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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L838	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis brandti	Myotis brandti	Myotis brandtii	Myotis brandti	Myotis brandti	Myotis brandtii	Myotis brandtii	Myotis brandtii	Myotis brandtii	Myotis brandtii	Myotis brandtii	Myotis brandtii	Myotis brandtii	Myotis brandtii	Myotis brandtii		[MSW2] Subgenus Selysius. Listed as a subspecies of mystacinus by Ellerman and Morrison-Scott (1951:139); but also see review by Strelkov and Buntova (1982). Includes fujiensis and gracilis, but see Yoshiyuki (1989).; [MSW3] Listed as a subspecies of mystacinus by Ellerman and Morrison-Scott (1951), but see Strelkov and Buntova (1982) and Benda and Tsytsulina (2000). Does not include fujiensis, see Yoshiyuki (1989), Benda and Tsytsulina (2000), and Horácek et al. (2000). Includes gracilis, see Benda and Tsytsulina (2000), but also see Yoshiyuki (1989). Horácek et al. (2000) provisionally treated gracilis as distinct from brandtii and fujiensis pending further study. See also Yoon and Son (1989). Sometimes misspelled brandti, but brandtii is the original spelling.; [HMW] Vespertilio brandtii Eversmann, 1845 , Orenburgsk, Russia . Subgenus Pizonyx; brandtii species group (2 species). See M. sibiricus . Monotypic.; [batnames2022] Listed as a subspecies of mystacinus by Ellerman and Morrison-Scott (1951), but see Strelkov and Buntova (1982) and Benda andTsytsulina (2000). Does not include fujiensis , see Yoshiyuki (1989), Benda and Tsytsulina (2000), and Horácek et al. (2000). Does notinclude gracilis or sibiricus , see Kruskop et al. 2012 who suggested that sibiricus may be the correct name for gracilis. Horácek et al. (2000) provisionally treated gracilis as distinct from brandtii and fujiensis pending further study. See also Yoon and Son (1989). Sometimes misspelled brandti , but brandtii is the original spelling.; [MDD2022] previously included M. sibiricus (including gracilis); [IUCN] <span>In previous assessments it included also Myotis brandtii gracilis now raised to species level, as M. sibiricus (Kruskop et al . 2012).; [batnames2023] Listed as a subspecies of mystacinus by Ellerman and Morrison-Scott (1951), but see Strelkov and Buntova (1982) and Benda andTsytsulina (2000). Does not include fujiensis , see Yoshiyuki (1989), Benda and Tsytsulina (2000), and Horácek et al. (2000). Does notinclude gracilis or sibiricus , see Kruskop et al. 2012 who suggested that sibiricus may be the correct name for gracilis. Horácek et al. (2000) provisionally treated gracilis as distinct from brandtii and fujiensis pending further study. See also Yoon and Son (1989). Sometimes misspelled brandti , but brandtii is the original spelling.; [MDD2023] previously included M. sibiricus (including gracilis); [MDD2025_2.0] previously included M. sibiricus (including gracilis); [batnames2025_1.7] Listed as a subspecies of mystacinus by Ellerman and Morrison-Scott (1951), but see Strelkov and Buntova (1982) and Benda andTsytsulina (2000). Does not include fujiensis, see Yoshiyuki (1989), Benda and Tsytsulina (2000), and Horácek et al. (2000). Does notinclude gracilis or sibiricus, see Kruskop et al. 2012 who suggested that sibiricus may be the correct name for gracilis. Horácek et al. (2000) provisionally treated gracilis as distinct from brandtii and fujiensispending further study. See also Yoon and Son (1989). Sometimes misspelled brandti, but brandtii is the original spelling.; [MDD2025_2.2] previously included M. sibiricus (including gracilis)						aureus, fujiensis, gracilis, sibiricus.	brandti, gracilis, fujiensis	brandtii, gracilis	aureus, coluotus, sibiricus			brandtii 	brandtii - aureus, coluotus	brandtii, aureus, coluotus	<span>In previous assessments it included also Myotis brandtii gracilis now raised to species level, as M. sibiricus (Kruskop et al . 2012).	brandtii 	brandtii - aureus, coluotus	brandtii, aureus, coluotus	brandtii, aureus, coluotus, brandti	brandtii 	brandtii - aureus, coluotus	brandtii (Eversmann, 1845)|aureus (C. Koch, 1863)|coluotus Kostroň, 1943|brandti Corbet & J. Edwards Hill, 1980 [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Spain, Britain – Urals	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis brandti	U.S.S.R., Orenburgsk. Obi., S. Ural, Sakmara River (OLR). "Foothills of the Ural Mountains."	Eversmann	1845	Bull. Soc. Nat. Moscow, 18:505.	Distribution: Ranging across Eurasia from Britain and Spain to eastern Siberia and Korea, also Sakhalin, Japan, and the Kuriles.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Brandt's bat	France, Britain – E Siberia, Hokkaido; ref. 4.127	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Eversmann	1845	Bull. Soc. Nat. Moscow, 18:505.	Subgenus Selysius. Listed as a subspecies of mystacinus by Ellerman and Morrison-Scott (1951:139); but also see review by Strelkov and Buntova (1982). Includes fujiensis and gracilis, but see Yoshiyuki (1989).	Britain to Kazakhstan, E Siberia including Sakhalin Isis, Kamchatka Peninsula and Kurile Isis (Russia), Mongolia south to Spain, Greece, Korea, and Ussuri region (Russia); Japan.	Russia, Orenburgsk. Obi., S. Ural, Bolshoi-Ik River, Spasskoie. Foothills of the Ural Mountains.		EVERSMANN	1845	Size small (forearm length, 32-38 mm; condylobasal length, 13-14 mm). Penis clubshaped and baculum with a basal notch.	Distribution: Ranging across Eurasia from Britain and Spain to eastern Siberia and Korea, also Sakhalin, Japan, and the Kuriles.	Three subspecies are here recognized:	M. b. brandti (Europe and western Siberia), M. b. gracilis (eastern Siberia to Korea, Sakhalin, Kuriles, and Hokkaido), M. b.fujiensis (Honshu).	103	species	M. brandti	EVERSMANN	1845	Selysius	subgenus	Myotis brandti				Size small (forearm length, 32-38 mm; condylobasal length, 13-14 mm). Penis clubshaped and baculum with a basal notch.	Three subspecies are here recognized:		22. M. brandti (EVERSMANN 1845) [mystacinus group].	22	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis brandtii	Myotis		brandtii	Eversmann	y	1845		Bull. Soc. Nat. Moscow	18	1	505		Brandt's Myotis	Russia, Orenburgsk. Obl., S. Ural, Bolshoi-Ik River, Spasskoie. Foothills of the Ural Mountains.	Britain south to Italy, Greece, and Bulgaria; east to Kazakhstan and Mongolia, E Siberia including Sakhalin Isls, Kamchatka Peninsula and Kurile Isls; Ussuri region (Russia); Korea.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	aureus Koch, 1865; coluotus Kostron, 1943; sibiricus Kastschenko, 1905; gracilis Ognev, 1927.	Listed as a subspecies of mystacinus by Ellerman and Morrison-Scott (1951), but see Strelkov and Buntova (1982) and Benda and Tsytsulina (2000). Does not include fujiensis, see Yoshiyuki (1989), Benda and Tsytsulina (2000), and Horácek et al. (2000). Includes gracilis, see Benda and Tsytsulina (2000), but also see Yoshiyuki (1989). Horácek et al. (2000) provisionally treated gracilis as distinct from brandtii and fujiensis pending further study. See also Yoon and Son (1989). Sometimes misspelled brandti, but brandtii is the original spelling.	4C3D87E8FF436AFCFF4E90BA16A2BFD9	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	946	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF436AFCFF4E90BA16A2BFD9.xml	Myotis brandtii	Vespertilionidae	Myotis	brandtii		1845	Murin de Brandt @fr | Gro Re Bartfledermaus @de | Ratonero de Brandt @es | Brandt's Bat @en	Vespertilio brandtii Eversmann, 1845 , Orenburgsk, Russia . Subgenus Pizonyx; brandtii species group (2 species). See M. sibiricus . Monotypic.	N & C Europe from S Fennoscandia and Great Britain to SE Europe and W Siberia, isolated populations in C Italy , Turkey , Caucasus, and C Asia. Exact distribution is uncertain because of confusion with field identification with the Common Whiskered Myotis ( M. mystacinus ).	Head-body 39-51 mm , tail 32-44 mm , ear 12-17 mm , hindfoot 7-9, forearm 33-38- 2 mm ; weight 5-7 g . Brandt's Myotis has remarkably long and wide-spaced ears, resembling its sister species, the Common Whiskered Myotis . It lacks postcalcariallobe to membrane. Fur is typically brownish or blackish, with golden tips. Venteris only slightly paler than dorsum. Brownish color becomes more evident with age; juveniles are almost completely black. Bare skin is darker than on other species such as Daubenton’s Myotis (M. daubentonit). Male Brandt's Myotis can be easily distinguished from other cryptic species such as the Common Whiskered Myotis by its uniformly thinner penis. Skull is slender and delicate, but large with flat frontal and parietal regions. I* has shallow groove or tiny protuberance on palatal edge of cingulum; P? and P, are comparatively robust and high, P, usually reaching two-thirds height of P,; P_ is slightly intruded from tooth row; paraconules are usually present on molars. Chromosomal complement has 2n = 44 and FNa = 52 ( Germany ).	Mixed, coniferous, and broadleaf forests, usually close to aquatic habitats such as swamps, lakes, moors, or damp areas, from sea level up to elevations of ¢. 1800 m . In the southernmost part of its distribution, Brandt's Myotis is more commonly found in the mountain areas.	Although Brandt's Myotis feeds close to aquatic habitats, it feeds on terrestrial insects. Typical foraging habitats include forest edges and tree lines. It uses aerial hawking to capture insects in flight. It is very agile and maneuverable, especially in light riparian, beech, and oak forests. It also hunts above tree canopies. Its most common prey is moths, spiders, midges, mosquitoes, harvestmen, and earwigs.	Maternity colonies of Brandt's Myotis are commonly found in buildings, attics of houses, or artificial empty cavities in roofs but also tree holes and bat boxes. Maternity colonies in natural roosts are formed by several tens of females, with some reports with up to 200 individuals. In bat boxes, they rarely exceed 20 individuals, sometimes being mixed with some pipistrelle bats. Young (normally single but sometimes twins) are usually born during in June, with dates varying latitudinally. Females became sexually active only in their second year oflife. Brandt's Myotis holds the record for bat longevity in the wild at 41 years in Siberia.	In summer, Brandt’s Myotis roost in tree holes, cracks, crevices in trunks, between timbers, or behind peeling bark. It seems to prefer roosting far from human settlements. Echolocation is clearly Myotislike, with broadband, highly modulated pulses. Calls are c.4-7 milliseconds long at 20-100 kHz.	Brandt's Myotis is reported to migrate up to 300 km , or it can be locally sedentary. It usually moves from summer roosts to hibernation sites that tend to be caves, mines, or cellars with small numbers and clusters. It starts dispersing in July, earlier than in other species, and males occupy swarmingsites early in August. During summer, one colony foraged in up to 13 different areas of 1-4 ha each, totally more than 100 ha.	Classified as Least Concern on The IUCN Red List. Brandt's Myotis is widespread, occurs at high densities, and is fairly common in most of its distribution, especially in northern areas. It is decreasing only in northern Turkey at the border ofits distribution.	Abe et al. (2005) | Agnelli et al. (2004) | Bannikov (1954) | Benda &Tsytsulina (2000) | Boston, Buckley et al. (2010) | Boston, Hanrahan et al. (2011) | Ekman & de Jong (1996) | Gerell (1999) | von Helversen et al. (2001) | Horacek et al. (2000) | Hutson, Spitzenberger, Coroiu et al. (2008) | Hutterer et al. (2005) | Jiang Jianjun et al. (2016) | Koopman (1993) | Podlutsky et al. (2005) | Ruprecht (1974) | Seim et al. (2013) | Smith & Xie Yan (2008) | Sokolov & Orlov (1980) | Tinnin et al. (2002) | Toffoli (2007) | Tsytsulina (2001) | Tupinier (2001) | Vlaschenko, Hukov et al. (2016) | Volleth & Heller (2012) | Won Chang-Man & Smith (1999)	https://zenodo.org/record/6398813/files/figure.png	418. Brandt's Myotis Myotis brandtii French: Murin de Brandt / German: GroRe Bartfledermaus / Spanish: Ratonero de Brandt Other common names: Brandt's Bat Taxonomy. Vespertilio brandtii Eversmann, 1845 , Orenburgsk, Russia . Subgenus Pizonyx; brandtii species group (2 species). See M. sibiricus . Monotypic. Distribution. N & C Europe from S Fennoscandia and Great Britain to SE Europe and W Siberia, isolated populations in C Italy , Turkey , Caucasus, and C Asia. Exact distribution is uncertain because of confusion with field identification with the Common Whiskered Myotis ( M. mystacinus ). Descriptive notes. Head-body 39-51 mm , tail 32-44 mm , ear 12-17 mm , hindfoot 7-9, forearm 33-38- 2 mm ; weight 5-7 g . Brandt's Myotis has remarkably long and wide-spaced ears, resembling its sister species, the Common Whiskered Myotis . It lacks postcalcariallobe to membrane. Fur is typically brownish or blackish, with golden tips. Venteris only slightly paler than dorsum. Brownish color becomes more evident with age; juveniles are almost completely black. Bare skin is darker than on other species such as Daubenton’s Myotis (M. daubentonit). Male Brandt's Myotis can be easily distinguished from other cryptic species such as the Common Whiskered Myotis by its uniformly thinner penis. Skull is slender and delicate, but large with flat frontal and parietal regions. I* has shallow groove or tiny protuberance on palatal edge of cingulum; P? and P, are comparatively robust and high, P, usually reaching two-thirds height of P,; P_ is slightly intruded from tooth row; paraconules are usually present on molars. Chromosomal complement has 2n = 44 and FNa = 52 ( Germany ). Habitat. Mixed, coniferous, and broadleaf forests, usually close to aquatic habitats such as swamps, lakes, moors, or damp areas, from sea level up to elevations of ¢. 1800 m . In the southernmost part of its distribution, Brandt's Myotis is more commonly found in the mountain areas. Food and Feeding. Although Brandt's Myotis feeds close to aquatic habitats, it feeds on terrestrial insects. Typical foraging habitats include forest edges and tree lines. It uses aerial hawking to capture insects in flight. It is very agile and maneuverable, especially in light riparian, beech, and oak forests. It also hunts above tree canopies. Its most common prey is moths, spiders, midges, mosquitoes, harvestmen, and earwigs. Breeding. Maternity colonies of Brandt's Myotis are commonly found in buildings, attics of houses, or artificial empty cavities in roofs but also tree holes and bat boxes. Maternity colonies in natural roosts are formed by several tens of females, with some reports with up to 200 individuals. In bat boxes, they rarely exceed 20 individuals, sometimes being mixed with some pipistrelle bats. Young (normally single but sometimes twins) are usually born during in June, with dates varying latitudinally. Females became sexually active only in their second year oflife. Brandt's Myotis holds the record for bat longevity in the wild at 41 years in Siberia. Activity patterns. In summer, Brandt’s Myotis roost in tree holes, cracks, crevices in trunks, between timbers, or behind peeling bark. It seems to prefer roosting far from human settlements. Echolocation is clearly Myotislike, with broadband, highly modulated pulses. Calls are c.4-7 milliseconds long at 20-100 kHz. Movements, Home range and Social organization. Brandt's Myotis is reported to migrate up to 300 km , or it can be locally sedentary. It usually moves from summer roosts to hibernation sites that tend to be caves, mines, or cellars with small numbers and clusters. It starts dispersing in July, earlier than in other species, and males occupy swarmingsites early in August. During summer, one colony foraged in up to 13 different areas of 1-4 ha each, totally more than 100 ha. Status and Conservation. Classified as Least Concern on The IUCN Red List. Brandt's Myotis is widespread, occurs at high densities, and is fairly common in most of its distribution, especially in northern areas. It is decreasing only in northern Turkey at the border ofits distribution. Bibliography. Abe et al. (2005), Agnelli et al. (2004), Bannikov (1954), Benda &Tsytsulina (2000), Boston, Buckley et al. (2010), Boston, Hanrahan et al. (2011), Ekman & de Jong (1996), Gerell (1999), von Helversen et al. (2001), Horacek et al. (2000), Hutson, Spitzenberger, Coroiu et al. (2008), Hutterer et al. (2005), Jiang Jianjun et al. (2016), Koopman (1993), Podlutsky et al. (2005), Ruprecht (1974), Seim et al. (2013), Smith & Xie Yan (2008), Sokolov & Orlov (1980), Tinnin et al. (2002), Toffoli (2007), Tsytsulina (2001), Tupinier (2001), Vlaschenko, Hukov et al. (2016), Volleth & Heller (2012), Won Chang-Man & Smith (1999).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis brandtii	Myotis	Unassigned-Myotis	brandtii	Eversmann	1845	1	Bull. Soc. Nat. Moscow	18(1): 505	Brandt's Myotis	 aureus Koch, 1865; coluotus Kostron, 1943.	Russia, Orenburgsk. Obl., S. Ural, Bolshoi-Ik River, Spasskoie. Foothills of the Ural Mountains.	Britain south to Italy, Greece, and Bulgaria; east to Kazakhstan and Mongolia, E Siberia including Sakhalin Isls, Kamchatka Peninsula and Kurile Isls; Ussuri region (Russia); Korea.	Not listed.	Least Concern	Listed as a subspecies of mystacinus by Ellerman and Morrison-Scott (1951), but see Strelkov and Buntova (1982) and Benda andTsytsulina (2000). Does not include fujiensis , see Yoshiyuki (1989), Benda and Tsytsulina (2000), and Horácek et al. (2000). Does notinclude gracilis or sibiricus , see Kruskop et al. 2012 who suggested that sibiricus may be the correct name for gracilis. Horácek et al. (2000) provisionally treated gracilis as distinct from brandtii and fujiensis pending further study. See also Yoon and Son (1989). Sometimes misspelled brandti , but brandtii is the original spelling.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis brandtii	23	Brandt's Myotis	Brandt's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	brandtii	Eversmann	1845	1						Orenburgsk, Russia.			brandtii (Eversmann, 1845)|aureus (L. Koch, 1865)|coluotus Kostron, 1943	previously included M. sibiricus (including gracilis)	Ohdachi, S. D. I, Ishibashi, Y., Iwasa, M. A., & Saitoh, Takashi (2009). The Wild Mammals of Japan, Shoukadoh, Kyoto.	Norway|Sweden|Finland|United Kingdom|France|Netherlands|Belgium|Luxembourg|Germany|Denmark|Switzerland|Liechtenstein?|Italy|Austria|Czech Republic|Poland|Slovakia|Hungary|Slovenia|Croatia|Bosnia & Herzegovina?|Serbia|Kosovo|Albania|North Macedonia?|Albania|Greece|Bulgaria|Romania|Moldova|Ukraine|Belarus|Lithuania|Latvia|Estonia|Russia|Georgia|Turkey|Kazakhstan	Asia|Europe	Palearctic	LC	0	0	0	Myotis_brandtii	0	sciname match	Myotis_brandtii	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Myotis brandtii	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	brandtii	(Eversmann, 1845)	<span>In previous assessments it included also Myotis brandtii gracilis now raised to species level, as M. sibiricus (Kruskop et al . 2012).	200000000	Myotis brandtii	Least Concern		2020	2019-05-13 00:00:00 UTC	3.1	English	As there are no obvious threats at the global level the species is listed as Least Concern. Only negative population trend or reduction of the range were reported for Poland, Russia and Ukraine.	Maternity colonies can comprise up to 350 females and use voids and crevices in buildings (e.g. behind open shatters) or trees, less frequently bat and bird boxes for roosting. These colonies may change roosts, therefore their availability is very important. The species feeds in woodlands, also above and along stagnant water, rivers and streams. In northern Europe coniferous forests are favoured over other forest types as well as over grassland and arable land. In the south of Europe the species can only be found in mountainous woodlands and rarely in urbanised habitats. It forages in areas from 1.5 km up to 10 km from the roost. M. brandtii preys mainly on small Lepidoptera and Diptera (see the literature review in Kyheröinen et al. 2019). Seasonal movements from summer roosts to hibernacula usually don't exceed several tens of kilometres, with the longest distance of 618 km (Hutterer et al. 2005).	Loss of roosting opportunities in building during renovation and refurbishment works, intentional evictions and closure of roost entrances. Removal of standing old trees with cavities and dead wood in forestry operations reduces the number of occupied and alternative roosts. Disturbance in winter quarters (caves, tunnels, abandoned mines) by visitors/vandalism, pesticide use and wood treatment in attics and roofs. The species is sensitive to light pollution at roosts, commuting routes and habitats (Simon et al. 2012, Voigt et al. 2018). Fragmentation of foraging areas and loss of linear structures which are utilised for commuting may impair habitats, especially around maternity colonies (Kyheröinen et al. 2019). At high risk of collisions with vehicles (Fensome, Mathews 2016).	In the EU, the species has unfavourable-inadequate status in Continental and Pannonian biogeographical regions, unfavourable-bad in Atlantic and unknown in Alpine, Boreal, and Mediterranean regions. Only the Netherlands indicated unfavourable-bad status for the Atlantic region (EEA 2013). A negative trend was observed in Belarus (Shpak 2018), while it is positive in Poland (Poland 2018) and rather stable in Ukraine (Ukraine 2018) and in Russia (S. Gazaryan, pers. comm. 2019).	Stable	A woodland, predominantly tree-dwelling Palaearctic species, distributed from Great Britain, western and central Europe and Fennoscandia across the Russian Plain, over the Urals up to the Ob River basin in the east. A distinct Asian species Myotis sibiricus that was considered earlier within M. brandtii occurs further eastward. The identity of M. sibiricu s was ascertained by the mtDNA analysis (Kawai et al.  2003, Kruskop et al.  2012) and recently confirmed also by the nuclear DNA markers (S. Kruskop, pers. comm.) M. brandtii occurs in woodlands across the Caucasus, including Hyrcanian forests in Azerbaijan at the border with Iran (Gazaryan 2003). Based on the skull morphology, Benda and Karatash (2005) mention a finding from central Anatolia. In the Balkans, presence of this species has not yet been confirmed only in Macedonia (M. Paunovic pers. comm. 2016). There are few records from Italy, from the Alpe Veglia and Alpe Devero Natural Park (Piedmont, northern Italy, Toffoli 2006) and from the Abruzzo, Lazio and Molise National Park (central Italy, Agnelli et al.  2004, Russo pers. comm. 2006).		Terrestrial	It is protected by national legislation in most of the main ; range states, except Russia. There are also international legal obligations for its protection through the Convention on Migratory Species (CMS) and EUROBATS Agreement. It is included in Appendix II ; of the Council of Europe’s Convention on the Conservation of European Wildlife and Natural Habitats (Bern Convention). Also included in Annex IV of EU Habitats and Species Directive. Important habitats partially protected by Natura 2000 sites and national networks of protected areas. ;Real population trends outside the EU are barely known due to the lack of research.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	brandtii	Eversmann	1845	1	Bull. Soc. Nat. Moscow	18(1): 505	Brandt's Myotis	 aureus Koch, 1865; coluotus Kostron, 1943.	Russia, Orenburgsk. Obl., S. Ural, Bolshoi-Ik River, Spasskoie. Foothills of the Ural Mountains.	Britain south to Italy, Greece, and Bulgaria; east to Kazakhstan and Mongolia, E Siberia including Sakhalin Isls, Kamchatka Peninsula and Kurile Isls; Ussuri region (Russia); Korea.	Not listed.	Least Concern	Listed as a subspecies of mystacinus by Ellerman and Morrison-Scott (1951), but see Strelkov and Buntova (1982) and Benda andTsytsulina (2000). Does not include fujiensis , see Yoshiyuki (1989), Benda and Tsytsulina (2000), and Horácek et al. (2000). Does notinclude gracilis or sibiricus , see Kruskop et al. 2012 who suggested that sibiricus may be the correct name for gracilis. Horácek et al. (2000) provisionally treated gracilis as distinct from brandtii and fujiensis pending further study. See also Yoon and Son (1989). Sometimes misspelled brandti , but brandtii is the original spelling.	Myotis brandtii	1005380	23	Brandt's Myotis	Brandt's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	brandtii	Eversmann	1845	1						Orenburgsk, Russia.			brandtii (Eversmann, 1845)|aureus (L. Koch, 1865)|coluotus Kostron, 1943	previously included M. sibiricus (including gracilis)	Ohdachi, S. D. I, Ishibashi, Y., Iwasa, M. A., & Saitoh, Takashi (2009). The Wild Mammals of Japan, Shoukadoh, Kyoto.				Norway|Sweden|Finland|United Kingdom|France|Netherlands|Belgium|Luxembourg|Germany|Denmark|Switzerland|Liechtenstein?|Italy|Austria|Czech Republic|Poland|Slovakia|Hungary|Slovenia|Croatia|Bosnia & Herzegovina?|Serbia|Kosovo|Albania|North Macedonia?|Albania|Greece|Bulgaria|Romania|Moldova|Ukraine|Belarus|Lithuania|Latvia|Estonia|Russia|Georgia|Turkey|Kazakhstan	Asia|Europe	Palearctic	LC	0	0	0	Myotis_brandtii	0	sciname match	Myotis_brandtii	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_brandtii	1005380	23	Brandt's Myotis	Brandt's Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	brandtii	Eversmann	1	Vespertilio Brandtii	Eversmann, E.F. 1845. Vespertiliones, in promontoriis Uralensibus tractibusque confinibus observati. Bulletin de la Société Impériale des Naturalistes de Moscou 18(1):489-516.	https://www.biodiversitylibrary.org/page/44165381	ZIN S. 41687	lectotype		Orenburgsk, Russia.			previously included M. sibiricus (including gracilis)	Ohdachi, S. D. I, Ishibashi, Y., Iwasa, M. A., & Saitoh, Takashi (2009). The Wild Mammals of Japan, Shoukadoh, Kyoto.				Norway|Sweden|Finland|United Kingdom|France|Netherlands|Belgium|Luxembourg|Germany|Denmark|Switzerland|Liechtenstein?|Italy|Austria|Czech Republic|Poland|Slovakia|Hungary|Slovenia|Croatia|Bosnia and Herzegovina?|Serbia|Kosovo|North Macedonia?|Albania|Greece|Bulgaria|Romania|Moldova|Ukraine|Belarus|Lithuania|Latvia|Estonia|Russia|Georgia|Turkey|Kazakhstan	Asia|Europe	Palearctic	LC	0	0	0	Myotis_brandtii	0	sciname match	Myotis_brandtii	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	brandtii	Eversmann	1845	1	Bull. Soc. Nat. Moscow	18(1): 505	Brandt's Myotis	aureus Koch, 1865; coluotus Kostron, 1943.	Russia, Orenburgsk. Obl., S. Ural, Bolshoi-Ik River, Spasskoie. Foothills of the Ural Mountains.	Britain south to Italy, Greece, and Bulgaria; east to Kazakhstan and Mongolia, E Siberia including Sakhalin Isls, Kamchatka Peninsula and Kurile Isls; Ussuri region (Russia); Korea.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/85566997/195857637/' target='_blank'>Least Concern</a>	Listed as a subspecies of mystacinus by Ellerman and Morrison-Scott (1951), but see Strelkov and Buntova (1982) and Benda andTsytsulina (2000). Does not include fujiensis, see Yoshiyuki (1989), Benda and Tsytsulina (2000), and Horácek et al. (2000). Does notinclude gracilis or sibiricus, see Kruskop et al. 2012 who suggested that sibiricus may be the correct name for gracilis. Horácek et al. (2000) provisionally treated gracilis as distinct from brandtii and fujiensispending further study. See also Yoon and Son (1989). Sometimes misspelled brandti, but brandtii is the original spelling.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis brandtii; Myotis brandtii; Myotis brandtii; Myotis brandtii; Myotis brandtii; Myotis brandtii; brandtii; gracilis; aureus; coluotus; sibiricus; aureus; coluotus; brandtii; aureus; coluotus; Murin de Brandt; Gro Re Bartfledermaus; Ratonero de Brandt; Brandt's Bat; Brandt's Myotis; Brandt's Bat; Brandt's Myotis; Brandt's Myotis; M. brandtii