), the Western Small-footed Myotis hasless steeply sloping frontal bone, flatter and broader skull, and greater rostral breadth. Parietal is normally inclined forward; occipital is rounded posteriorly; braincase is flattened; sagittal crest is generally absent but, when present,is very low; and lambdoidal crests are generally present and low. Chromosomal complement has 2n = 44 and FN = 50, with four pairs of metacentric and 17 pairs of acrocentric autosomes. X- and Y-chromosomes are submetacentric.	Grasslands with rock outcrops, coniferous forests, deserts, and dry shrublands, at elevations of 300-3300 m . Along the Rockies and adjacent plains, Western Smallfooted Myotis are generally found in broken terrain of canyons and foothills, commonly in places with cover of trees or shrubs.	The Western Small-footed Myotis is an aerial insectivore that forages only c. 1 m aboveground;it flights is erratic and slow. One-half of its foraging time is spent near rocks and cliffs and the remainder over open areas, forest canopy, and water. Diets contain Lepidoptera but also Diptera , Hemiptera , Hymenoptera , and Coleoptera .	The Western Small-footed Myotisis polygynandrous and reproduces seasonally, generally starting in early July. Mating season begins shortly before hibernation, and females store sperm until spring when ovulation and fertilization occur. Gestation lasts ¢.2 months. Most females give birth to one young, but twins have been reported. Parturition seems to occur in July-September. Sexual maturity typically occurs at just over a year of age.	The Western Small-footed Myotis begins foraging before full dark, and activity is concentrated in the three first hours of the night. It hibernates from early autumn through winter. It roosts in rock crevices,cliffs, abandoned mines, caves, and occasionally tunnels and buildings and under tree bark and rocks scattered on the ground. In Wyoming , calls had mean maximum frequency of 62-4 kHz, minimum frequency of 40-3 kHz, and duration of 3-5 milliseconds.	While foraging, no Western Smallfooted Myotis were reported further than 1 km from their roost or foraging areas. Western Small-footed Myotis tend to be more solitary than congeners. In summer, pregnant females form maternity colonies of up to 20 individuals (adults and young), but sometimes they are solitary.	Classified as Least Concern on The IUCN Red List. The Western SmallHfooted Myotis is widespread, common over most of its distribution, occurs in protected areas, and is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	Armstrong et al. (2011) | Arroyo-Cabrales & Alvarez-Castafeda (2017h) | Halloway & Barclay (2001) | Landry (2004) | O'Farrell et al. (1999) | Simmons (2005) | Wilson & Ruff (1999)	https://zenodo.org/record/6398648/files/figure.png	372. Western Small-footed Myotis Myotis ciliolabrum French: Murin pygmée / German: Westliches Kleinful3-Mausohr / Spanish: Ratonero pigmeo Taxonomy. Vespertilio ciliolabrum Merriam, 1886 , “near the town of Banner,” Trego County, Kansas , USA . Subgenus Pyzonux; lucifugus species group. Populations of M. ciliolabrum were previously called M. keenii , M. leibir, M. subulatus , and M. melanorhinus . Monotypic. Distribution. From SE Alberta and SW Saskatchewan ( Canada ) S to SC USA , including E Idaho , Montana , SW North Dakota , Wyoming , W & S South Dakota , NE Utah , E Colorado , Nebraska , W Kansas , and extreme NE New Mexico . Descriptive notes. Head-body ¢.39 41 mm , tail 37-49 mm , ear 12-16 mm , hindfoot 6-9 mm , forearm 31:2-35- 7 mm ; weight 3-6 g . Females are slightly larger than males. The Western Small-footed Myotis is morphologically similar to the Eastern Smallfooted Myotis (M. leibir) and the Dark-nosed Small-footed Myotis ( M. melanorhinus ). Fur is long and silky. Pelage is generally paler than in other North American Myotis . Dorsal hairs are bicolored, with blackish bases and flaxen to pale yellow tips; ventral hairs are is slightly paler, with dark brown bases and light yellow or nearly white tips. Ears are moderately long and extend slightly beyond nose when laid forward; tragus is long and narrow, being ¢.50% of ear length. Membranes, ears, and skin are blackish. Plagiopatagium is attached to feet by a broad band of membrane; calcar is distinctly keeled. Skull is moderate in size (greatest skull lengths 13-5-14- 7 mm ), and compared with the Eastern SmallHfooted Myotis , the Dark-nosed Small-footed Myotis , and the California Myotis ( M. californicus ), the Western Small-footed Myotis hasless steeply sloping frontal bone, flatter and broader skull, and greater rostral breadth. Parietal is normally inclined forward; occipital is rounded posteriorly; braincase is flattened; sagittal crest is generally absent but, when present,is very low; and lambdoidal crests are generally present and low. Chromosomal complement has 2n = 44 and FN = 50, with four pairs of metacentric and 17 pairs of acrocentric autosomes. X- and Y-chromosomes are submetacentric. Habitat. Grasslands with rock outcrops, coniferous forests, deserts, and dry shrublands, at elevations of 300-3300 m . Along the Rockies and adjacent plains, Western Smallfooted Myotis are generally found in broken terrain of canyons and foothills, commonly in places with cover of trees or shrubs. Food and Feeding. The Western Small-footed Myotis is an aerial insectivore that forages only c. 1 m aboveground;it flights is erratic and slow. One-half of its foraging time is spent near rocks and cliffs and the remainder over open areas, forest canopy, and water. Diets contain Lepidoptera but also Diptera , Hemiptera , Hymenoptera , and Coleoptera . Breeding. The Western Small-footed Myotisis polygynandrous and reproduces seasonally, generally starting in early July. Mating season begins shortly before hibernation, and females store sperm until spring when ovulation and fertilization occur. Gestation lasts ¢.2 months. Most females give birth to one young, but twins have been reported. Parturition seems to occur in July-September. Sexual maturity typically occurs at just over a year of age. Activity patterns. The Western Small-footed Myotis begins foraging before full dark, and activity is concentrated in the three first hours of the night. It hibernates from early autumn through winter. It roosts in rock crevices,cliffs, abandoned mines, caves, and occasionally tunnels and buildings and under tree bark and rocks scattered on the ground. In Wyoming , calls had mean maximum frequency of 62-4 kHz, minimum frequency of 40-3 kHz, and duration of 3-5 milliseconds. Movements, Home range and Social organization. While foraging, no Western Smallfooted Myotis were reported further than 1 km from their roost or foraging areas. Western Small-footed Myotis tend to be more solitary than congeners. In summer, pregnant females form maternity colonies of up to 20 individuals (adults and young), but sometimes they are solitary. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Western SmallHfooted Myotis is widespread, common over most of its distribution, occurs in protected areas, and is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category. Bibliography. Armstrong et al. (2011), Arroyo-Cabrales & Alvarez-Castafeda (2017h), Halloway & Barclay (2001), Landry (2004), O'Farrell et al. (1999), Simmons (2005), Wilson & Ruff (1999).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis ciliolabrum	Myotis	Unassigned-Myotis	ciliolabrum	Merriam	1886	0	Proc. Biol. Soc. Wash.	4:02	Western Small-footed Myotis	Present	United States, Kansas, Trego Co., near Banner, about 1 mi. (1.5 km) from Castle Rock, bluff on Hackberry Creek	British Columbia, S Alberta. and Saskatchewan (Canada) south through E Colorado and Oklahoma (USA) to C Mexico	Not listed.	Least Concern	Formerly included in leibii (for which Hall [1981] used the name subulatus ), but see van Zyll de Jong (1984). Includes melanorhinus ; see Ammerman et al. (2016).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis ciliolabrum	23	Western Small-footed Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	ciliolabrum	Merriam	1886	1	Vespertilio_ciliolabrum	Merriam, C. H. (1886). Description of a New Species of Bat from the Western United States. Proceedings of the Biological Society of Washington, 4, 2.	https://www.biodiversitylibrary.org/page/3688929#page/31/mode/1up	USNM 186444		"near the town of Banner," Trego County, Kansas, USA.			subulatus (Say, 1823) [name either represents Myotis ciliolabrum or Myotis yumanensis and has been suggested for suppression to avoid further confusion; tentatively retained under Myotis ciliolabrum here]|ciliolabrum (Merriam, 1886)|melanorhinus Merriam, 1890|henshawii (H. Allen, 1894)|orinomus D. G. Elliot, 1903	includes melanorhinus	Ammerman, L. K., Lee, D. N., & Pfau, R. S. (2016). Patterns of genetic divergence among Myotis californicus, M. ciliolabrum, and M. leibii based on amplified fragment length polymorphism. Acta Chiropterologica, 18(2), 336-346.	Canada|United States|Mexico	North America	Palearctic	LC	0	0	0	Myotis_ciliolabrum	0	sciname match	Myotis_ciliolabrum	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14153	Myotis ciliolabrum	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	ciliolabrum	(Merriam, 1886)	Formerly included in leibii . Reviewed by Holloway and Barclay (2001), but note that they included melanorhinus as a subspecies of ciliolabrum .	20000000	Myotis ciliolabrum	Least Concern		2017	2016-08-29 00:00:00 UTC	3.1	English	This species is listed as Least Concern in because of its wide distribution, being common over most of its distribution, occurrence in a number of protected areas, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	The Western Small-footed Myotis has a wide ecological range, from rock outcrops on open grasslands to canyons in the foothills to lower mountains with yellow pine woodlands. Day roosts are variable, but include cracks and crevices in cliffs, beneath tree bark, in mines and caves, and occasionally in tunnels and dwellings of humans. Night roosts are under a variety of natural and human-induced structures. Reproductive M. ciliolabrum roost during the day solitarily or in small groups. Hibernacula include caves, mines, and tunnels. Myotis ciliolabrum feeds early in the evening on small flying insects such as flies (Diptera), bugs (Hemiptera), moths (Lepidoptera), small beetles (Coleoptera) and winged ants (Hymenoptera), but are known to consume Trichoptera as well (Holloway and Barclay 2001). ;They begin to forage well before full dark, but not as early as the pipistrelle (Pipistrellus hesperus ). ;This species is highly maneuverable in flight, often foraging among boulders, along cliffs or shrubs and trees. ;Presently, not much is known about reproductive behavior of this species. Parturition dates vary across its geographical range. Pregnant females have been encountered all through June; a single young is probably common. Scrotal (reproductive) males were captured in August and September.	There are no major threats throughout the species' range.	Myotis ciliolabrum is common in arid desert, badland, and semiarid habitats, although higher elevation, more mesic habitats are used in the southern part of the range (Holloway and Barclay 2001). In New Mexico, the species occurs at low to moderate elevations to as high as 9,500 feet elevation.	Stable	This species is distributed from South Alberta and Saskatchewan (Canada) south through Eastern Colorado and Western Kansas (USA) (Simmons 2005) to North and Central Mexico (Ceballos and Oliva 2005).		Terrestrial	It occurs in several protected areas through its geographic range. ;Myotis ciliolabrum is on the Blue List in Alberta and British Columbia, indicating it is at risk or vulnerable (Holloway and Barclay 2001). In the United States, the species was a Federal Category 2 candidate species (Fish and Wildlife Service 1994), meaning that it is a species of concern, but with insufficient information on population numbers to add them to the endangered species list.	Nearctic|Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	ciliolabrum	Merriam	1886	0	Proc. Biol. Soc. Wash.	4:02	Western Small-footed Myotis	Present	United States, Kansas, Trego Co., near Banner, about 1 mi. (1.5 km) from Castle Rock, bluff on Hackberry Creek	British Columbia, S Alberta. and Saskatchewan (Canada) south through E Colorado and Oklahoma (USA) to C Mexico	Not listed.	Least Concern	Formerly included in leibii (for which Hall [1981] used the name subulatus ), but see van Zyll de Jong (1984). Includes melanorhinus ; see Ammerman et al. (2016).	Myotis ciliolabrum	1005389	23	Western Small-footed Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	ciliolabrum	Merriam	1886	1	Vespertilio_ciliolabrum	Merriam, C. H. (1886). Description of a New Species of Bat from the Western United States. Proceedings of the Biological Society of Washington, 4, 2.	https://www.biodiversitylibrary.org/page/3688929#page/31/mode/1up	USNM 186444		"near the town of Banner," Trego County, Kansas, USA.			subulatus (Say, 1823) [name either represents Myotis ciliolabrum or Myotis yumanensis and has been suggested for suppression to avoid further confusion; tentatively retained under Myotis ciliolabrum here]|ciliolabrum (Merriam, 1886)|melanorhinus Merriam, 1890|henshawii (H. Allen, 1894)|orinomus D. G. Elliot, 1903	includes melanorhinus	Ammerman, L. K., Lee, D. N., & Pfau, R. S. (2016). Patterns of genetic divergence among Myotis californicus, M. ciliolabrum, and M. leibii based on amplified fragment length polymorphism. Acta Chiropterologica, 18(2), 336-346.			USA(KS,NM,OK,CO,NE,UT,WY,SD,ND,MT,ID,WA,OR,CA,NV,AZ,TX)	Canada|United States|Mexico	North America	Palearctic	LC	0	0	0	Myotis_ciliolabrum	0	sciname match	Myotis_ciliolabrum	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_ciliolabrum	1005389	23	Western Small-footed Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	ciliolabrum	C. H. Merriam	1	Vespertilio ciliolabrum	Merriam, C.H. 1886-12-17. Description of a new species of bat from the western United States (_Vespertilio ciliolabrum_ sp. nov.). Proceedings of the Biological Society of Washington 4:1-4.	https://www.biodiversitylibrary.org/page/3842030	USNM:MAMM:186444	holotype	http://n2t.net/ark:/65665/3290d0c0c-86f1-4794-b0da-b2011a82a1c6	"near the town of Banner," Trego County, Kansas, USA.			includes melanorhinus	Ammerman, L. K., Lee, D. N., & Pfau, R. S. (2016). Patterns of genetic divergence among Myotis californicus, M. ciliolabrum, and M. leibii based on amplified fragment length polymorphism. Acta Chiropterologica, 18(2), 336-346.			USA(KS,NM,OK,CO,NE,UT,WY,SD,ND,MT,ID,WA,OR,CA,NV,AZ,TX)	Canada|United States|Mexico	North America	Nearctic	LC	0	0	0	Myotis_ciliolabrum	0	sciname match	Myotis_ciliolabrum	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	ciliolabrum	Merriam	1886	1	Proc. Biol. Soc. Wash.	4:02	Western Small-footed Myotis	Present	United States, Kansas, Trego Co., near Banner, about 1 mi. (1.5 km) from Castle Rock, bluff on Hackberry Creek	British Columbia, S Alberta. and Saskatchewan (Canada) south through E Colorado and Oklahoma (USA) to C Mexico	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14153/22058110/' target='_blank'>Least Concern</a>	Formerly included in leibii (for which Hall [1981] used the name subulatus), but see van Zyll de Jong (1984). Includes melanorhinus; see Ammerman et al. (2016).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis ciliolabrum; Myotis ciliolabrum; Myotis ciliolabrum; Myotis ciliolabrum; Myotis ciliolabrum; Myotis ciliolabrum; subulatus; Myotis ciliolabrum; Myotis yumanensis; ciliolabrum; melanorhinus; henshawii; orinomus; Murin pygmée; Westliches Kleinful3-Mausohr; Ratonero pigmeo; Western Small-footed Myotis; Western Small-footed Myotis; Western Small-footed Myotis; M. ciliolabrum