http://www.w3.org/ns/prov#wasDerivedFrom	http://purl.org/dc/elements/1.1/format	name_CH1_1980	name_MSW1_1982	name_CH3_1991	name_MSW2_1993	name_Koopman_1994	name_MSW3_2005	name_HMW_2019	name_BatNames_2022	name_MDD_2022	name_IUCN_2022	name_BatNames_2023	name_MDD_2023	name_MDD_2025_2.0	name_batnames_2025_1.7	name_MDD_2025_2.2	column151	taxonomic_notes_concatenated	column171	synonyms_CH1	subspecies__MSW2	synonyms__MSW1	synonyms_CH3	synonyms_MSW2	subspecies_Koopman94_interpreted	subspecies_MSW3_interpreted	synonym_MSW3_interpreted	subspecies_HMW_interpreted	synonym_HMW_interpreted	subspecies_batnames_interpreted	synonym_batnames_interpreted	synonym_MDD_interpreted	synonym_IUCN_interpreted	subspecies_batnames2023_interpreted	synonym_batnames2023_interpreted	synonym_MDD2023_interpreted	synonym_MDD2025_interpreted	subspecies_batnames2025_interpreted	synonyms_batnames2025_interpreted	nominalNames	column391	docOrigin_CH1	commonName_CH1	distribution_CH1	docOrigin_MSW1	column451	typeLocality_MSW1	authority_MSW1	year_MSW1	citation_MSW1	distribution	comment_MSW1	docOrigin_CH3	commonName_CH3	distribution_CH3	docOrigin_MSW2	authority_MSW2	year_MSW2	citation_MSW2	comments_MSW2	distribution_MSW2	typeLocality_MSW2	docOrigin_Koopman94	authority_Koopman94	year_Koopman94	description_Koopman94	distribution_Koopman94	diversity_Koopman94	subspecies_Koopman94	page	rank	name	authority	year	parent	parent_rank	corrected_name	actual_species_count	claimed_species_count	dental_formula	description	diversity	full_subspecies_text	name_line	species_index	subspecies	synonym	text	docOrigin_MSW3	order_MSW3	family_MSW3	subfamily_MSW3	tribe_MSW3	name_MSW3	genus_MSW3	subgenus_MSW3	species_MSW3	authoritySpeciesAuthor_MSW3	(parentheses (1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common Name_batnames	synonyms_batnames	type_locality_batnames	Distribution_batnames	CITES_batnames	IUCN_batnames	comments_batnames	docOrigin_MDD	name_MDD	phylosort_MDD	mainCommonName_MDD	otherCommonNames_MDD	subclass_MDD	infraclass_MDD	magnorder_MDD	superorder_MDD	order_MDD	suborder_MDD	infraorder_MDD	parvorder_MDD	superfamily_MDD	family_MDD	subfamily_MDD	tribe_MDD	genus_MDD	subgenus_MDD	specificEpithet_MDD	authoritySpeciesAuthor_MDD	authoritySpeciesYear_MDD	authorityParentheses_MDD	originalNameCombination_MDD	authoritySpeciesCitation_MDD	authoritySpeciesLink_MDD	holotypeVoucher_MDD	holotypeVoucherURIs_MDD	typeLocality_MDD	typeLocalityLatitude_MDD	typeLocalityLongitude_MDD	nominalNames_MDD	taxonomyNotes_MDD	taxonomyNotesCitation_MDD	countryDistribution_MDD	continentDistribution_MDD	biogeographicRealm_MDD	iucnStatus_MDD	extinct_MDD	domestic_MDD	flagged_MDD	CMW_sciName_MDD	diffSinceCMW_MDD	MSW3_matchtype_MDD	MSW3_sciName_MDD	diffSinceMSW3_MDD	docOrigin_IUCN	internalTaxonId_IUCN	NAME_IUCN	kingdomName_IUCN	phylumName_IUCN	className_IUCN	orderName_IUCN	familyName_IUCN	genusName_IUCN	speciesName_IUCN	authoritySpeciesAuthorYear_IUCN	taxonomicNotes_IUCN	assessmentId_IUCN	scientificName_IUCN	redlistCategory_IUCN	redlistCriteria_IUCN	yearPublished_IUCN	assessmentDate_IUCN	criteriaVersion_IUCN	language_IUCN	rationale_IUCN	habitat_IUCN	threats_IUCN	population_IUCN	populationTrend_IUCN	range_IUCN	useTrade_IUCN	systems_IUCN	conservationActions_IUCN	realm_IUCN	yearLastSeen_IUCN	possiblyExtinct_IUCN	possiblyExtinctInTheWild_IUCN	scopes_IUCN	docOrigin_batnames2023	FAMILY_batnames2023	GENUS_batnames2023	SUBGENUS_batnames2023	SPECIES_batnames2023	authoritySpeciesAuthor_batnames2023	authoritySpeciesYearbatnames2023	PARENTHESES_batnames2023 (1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name MDD2023	id_MDD2023	phylosort_MDD2023	mainCommonName_MDD2023	otherCommonNames_MDD2023	subclass_MDD2023	infraclass_MDD2023	magnorder_MDD2023	superorder_MDD2023	order_MDD2023	suborder_MDD2023	infraorder_MDD2023	parvorder_MDD2023	superfamily_MDD2023	Family_mdd2023	subfamily_MDD2023	tribe_MDD2023	genus_MDD2023	subgenus_MDD2023	specificEpithet_MDD2023	authoritySpeciesAuthor_MDD2023	authoritySpeciesYear_MDD2023	authorityParentheses_MDD2023	originalNameCombination_MDD2023	authoritySpeciesCitation_MDD2023	authoritySpeciesLink_MDD2023	holotypeVoucher_MDD2023	holotypeVoucherURIs_MDD2023	typeLocality_MDD2023	typeLocalityLatitude_MDD2023	typeLocalityLongitude_MDD2023	nominalNames_MDD2023	taxonomyNotes_MDD2023	taxonomyNotesCitation_MDD2023	distributionNotes_MDD2023	distributionNotesCitation_MDD2023	subregionDistribution_MDD2023	countryDistribution_MDD2023	continentDistribution_MDD2023	biogeographicRealm_MDD2023	iucnStatus_MDD2023	extinct_MDD2023	domestic_MDD2023	flagged_MDD2023	CMW_sciName_MDD2023	diffSinceCMW_MDD2023	MSW3_matchtype_MDD2023	MSW3_sciName_MDD2023	diffSinceMSW3_MDD2023	docOrigin_MDD2025	sciName	id	phylosort	mainCommonName	otherCommonNames	subclass	infraclass	magnorder	superorder	order	suborder	infraorder	parvorder	superfamily	family	subfamily	tribe	genus	subgenus	specificEpithet	authoritySpeciesAuthor	authorityParentheses	originalNameCombination	authoritySpeciesCitation	authoritySpeciesLink	typeVoucher	typeKind	typeVoucherURIs	typeLocality	typeLocalityLatitude	typeLocalityLongitude	taxonomyNotes	taxonomyNotesCitation	distributionNotes	distributionNotesCitation	subregionDistribution	countryDistribution	continentDistribution	biogeographicRealm	iucnStatus	extinct	domestic	flagged	CMW_sciName	diffSinceCMW	MSW3_matchtype	MSW3_sciName	diffSinceMSW3	docOrigin_batnames2025	Family	Genus	Subgenus	Species	Author	Date	Parentheses (1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L85	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Asellia tridens	Asellia tridens	Asellia tridens	Asellia tridens	Asellia tridens	Asellia tridens	Asellia tridens	Asellia tridens	Asellia tridens	Asellia tridens	Asellia tridens	Asellia tridens	Asellia tridens	Asellia tridens	Asellia tridens		[MSW3] Reviewed in part by Owen and Qumiseyeh (1987), Harrison and Bates (1991), Bates and Harrison (1997), and Horácek et al. (2000). Subspecies are poorly delimited, see Owen and Qumiseyeh (1987) and Kock et al. (2002).; [HMW] Rhinolophus tridens É. Geoffroy Saint-Hilaire, 1813 , near Luxor , Egypt . Up to four subspecies have at times been recognized within this species, but recent molecular work suggests the existence of just two. Two subspecies recognized.; [batnames2022] Does not include italosomalica ; see Benda et al. (2011). Reviewed by Benda et al. (2011) and in part by Owen and Qumiseyeh (1987), Harrison and Bates (1991), Bates and Harrison (1997), and Horácek etal. (2000). Subspecies are poorly delimited, see Owen and Qumiseyeh (1987), Kock et al. (2002), and Benda et al. (2011).; [MDD2022] previously included A. italosomalica; [batnames2023] Does not include italosomalica ; see Benda et al. (2011). Reviewed by Benda et al. (2011) and in part by Owen and Qumiseyeh (1987), Harrison and Bates (1991), Bates and Harrison (1997), and Horácek etal. (2000). Subspecies are poorly delimited, see Owen and Qumiseyeh (1987), Kock et al. (2002), and Benda et al. (2011).; [MDD2023] previously included A. italosomalica; [MDD2025_2.0] previously included A. italosomalica; [batnames2025_1.7] Does not include italosomalica; see Benda et al. (2011). Reviewed by Benda et al. (2011) and in part by Owen and Qumiseyeh (1987), Harrison and Bates (1991), Bates and Harrison (1997), and Horácek etal. (2000). Subspecies are poorly delimited, see Owen and Qumiseyeh (1987), Kock et al. (2002), and Benda et al. (2011).; [MDD2025_2.2] previously included A. italosomalica						diluta, italosomalica, murraiana, pallida.	diluta, italosomalica, tridens, murraiana	tridens, diluta, italosomalica, murraiana ,	diluta - pallida	tridens, murraiana		tridens, murraiana	tridens - diluta, pallida	tridens, murraiana, diluta, pallida		tridens, murraiana	tridens - diluta, pallida	tridens, murraiana, diluta, pallida	tridens, murraiana, diluta, pallida	pallida, murraiana, tridens	tridens - diluta	tridens (É. Geoffroy Saint-Hilaire, 1813)|murraiana (J. Anderson, 1881)|diluta Andersen, 1918|pallida P. Laurent, 1937|murriana Tate, 1941 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Trident bat	Morocco, Senegal – Pakistan	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Asellia tridens	Egypt, Qena, near Luxor.	E. Geoffroy	1813	Ann. Mus. Hist. Nat. Paris, 20:265.	Distribution: Same as for genus.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Trident bat	Morocco, Senegal – Pakistan	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	E. Geoffroy	1813	Ann. Mus. Hist. Nat. Paris, 20:265.		Pakistan to Arabia, Sinai peninsula (NE Egypt) and Israel; Egypt to Morocco, Senegal, Chad, Sudan and S Somalia; Socotra (Yemen); perhaps Zanzibar.	Egypt, Qena, near Luxor.		E. GEOFFROY	1818	Size relatively large. Forearm length, 43-53 mm; forearm slight ly shorter than combined length of third metacar pal and its first phalanx.	Distribution: Same as for genus.	Four poorly delimited subspecies are here recognized:	A. t. diluta (Senegambia to Morocco), A. t. italosomalica (Somalia, southern Ethiopia), A. t. tridens (Algeria to Ye men and Egypt), A. t. murraiana (remainder of range).	67	species	A. tridens	E. GEOFFROY	1818	Asellia	genus	Asellia tridens				Size relatively large. Forearm length, 43-53 mm; forearm slight ly shorter than combined length of third metacar pal and its first phalanx.	Four poorly delimited subspecies are here recognized:		1. A. tridens (E. GEOFFROY 1818).	1	_A. t. murraiana_ (Anderson, 1881); _A. t. tridens_ (Geoffroy Saint-Hilaire, 1813) (synonyms: _diluta_ Andersen, 1918, _pallida_ Laurent, 1937)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Hipposideridae			Asellia tridens	Asellia		tridens	E. Geoffroy	y	1813		Ann. Mus. Natn. Hist. Nat. Paris	20		265		Geoffroy's Trident Leaf-nosed Bat	Egypt, Qena, near Luxor.	Pakistan and Afganistan to Israel and Jordan, Iran, Iraq, Syria, Saudi Arabia, Sinai peninsula (NE Egypt), Socotra (Yemen) and Oman; Egypt to Morocco including S Lybia, Tunisia, and Algeria; Senegal, Mauritania, Gambia, Burkina Faso, Mali, Niger, Chad, Sudan, S Somalia, and Eritrea; perhaps Zanzibar.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	diluta Anderson, 1881; pallida Laurent, 1937; italosomalica De Beaux, 1931; murraiana Anderson, 1881.	Reviewed in part by Owen and Qumiseyeh (1987), Harrison and Bates (1991), Bates and Harrison (1997), and Horácek et al. (2000). Subspecies are poorly delimited, see Owen and Qumiseyeh (1987) and Kock et al. (2002).	03BD87A2C674A207F890F8BFFCE1472F	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Hipposideridae_210.pdf.imf	hash://md5/ff84ffdac676a204fff8ff9affef4346	229	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/BD/87/03BD87A2C674A207F890F8BFFCE1472F.xml	Asellia tridens	Hipposideridae	Asellia	tridens		1813	Aselliatrident @fr | Nordafrikanische Dreizackblattnase @de | Aselliatridente @es | Geoffroy’s Trident Bat @en	Rhinolophus tridens É. Geoffroy Saint-Hilaire, 1813 , near Luxor , Egypt . Up to four subspecies have at times been recognized within this species, but recent molecular work suggests the existence of just two. Two subspecies recognized.	A. t. tridens E. Geoffroy Saint-Hilaire, 1813 widespread in N Africa, from Morocco , Mauritania , and Senegal E to Egypt , Sudan , Eritrea , Ethiopia , Djibouti , and Somalia . A. t. murraianaj. Anderson, 1881 widely in the Middle East, including Syria , Iraq , Israel , Palestine , Jordan , Arabian Peninsula, and Iran , and E to W Afghanistan and S Pakistan .	Head-body 41-60 mm, tail 16-29 mm, ear 14-22 mm, hindfoot 8-10 mm, forearm 45-55 mm; weight 6-13 g. Geoffroy’s Trident Leaf-nosed Bat is the largest species in the genus. Muzzle is relatively short with large and distinctive noseleaf that has three subtriangular projections from its posterior margin. Pelage is beige or pale brownish gray dorsally, somewhat paler ventrally. Chromosomal complement has 2n = 50 and FN = 62.	Geoffroy’s Trident Leaf-nosed Bat inhabits a variety of desert and semi-desert habitats where it is mostly associated with oases and dry riverbeds. It generally occurs at low elevations, but may occur up to 2000 m .	Geoffroy’s Trident Leaf-nosed Bat forages by flying low and hawking a variety of large insects including beetles, moths, flies, and grasshoppers.	Apparently, Geoffroy’s Trident Leaf-nosed Bat is a seasonal breeder, giving birth to a single young in the boreal summer. In Africa, pregnant females have been reported in April and May, and dependent young in July. In the Middle East, females are pregnant in April-June, with births in June-July followed by 40 days of lactation.	Geoffroy’s Trident Leaf-nosed Bat roosts during the day in a wide range of situations including caves, mines, rock caverns, underground irrigation channels, and anthropogenic structures such as buildings, cellars, and tombs. It typically departs from its roost shortly after sunset, with males showing two peaks of activity during the night but females only one; both sexes have the greatest activity period in the hour immediately after sunset, with males showing a second peak in the early hours of the morning. Echolocation call includes a variable CF component of 115-120 kHz that varies individually and is negatively correlated with the size of the bat.	Geoffroy’s Trident Leaf-nosed Bat roosts in small to very large colonies, with up to 5000 individuals having been reported from a cave in Iran . Roosts are usually occupied by this species alone, although it may occasionally share its roost with other bats including the Egyptian Rousette ( Rousettus aegyptiacus), the Persian Trident Bat {Triaenopspersicus), and mouse-tailed bats {Rhinopoma spp.). Numbers of bats roosting typically vary with season, and sexual segregation has also been reported. Pregnant females form maternity roosts. Throughout much of the Middle East, bats disappear during the boreal winter, reappearing in April-May presumably because they migrate between summer and winter sites. Both sexes put on fat in the boreal autumn before migration. Bats move c.1-5-2 km between roosting and foraging sites. On following pages: 5. Greater Leaf-nosed Bat ( Doryrhina camerunensis }; 6.Telefomin Leaf-nosed Bat ( Doryrhina corynophyllus ); 7. Cyclops Leaf-nosed Bat ( Doryrhina cyclops ); 8. Hill's Leaf-nosed Bat ( Doryrhina edwardshillil; 9. Fly River Leaf-nosed Bat { Doryrhina muscinus ); 10. Semon's Leaf-nosed Bat [ Doryrhina semoni ); 11. Northern Leaf-nosed Bat [ Doryrhina stenotis }; 12. Wollaston's Leaf-nosed Bat ( Doryrhina wollastom); 13. Commerson's Leaf-nosed Bat ( Macronycteris commersoniii; 14. Madagascar Cryptic Leaf-nosed Bat ( Macronycteris cryptovalorona); 15. Giant Leaf-nosed Bat ( Macronycteris gigas ); 16. SaoTome Leaf-nosed Bat ( Macronycteris thomensis }; 17. Striped Leafnosed Bat ( Macronycteris vittatus ); 18. Dong BacTrident Bat ( Aselliscus dongbacanus ); 19. Stoliczka's Trident Bat ( Aselliscus stoliczkanusY, 20. Temminck ’s Trident Bat ( Aselliscus tricuspidatus ); 21. East Asian Tailless Leaf-nosed Bat ( Coelops frithii ); 22. Malayan Tailless Leaf-nosed Bat ( Coelops robinsoni ); 23. Solomons Leaf-nosed Bat ( Anthops ornatus }.	Classified as Least Concern on The IUCN Red, List. Its large distribution, often in remote areas and harsh landscapes, suggests that this species ought to be relatively secure.	Al-Robaae (1966) | Amichai et al. (2013) | Aulagnier (2013b) | Benda, Andreas et al. (2006) | Benda, Dietz et al. (2008) | Benda, Faizolâhi et al. (2012) | Benda, Lucan et al. (2010) | Benda, Spitzenberger et al. (2014) | Benda, Vallo & Reiter (2011) | Bogdanowicz & Owen (1998) | Bray & Benda (2016) | Brasset & Caubère (1960) | De-Blase (1980) | Feldman et al. (2000) | Gustafson & Schnitzler (1979) | Harrison & Bates (1991) | Jones et al. (1993) | Koch-Weser (1984) | Kock (1969d) | Kowalski & Rzebik-Kowalska (1991) | Fye (1972) | Qumsiyeh (1985) | Qumsiyeh & Schütter (1981) | Whitaker et al. (1994)	https://zenodo.org/record/3749047/files/figure.png	4. Geoffroy’s Trident Leaf-nosed Bat Asellia tridens French: Asellia trident / German: Nordafrikanische Dreizackblattnase / Spanish: Asellia tridente Other common names: Geoffroy’s Trident Bat Taxonomy. Rhinolophus tridens É. Geoffroy Saint-Hilaire, 1813 , near Luxor , Egypt . Up to four subspecies have at times been recognized within this species, but recent molecular work suggests the existence of just two. Two subspecies recognized. Subspecies and Distribution. A. t. tridens E. Geoffroy Saint-Hilaire, 1813 widespread in N Africa, from Morocco , Mauritania , and Senegal E to Egypt , Sudan , Eritrea , Ethiopia , Djibouti , and Somalia . A. t. murraianaj. Anderson, 1881 widely in the Middle East, including Syria , Iraq , Israel , Palestine , Jordan , Arabian Peninsula, and Iran , and E to W Afghanistan and S Pakistan . Descriptive notes. Head-body 41-60 mm, tail 16-29 mm, ear 14-22 mm, hindfoot 8-10 mm, forearm 45-55 mm; weight 6-13 g. Geoffroy’s Trident Leaf-nosed Bat is the largest species in the genus. Muzzle is relatively short with large and distinctive noseleaf that has three subtriangular projections from its posterior margin. Pelage is beige or pale brownish gray dorsally, somewhat paler ventrally. Chromosomal complement has 2n = 50 and FN = 62. Habitat. Geoffroy’s Trident Leaf-nosed Bat inhabits a variety of desert and semi-desert habitats where it is mostly associated with oases and dry riverbeds. It generally occurs at low elevations, but may occur up to 2000 m . Food and Feeding. Geoffroy’s Trident Leaf-nosed Bat forages by flying low and hawking a variety of large insects including beetles, moths, flies, and grasshoppers. Breeding. Apparently, Geoffroy’s Trident Leaf-nosed Bat is a seasonal breeder, giving birth to a single young in the boreal summer. In Africa, pregnant females have been reported in April and May, and dependent young in July. In the Middle East, females are pregnant in April-June, with births in June-July followed by 40 days of lactation. Activity patterns. Geoffroy’s Trident Leaf-nosed Bat roosts during the day in a wide range of situations including caves, mines, rock caverns, underground irrigation channels, and anthropogenic structures such as buildings, cellars, and tombs. It typically departs from its roost shortly after sunset, with males showing two peaks of activity during the night but females only one; both sexes have the greatest activity period in the hour immediately after sunset, with males showing a second peak in the early hours of the morning. Echolocation call includes a variable CF component of 115-120 kHz that varies individually and is negatively correlated with the size of the bat. Movements, Home range and Social organization. Geoffroy’s Trident Leaf-nosed Bat roosts in small to very large colonies, with up to 5000 individuals having been reported from a cave in Iran . Roosts are usually occupied by this species alone, although it may occasionally share its roost with other bats including the Egyptian Rousette ( Rousettus aegyptiacus), the Persian Trident Bat {Triaenopspersicus), and mouse-tailed bats {Rhinopoma spp.). Numbers of bats roosting typically vary with season, and sexual segregation has also been reported. Pregnant females form maternity roosts. Throughout much of the Middle East, bats disappear during the boreal winter, reappearing in April-May presumably because they migrate between summer and winter sites. Both sexes put on fat in the boreal autumn before migration. Bats move c.1-5-2 km between roosting and foraging sites. On following pages: 5. Greater Leaf-nosed Bat ( Doryrhina camerunensis }; 6.Telefomin Leaf-nosed Bat ( Doryrhina corynophyllus ); 7. Cyclops Leaf-nosed Bat ( Doryrhina cyclops ); 8. Hill's Leaf-nosed Bat ( Doryrhina edwardshillil; 9. Fly River Leaf-nosed Bat { Doryrhina muscinus ); 10. Semon's Leaf-nosed Bat [ Doryrhina semoni ); 11. Northern Leaf-nosed Bat [ Doryrhina stenotis }; 12. Wollaston's Leaf-nosed Bat ( Doryrhina wollastom); 13. Commerson's Leaf-nosed Bat ( Macronycteris commersoniii; 14. Madagascar Cryptic Leaf-nosed Bat ( Macronycteris cryptovalorona); 15. Giant Leaf-nosed Bat ( Macronycteris gigas ); 16. SaoTome Leaf-nosed Bat ( Macronycteris thomensis }; 17. Striped Leafnosed Bat ( Macronycteris vittatus ); 18. Dong BacTrident Bat ( Aselliscus dongbacanus ); 19. Stoliczka's Trident Bat ( Aselliscus stoliczkanusY, 20. Temminck ’s Trident Bat ( Aselliscus tricuspidatus ); 21. East Asian Tailless Leaf-nosed Bat ( Coelops frithii ); 22. Malayan Tailless Leaf-nosed Bat ( Coelops robinsoni ); 23. Solomons Leaf-nosed Bat ( Anthops ornatus }. Status and Conservation. Classified as Least Concern on The IUCN Red, List. Its large distribution, often in remote areas and harsh landscapes, suggests that this species ought to be relatively secure. Bibliography. Al-Robaae (1966), Amichai et al. (2013), Aulagnier (2013b), Benda, Andreas et al. (2006), Benda, Dietz et al. (2008), Benda, Faizolâhi et al. (2012), Benda, Lucan et al. (2010), Benda, Spitzenberger et al. (2014), Benda, Vallo & Reiter (2011), Bogdanowicz & Owen (1998), Bray & Benda (2016), Brasset & Caubère (1960), De-Blase (1980), Feldman et al. (2000), Gustafson & Schnitzler (1979), Harrison & Bates (1991), Jones et al. (1993), Koch-Weser (1984), Kock (1969d), Kowalski & Rzebik-Kowalska (1991), Fye (1972), Qumsiyeh (1985), Qumsiyeh & Schütter (1981), Whitaker et al. (1994).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Hipposideridae	Asellia tridens	Asellia		tridens	E. Geoffroy	1813	1	Ann. Mus. Natn. Hist. Nat. Paris	20: 265, pl. 5	Geoffroy's Trident Leaf-nosed Bat	 diluta </b>Anderson, 1881; pallida </b>Laurent, 1937; <b> murraiana </b>Anderson, 1881.	Egypt, Qena, near Luxor.	Pakistan and Afganistan to Israel and Jordan, Iran, Iraq, Syria, Saudi Arabia, Sinai peninsula (NE Egypt), Socotra (Yemen) and Oman; Egypt to Morocco including S Lybia, Tunisia, and Algeria; Senegal, Mauritania, Gambia, Burkina Faso, Mali, Niger, Chad, Sudan, S Somalia, and Eritrea; perhaps Zanzibar.	Not listed.	Least Concern	Does not include italosomalica ; see Benda et al. (2011). Reviewed by Benda et al. (2011) and in part by Owen and Qumiseyeh (1987), Harrison and Bates (1991), Bates and Harrison (1997), and Horácek etal. (2000). Subspecies are poorly delimited, see Owen and Qumiseyeh (1987), Kock et al. (2002), and Benda et al. (2011).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Asellia tridens	23	Geoffroy's Trident Leaf-nosed Bat	Geoffroy's Trident Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	HIPPOSIDERIDAE	NA	NA	Asellia	NA	tridens	É. Geoffroy Saint-Hilaire	1813	1	Rhinolophus_tridens	Geoffroy Saint-Hilaire, É. (1813). Sur un genre de chauve-souris, sous le nom de Rhinolophes (1). Annales du Muséum d'histoire naturelle, 20, 260.	https://www.biodiversitylibrary.org/item/23442#page/278/mode/1up	MNHN 1986-1068		near Luxor, Egypt.			tridens (É. Geoffroy Saint-Hilaire, 1813)|murraiana (J. Anderson, 1881)|diluta K. Andersen, 1918|pallida Laurent, 1937	previously included A. italosomalica	Benda, P., Vallo, P., & Reiter, A. (2011). Taxonomic revision of the genus Asellia (Chiroptera: Hipposideridae) with a description of a new species from southern Arabia. Acta Chiropterologica, 13(2), 245-270.	Senegal|Gambia|Mauritania|Mali|Morocco|Burkina Faso|Algeria|Tunisia|Niger|Libya|Chad|Sudan|South Sudan|Ethiopia|Somalia|Eritrea|Djibouti|Egypt|Israel|Palestine|Jordan|Lebanon?|Saudi Arabia|Oman|Yemen|United Arab Emirates?|Qatar?|Kuwait?|Iraq|Syria|Iran|Afghanistan|Pakistan	Africa|Asia	Afrotropic|Palearctic	LC	0	0	0	Asellia_tridens	0	sciname match	Asellia_tridens	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Asellia tridens	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	HIPPOSIDERIDAE	Asellia	tridens	(É. Geoffroy, 1813)		20000000	Asellia tridens	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	This bat is listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This is a gregarious and colonial species which occurs in crevices or in cliffs in arid and semi-desert habitats. It roosts in temples, caves, mines, open-wells, underground irrigation tunnels and old tombs and buildings. Forages over desert and semi-desert vegetation zones, mainly in oases. Forages by slow hawking, has been observed foraging around palm trees and buildings, and over water.	The main threat is the widespread use of pesticides against locusts. Human disturbance in caves and old buildings is affecting some populations.	It is a very common species, found in colonies of up to several hundred in North Africa, and in its Asian distribution it has been found in groups up to 5,000 animals.	Stable	Geoffroy's Trident Leaf-nosed Bat ranges widely in the Sahara, through the Arabian peninsula and the Middle East, to Afghanistan and Pakistan. It is absent from the northern parts of Morocco and Algeria, and Tunisia, and occurs south to Ethiopia and Somalia.		Terrestrial	It presumably occurs in several protected areas. Underground roost management is needed in some places. A study on the impacts of pesticides is required, especially ways in which the impact might be minimised.	Afrotropical|Indomalayan|Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Hipposideridae	Asellia		tridens	É. Geoffroy Saint-Hilaire	1813	1	Ann. Mus. Natn. Hist. Nat. Paris	20: 265, pl. 5	Geoffroy's Trident Leaf-nosed Bat	 diluta </b>Anderson, 1881; pallida </b>Laurent, 1937; <b> murraiana </b>Anderson, 1881.	Egypt, Qena, near Luxor.	Pakistan and Afganistan to Israel and Jordan, Iran, Iraq, Syria, Saudi Arabia, Sinai peninsula (NE Egypt), Socotra (Yemen) and Oman; Egypt to Morocco including S Lybia, Tunisia, and Algeria; Senegal, Mauritania, Gambia, Burkina Faso, Mali, Niger, Chad, Sudan, S Somalia, and Eritrea; perhaps Zanzibar.	Not listed.	Least Concern	Does not include italosomalica ; see Benda et al. (2011). Reviewed by Benda et al. (2011) and in part by Owen and Qumiseyeh (1987), Harrison and Bates (1991), Bates and Harrison (1997), and Horácek etal. (2000). Subspecies are poorly delimited, see Owen and Qumiseyeh (1987), Kock et al. (2002), and Benda et al. (2011).	Asellia tridens	1004559	23	Geoffroy's Trident Leaf-nosed Bat	Geoffroy's Trident Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	PTEROPODIFORMES	NA	NA	RHINOLOPHOIDEA	Hipposideridae	NA	NA	Asellia	NA	tridens	É. Geoffroy Saint-Hilaire	1813	1	Rhinolophus_tridens	Geoffroy Saint-Hilaire, É. (1813). Sur un genre de chauve-souris, sous le nom de Rhinolophes (1). Annales du Muséum d'histoire naturelle, 20, 260.	https://www.biodiversitylibrary.org/item/23442#page/278/mode/1up	MNHN 1986-1068		near Luxor, Egypt.			tridens (É. Geoffroy Saint-Hilaire, 1813)|murraiana (J. Anderson, 1881)|diluta K. Andersen, 1918|pallida Laurent, 1937	previously included A. italosomalica	Benda, P., Vallo, P., & Reiter, A. (2011). Taxonomic revision of the genus Asellia (Chiroptera: Hipposideridae) with a description of a new species from southern Arabia. Acta Chiropterologica, 13(2), 245-270.				Senegal|Gambia|Mauritania|Mali|Morocco|Burkina Faso|Algeria|Tunisia|Niger|Libya|Chad|Sudan|South Sudan|Ethiopia|Somalia|Eritrea|Djibouti|Egypt|Israel|Palestine|Jordan|Lebanon?|Saudi Arabia|Oman|Yemen|United Arab Emirates?|Qatar?|Kuwait?|Iraq|Syria|Iran|Afghanistan|Pakistan	Africa|Asia	Afrotropic|Palearctic	LC	0	0	0	Asellia_tridens	0	sciname match	Asellia_tridens	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Asellia_tridens	1004559	23	Geoffroy's Trident Roundleaf Bat	Geoffroy's Trident Bat|Geoffroy's Trident Leaf-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yinpterochiroptera	NA	NA	Rhinolophoidea	Hipposideridae	NA	NA	Asellia	NA	tridens	É. Geoffroy Saint-Hilaire	1	Rhinolophus tridens	Geoffroy Saint-Hilaire, É. 1813-03. Description des mammifères qui se trouvent en Égypte. Pp. 99–144 in Anonymous. 1818. Description de l'Égypte, ou Recueil des observations et des recherches qui ont été faites en Égypte pendant l'expédition de l'Armée française, publié par les ordres de sa Majesté l'Empéreur Napoléon le Grand. Histoire naturelle. Tome second. Imprimerie Impériale, Paris, 752 pp.	https://www.biodiversitylibrary.org/page/15953719	MNHN-ZM-MO-1986-1068 (= MNHN A. 235)	syntypes	http://coldb.mnhn.fr/catalognumber/mnhn/zm/mo-1986-1068	near Luxor, Egypt.	25.74028	32.60167	previously included A. italosomalica	Benda, P., Vallo, P., & Reiter, A. (2011). Taxonomic revision of the genus Asellia (Chiroptera: Hipposideridae) with a description of a new species from southern Arabia. Acta Chiropterologica, 13(2), 245-270.				Senegal|Gambia|Mauritania|Mali|Morocco|Burkina Faso|Algeria|Tunisia|Niger|Libya|Chad|Sudan|South Sudan|Ethiopia|Somalia|Eritrea|Djibouti|Egypt|Israel|Palestine|Jordan|Lebanon?|Saudi Arabia|Oman|Yemen|United Arab Emirates?|Qatar?|Kuwait?|Iraq|Syria|Iran|Afghanistan|Pakistan	Africa|Asia	Afrotropic|Palearctic	LC	0	0	0	Asellia_tridens	0	sciname match	Asellia_tridens	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Hipposideridae	Asellia		tridens	É. Geoffroy Saint-Hilaire	1813	1	Ann. Mus. Natn. Hist. Nat. Paris	20: 265, pl. 5	Geoffroy's Trident Leaf-nosed Bat	diluta Anderson, 1881; pallida Laurent, 1937; murraiana Anderson, 1881.	Egypt, Qena, near Luxor.	Pakistan and Afganistan to Israel and Jordan, Iran, Iraq, Syria, Saudi Arabia, Sinai peninsula (NE Egypt), Socotra (Yemen) and Oman; Egypt to Morocco including S Lybia, Tunisia, and Algeria; Senegal, Mauritania, Gambia, Burkina Faso, Mali, Niger, Chad, Sudan, S Somalia, and Eritrea; perhaps Zanzibar.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/80221529/21975715/' target='_blank'>Least Concern</a>	Does not include italosomalica; see Benda et al. (2011). Reviewed by Benda et al. (2011) and in part by Owen and Qumiseyeh (1987), Harrison and Bates (1991), Bates and Harrison (1997), and Horácek etal. (2000). Subspecies are poorly delimited, see Owen and Qumiseyeh (1987), Kock et al. (2002), and Benda et al. (2011).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Asellia tridens; Asellia tridens; Asellia tridens; Asellia tridens; Asellia tridens; Asellia tridens; tridens ; diluta ; italosomalica ; murraiana ; diluta - pallida; tridens; murraiana; murraiana; diluta; pallida; tridens; murraiana; diluta; pallida; Aselliatrident; Nordafrikanische Dreizackblattnase; Aselliatridente; Geoffroy’s Trident Bat; Geoffroy's Trident Leaf-nosed Bat; Geoffroy's Trident Bat; Geoffroy's Trident Leaf-nosed Bat; Geoffroy's Trident Leaf-nosed Bat; A. tridens