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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L837	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Myotis montivagus [synonym of]	Myotis montivagus borneoensis	Myotis montivagus borneoensis	Myotis borneoensis	Myotis borneoensis	Myotis borneoensis	Myotis borneoensis	Myotis borneoensis	Myotis borneoensis	Myotis borneoensis	Myotis borneoensis	Myotis borneoensis		[HMW] Myotis montivagus borneoensis Hill & Francis, 1984 , Sepilok, Sabah , Borneo. Subgenus Myotis ; montivagus species group. Based on morphological and morphometric differences, M. borneoensis was separated from M. montivagus in 2013. Monotypic.; [batnames2022] Distinct from montivagus; see Gorfol et al. (2013).; [MDD2022] split from M. montivagus; [IUCN] <span lang="EN-US">This taxon was generally regarded until recently as a subspecies of M. montivagus but was elevated to species rank based on morphology and morphometrics by Görföl et al . (2013)<a name="_GoBack"></a>.; [batnames2023] Distinct from montivagus; see Gorfol et al. (2013).; [MDD2023] split from M. montivagus; [MDD2025_2.0] split from M. montivagus; [batnames2025_1.7] Distinct from montivagus; see Gorfol et al. (2013).; [MDD2025_2.2] split from M. montivagus														borneoensis	<span lang="EN-US">This taxon was generally regarded until recently as a subspecies of M. montivagus but was elevated to species rank based on morphology and morphometrics by Görföl et al . (2013)<a name="_GoBack"></a>.			borneoensis	borneoensis			borneoensis J. Edwards Hill & C. M. Francis, 1984						N/A																																								NA																											4C3D87E8FF336A8CFA5D9443163CB220	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	962	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF336A8CFA5D9443163CB220.xml	Myotis borneoensis	Vespertilionidae	Myotis	borneoensis	Hill & Francis	1984	Murin de Bornéo @fr | Borneo-Bartfledermaus @de | Ratonero bigotudo de Borneo @es	Myotis montivagus borneoensis Hill & Francis, 1984 , Sepilok, Sabah , Borneo. Subgenus Myotis ; montivagus species group. Based on morphological and morphometric differences, M. borneoensis was separated from M. montivagus in 2013. Monotypic.	N Borneo and scattered localities in Sabah , Kalimantan, and Sarawak . Recently, a medium-large brown Myotis (tentatively assigned to this species) was collected in Bukit Barisan Selatan National Park, S Sumatra, but this needs to be confirmed by genetic analyses.	Tail 41-47 mm ,ear 15-1-16- 1 mm , hindfoot 8-9 mm , forearm 42-1— 44- 8 mm ; weight 9-135 g . Furis rather long and dense. Upperparts have dark blackish brown hair bases, with dark brown tips. Underparts have similar blackish brown hair bases and buffy brown tips. Ears long and narrow, with long tragusthat reaches ¢.50% of ear length and bendsslightly forward. Wings are attached at base offirst toe. Skull is broad and not especially elongated, braincase is broad, interorbital region is wide, and zygomata are widely flared. Depression between rostrum and braincase is rather shallow. Sagittal and lambdoid crests are well developed; anteorbital bridge is wide. Basal dimension of P? is one-third that of P*. P? very small and completely displaced from tooth row so that P? and P* are usually in tight contact. P, is intruded from tooth row and not visible laterally; P, and P, are not in contact. Condylo-canine lengths are 15-4-16- 1 mm ; maxillary wor rows are 6-7-7- 1 mm .	Alongside streamsin tall forests and plantations.	No information.	No information.	The Bornean Whiskered Myotis is nocturnal.	In a survey of small mammals in Bukit Sarang, Bintulu division, Sarawak , the Bornean Whiskered Myotis (under montivagus ) was captured syntopically with other four Myotis species—Horsfield’s Myotis ( M. cf. ater ), Gomantong Myotis ( M. gomantongensis ), and M. sp. undescribed. Bukit Sarang is a 700-m long and 80-m high limestone outcrop, surrounded by freshwater swamp forest inundated to a depth of 0-5- 1 m several times per year. In East Kalimantan, it collected in a cave in karst area.	Classified as Data Deficient on The IUCN Red List. The Bornean Whiskered Myotis is known from only a few specimens, and little is known aboutits distribution, abundance, ecology, or conservation threats.	Corbet & Hill (1992) | Gorfol & Csorba (2017a) | Gorfél et al. (2013) | Hill & Francis (1984) | Huang, J.C.C. et al. (2014) | Koopman (1994) | Payne et al. (1985) | Phillipps & Phillipps (2016) | Simmons (2005) | Suyanto & Struebig (2007) | Wilson et al. (2006) | Yasuma et al. (2003)	https://zenodo.org/record/6398896/files/figure.png	453. Bornean Whiskered Myotis Myotis borneoensis French: Murin de Bornéo / German: Borneo-Bartfledermaus / Spanish: Ratonero bigotudo de Borneo Taxonomy. Myotis montivagus borneoensis Hill & Francis, 1984 , Sepilok, Sabah , Borneo. Subgenus Myotis ; montivagus species group. Based on morphological and morphometric differences, M. borneoensis was separated from M. montivagus in 2013. Monotypic. Distribution. N Borneo and scattered localities in Sabah , Kalimantan, and Sarawak . Recently, a medium-large brown Myotis (tentatively assigned to this species) was collected in Bukit Barisan Selatan National Park, S Sumatra, but this needs to be confirmed by genetic analyses. Descriptive notes. Tail 41-47 mm ,ear 15-1-16- 1 mm , hindfoot 8-9 mm , forearm 42-1— 44- 8 mm ; weight 9-135 g . Furis rather long and dense. Upperparts have dark blackish brown hair bases, with dark brown tips. Underparts have similar blackish brown hair bases and buffy brown tips. Ears long and narrow, with long tragusthat reaches ¢.50% of ear length and bendsslightly forward. Wings are attached at base offirst toe. Skull is broad and not especially elongated, braincase is broad, interorbital region is wide, and zygomata are widely flared. Depression between rostrum and braincase is rather shallow. Sagittal and lambdoid crests are well developed; anteorbital bridge is wide. Basal dimension of P? is one-third that of P*. P? very small and completely displaced from tooth row so that P? and P* are usually in tight contact. P, is intruded from tooth row and not visible laterally; P, and P, are not in contact. Condylo-canine lengths are 15-4-16- 1 mm ; maxillary wor rows are 6-7-7- 1 mm . Habitat. Alongside streamsin tall forests and plantations. Food and Feeding. No information. Breeding. No information. Activity patterns. The Bornean Whiskered Myotis is nocturnal. Movements, Home range and Social organization. In a survey of small mammals in Bukit Sarang, Bintulu division, Sarawak , the Bornean Whiskered Myotis (under montivagus ) was captured syntopically with other four Myotis species—Horsfield’s Myotis ( M. cf. ater ), Gomantong Myotis ( M. gomantongensis ), and M. sp. undescribed. Bukit Sarang is a 700-m long and 80-m high limestone outcrop, surrounded by freshwater swamp forest inundated to a depth of 0-5- 1 m several times per year. In East Kalimantan, it collected in a cave in karst area. Status and Conservation. Classified as Data Deficient on The IUCN Red List. The Bornean Whiskered Myotis is known from only a few specimens, and little is known aboutits distribution, abundance, ecology, or conservation threats. Bibliography. Corbet & Hill (1992), Gorfol & Csorba (2017a), Gorfél et al. (2013), Hill & Francis (1984), Huang, J.C.C. et al. (2014), Koopman (1994), Payne et al. (1985), Phillipps & Phillipps (2016), Simmons (2005), Suyanto & Struebig (2007), Wilson et al. (2006), Yasuma et al. (2003).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis borneoensis	Myotis	Unassigned-Myotis	borneoensis	Hill & Francis	1984	0	Bull. Br. Mus. Nat. Hist. Zool.	47(5): 309	Bornean Whiskered Myotis	None.		Borneo	Not listed.	Data Deficient	Distinct from montivagus; see Gorfol et al. (2013).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis borneoensis	23	Bornean Whiskered Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Myotis	borneoensis	J. Edwards Hill & Francis	1984	0	Myotis_montivagus_borneoensis	Hill, J. E. & Francis, C. M. (1984) New bats (Mammalia: Chiroptera) and new records of bats from Borneo and Malaya. Bulletin of the British Museum (Natural History), Zoology Series, 47(5), 309.	https://www.biodiversitylibrary.org/item/19472#page/341/mode/1up	BM 1983.349		Sepilok, Sabah, Borneo.			borneoensis J. Edwards Hill & Francis, 1984	split from M. montivagus	Görföl, T., Estók, P., & Csorba, G. (2013). The subspecies of Myotis montivagus—taxonomic revision and species limits (Mammalia: Chiroptera: Vespertilionidae). Acta Zoologica Academiae Scientiarum Hungaricae, 59(1), 41-59.	Malaysia|Indonesia	Asia	Indomalaya	DD	0	0	0	Myotis_borneoensis	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	90000000	Myotis borneoensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	borneoensis	Hill and Francis, 1984	<span lang="EN-US">This taxon was generally regarded until recently as a subspecies of M. montivagus but was elevated to species rank based on morphology and morphometrics by Görföl et al . (2013)<a name="_GoBack"></a>.	90000000	Myotis borneoensis	Data Deficient		2017	2016-07-02 00:00:00 UTC	3.1	English	<span lang="EN-US">This species is listed as Data Deficient as it is known only from a few specimens and we have very few information on its distribution, abundance, ecology or threats.	<span lang="EN-US">Payne et al . (1985) and Wilson et al . (2006) recorded the species alongside forest streams, whereas specimens collected in Kalimantan emerged from a cave in a karst area.	The species may be a cave-dweller (Görföl et al . 2013), hence the disturbance of caves is a threat to the species. It was captured in tall forest; logging of forests may be also a threat.	<p><span lang="EN-US">The species is known from only a handful of specimens from Borneo.</p>	Unknown	<span lang="EN-US">Myotis borneoensis <span lang="EN-US"> is confined to Borneo. It occurs in Sabah, Malaysia (Hill and Francis 1984, Payne et al . 1985, Yasuma and Andau 2000 in Francis et al . 2008), Sarawak (Wilson et al . 2006), and also in Kalimantan, Indonesia (Suyanto and Struebig 2007).</span>		Terrestrial	No conservation actions are known to be in place for this species.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	borneoensis	Hill & Francis	1984	0	Bull. Br. Mus. Nat. Hist. Zool.	47(5): 309	Bornean Whiskered Myotis	None.		Borneo	Not listed.	Data Deficient	Distinct from montivagus; see Gorfol et al. (2013).	Myotis borneoensis	1005379	23	Bornean Whiskered Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Myotis	borneoensis	J. Edwards Hill & Francis	1984	0	Myotis_montivagus_borneoensis	Hill, J. E. & Francis, C. M. (1984) New bats (Mammalia: Chiroptera) and new records of bats from Borneo and Malaya. Bulletin of the British Museum (Natural History), Zoology Series, 47(5), 309.	https://www.biodiversitylibrary.org/item/19472#page/341/mode/1up	BM 1983.349		Sepilok, Sabah, Borneo.			borneoensis J. Edwards Hill & Francis, 1984	split from M. montivagus	Görföl, T., Estók, P., & Csorba, G. (2013). The subspecies of Myotis montivagus—taxonomic revision and species limits (Mammalia: Chiroptera: Vespertilionidae). Acta Zoologica Academiae Scientiarum Hungaricae, 59(1), 41-59.				Malaysia|Indonesia	Asia	Indomalaya	DD	0	0	0	Myotis_borneoensis	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_borneoensis	1005379	23	Bornean Whiskered Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Myotis	borneoensis	J. Edwards Hill & C. M. Francis	0	Myotis montivagus borneoensis	Hill, J.E. and Francis, C.M. 1984. New bats (Mammalia: Chiroptera) and new records of bats from Borneo and Malaya. Bulletin of the British Museum (Natural History) (Zoology)47(5):305-329.	https://www.biodiversitylibrary.org/page/2281955	BMNH:Mamm:1983.349	holotype	https://data.nhm.ac.uk/object/1df17d8c-02f5-467f-9fd3-72345163ea04 | https://data.nhm.ac.uk/object/abbf446e-01aa-4bd4-af77-26910452860e	Sepilok, Sabah, Borneo.	5.866667	117.9333	split from M. montivagus	Görföl, T., Estók, P., & Csorba, G. (2013). The subspecies of Myotis montivagus—taxonomic revision and species limits (Mammalia: Chiroptera: Vespertilionidae). Acta Zoologica Academiae Scientiarum Hungaricae, 59(1), 41-59.				Malaysia|Indonesia	Asia	Indomalaya	DD	0	0	0	Myotis_borneoensis	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Myotis	borneoensis	Hill & Francis	1984	0	Bull. Br. Mus. Nat. Hist. Zool.	47(5): 309	Bornean Whiskered Myotis	None.		Borneo	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/85568289/85568292/' target='_blank'>Data Deficient</a>	Distinct from montivagus; see Gorfol et al. (2013).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis borneoensis; Myotis borneoensis; Myotis borneoensis; Myotis borneoensis; Myotis borneoensis; borneoensis; Murin de Bornéo; Borneo-Bartfledermaus; Ratonero bigotudo de Borneo; Bornean Whiskered Myotis; Bornean Whiskered Myotis; M. borneoensis