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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L823	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis	Myotis atacamensis		[MSW2] Subgenus Selysius. Listed as a subspecies of chiloensis by Cabrera (1958:99). Includes nicholsoni; see LaVal (1973«:18-20).; [MSW3] Listed as a subspecies of chiloensis by Cabrera (1958). Includes nicholsoni; see LaVal (1973a).; [HMW] Vespertilio atacamensis Lataste, 1892 , San Pedro de “ Atacama ,” Antofagasta , Chile . Subgenus Pizonyx; albescens species group. Myotis atacamensis was considered a subspecies of M. chiloensis , but studies based on morphology and molecular data indicate clear distinction between the two. Monotypic.; [batnames2022] Listed as a subspecies of chiloensis by Cabrera (1958). Includes nicholsoni ; see LaVal (1973 a ). Neotype designated by Novaes et al. (2022), who also presented a new diagnosis.; [IUCN] Listed as a subspecies of Myotis ;chiloensis . Includes M. ;nicholsoni (LaVal, 1973) (Simmons 2005).; [batnames2023] Listed as a subspecies of chiloensis by Cabrera (1958). Includes nicholsoni ; see LaVal (1973 a ). Neotype designated by Novaes et al. (2022), who also presented a new diagnosis.; [MDD2023] recently re-described with the designation of a neotype; [MDD2025_2.0] recently re-described with the designation of a neotype; [batnames2025_1.7] Listed as a subspecies of chiloensis by Cabrera (1958). Includes nicholsoni; see LaVal (1973a). Neotype designated by Novaes et al. (2022), who also presented a new diagnosis.; [MDD2025_2.2] recently re-described with the designation of a neotype				nicholsoni		nicholsoni.			nicholsoni			atacamensis 	atacamensis - nicholsoni	atacamensis, nicholsoni	Listed as a subspecies of Myotis ;chiloensis . Includes M. ;nicholsoni (LaVal, 1973) (Simmons 2005).	atacamensis 	atacamensis - nicholsoni	atacamensis, nicholsoni 	atacamensis, nicholsoni 	atacamensis 	atacamensis - nicholsoni	atacamensis (Lataste, 1892)|nicholsoni Sanborn, 1941		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		S Peru, N Chile	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis atacamensis	Chile, Antofogasta, San Pedro de Atacama.	Lataste	1892	Actes Soc. Sei. Chile, 1:80.	Distribution: Confined to the Pacific coastal region of southern Peru and northern Chile.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		S Peru, N Chile	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Lataste	1892	Actes Soc. Sci. Chile, 1:80.	Subgenus Selysius. Listed as a subspecies of chiloensis by Cabrera (1958:99). Includes nicholsoni; see LaVal (1973«:18-20).	S Peru, N Chile.	Chile, Antofogasta, San Pedro de Atacama.		LATASTE	1891	Size relatively small (forearm length, 30-34 mm; condylobasal length, 11-12 mm). Occiput not raised above braincase and rostrum relatively slender. Sagittal crest absent.	Distribution: Confined to the Pacific coastal region of southern Peru and northern Chile.	No subspecies.		105	species	M. atacamensis	LATASTE	1891	Selysius	subgenus	Myotis atacamensis				Size relatively small (forearm length, 30-34 mm; condylobasal length, 11-12 mm). Oc- ciput not raised above braincase and rostrum relatively slender. Sagittal crest absent.	No subspecies.		46. M. atacamensis (LATASTE 1891) [nigricans group].	46	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis atacamensis	Myotis		atacamensis	Lataste	y	1892		Actes Soc. Sci. Chile	1		80		Atacaman Myotis	Chile, Antofogasta, San Pedro de Atacama.	S Peru, N Chile.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.	nicholsoni Sanborn, 1941.	Listed as a subspecies of chiloensis by Cabrera (1958). Includes nicholsoni; see LaVal (1973a).	4C3D87E8FF4E6AF2FA899FF21C6ABE05	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	943	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF4E6AF2FA899FF21C6ABE05.xml	Myotis atacamensis	Vespertilionidae	Myotis	atacamensis		1892	Murin d Atacama @fr | Atacama-Mausohr @de | Ratonero de Atacama @es | Atacaman Myotis @en	Vespertilio atacamensis Lataste, 1892 , San Pedro de “ Atacama ,” Antofagasta , Chile . Subgenus Pizonyx; albescens species group. Myotis atacamensis was considered a subspecies of M. chiloensis , but studies based on morphology and molecular data indicate clear distinction between the two. Monotypic.	From CW Peru to CWChile.	Head-body c.41 45 mm , tail 31-40 mm , ear 13-14 mm , hindfoot 5-8 mm , forearm 30-8-34:- 1 mm : weight 3-4 g . Fur is long and silky (dorsal fur 7-9 mm ; ventral fur 6-7 mm ). Ventral and dorsal hairs are strongly bicolored, ventral hairs are black on bases and whitish on tips, and dorsal hairs are black on bases and light ocherous buff on tips. Teeth are small. Ears are medium-sized, extending forward halfway from eye to nostril. Membranes are mummy brown; plagiopatagiumis attached to foot at toes by a broad band of membrane; and fringe ofhairs along trailing edge of uropatagium is present. Skull is small (greatest length ofskull 12:6-13- 8 mm ); occipital regionis rounded; forehead slopes steeply, with inflated braincase; rostrum is narrow and short; sagittal and lambdoidal crests are generally absent or, when present, are very low. P? is aligned in tooth row and visible in labial view. Bacula are small and slight in width from proximalto distal end, and average measurements are: 0-50 mm long, 0-20 mm deep, and 0-27 mm wide. Wing morphology and echolocation are typical of aerial insectivore that uses cluttered spaces.	Arid and semiarid scrublands and thorn and sclerophyll forests from sea level up to elevations of ¢. 3475 m .	The Atacama Myotis feeds exclusively on insects captured in flight in open habitats and over watercourses.	Atacama Myotis seems to reproduce in September-December, with females giving birth to one young per pregnancy.	In semiarid highlands, foraging activity of the Atacama Myotis begins one hour before dusk and lasts for c.3 hours. Tree cavities, crevices in rocky outcrops, caves, abandoned mines, and other human buildings (e.g. house roof,attic and drainage pipeline) are used as roosts. Mean call components are start frequency of 67 kHz, end frequency of 47-7 kHz, peak frequency of 52 kHz, duration of 3-3 milliseconds, and interval between pulses of 134-9 milliseconds.	Colonies of Atacama Myotis can have more than 30 individuals.	Classified as Endangered on The IUCNRed List. Although the Atacama Myotisis still reasonably widespread,it is very dependent on specific habitats that have become severely fragmented. Surveys along coastal deserts of Peru and Chile found only a few populations that were disjunct from each other. Area of occupancyis less than 200 km *. With predicted loss of habitat, population of the Atacama Myotisis expected to decline more than 30% in the next three generations.	Galaz & Yanez (2006) | Galaz et al. (2009) | Iriarte (2008) | LaVal (1973b) | Ossa et al. (2017) | Rodriguez-San Pedro, Allendes, Castillo et al. (2014) | Rodriguez-San Pedro, Allendes & Ossa (2016) | Stadelman et al. (2007) | Vargas-Rodriguez et al. (2016)	https://zenodo.org/record/6398797/files/figure.png	411. Atacama Myotis Myotis atacamensis French: Murin dAtacama / German: Atacama-Mausohr / Spanish: Ratonero de Atacama Other common names: Atacaman Myotis Taxonomy. Vespertilio atacamensis Lataste, 1892 , San Pedro de “ Atacama ,” Antofagasta , Chile . Subgenus Pizonyx; albescens species group. Myotis atacamensis was considered a subspecies of M. chiloensis , but studies based on morphology and molecular data indicate clear distinction between the two. Monotypic. Distribution. From CW Peru to CWChile. Descriptive notes. Head-body c.41 45 mm , tail 31-40 mm , ear 13-14 mm , hindfoot 5-8 mm , forearm 30-8-34:- 1 mm : weight 3-4 g . Fur is long and silky (dorsal fur 7-9 mm ; ventral fur 6-7 mm ). Ventral and dorsal hairs are strongly bicolored, ventral hairs are black on bases and whitish on tips, and dorsal hairs are black on bases and light ocherous buff on tips. Teeth are small. Ears are medium-sized, extending forward halfway from eye to nostril. Membranes are mummy brown; plagiopatagiumis attached to foot at toes by a broad band of membrane; and fringe ofhairs along trailing edge of uropatagium is present. Skull is small (greatest length ofskull 12:6-13- 8 mm ); occipital regionis rounded; forehead slopes steeply, with inflated braincase; rostrum is narrow and short; sagittal and lambdoidal crests are generally absent or, when present, are very low. P? is aligned in tooth row and visible in labial view. Bacula are small and slight in width from proximalto distal end, and average measurements are: 0-50 mm long, 0-20 mm deep, and 0-27 mm wide. Wing morphology and echolocation are typical of aerial insectivore that uses cluttered spaces. Habitat. Arid and semiarid scrublands and thorn and sclerophyll forests from sea level up to elevations of ¢. 3475 m . Food and Feeding. The Atacama Myotis feeds exclusively on insects captured in flight in open habitats and over watercourses. Breeding. Atacama Myotis seems to reproduce in September-December, with females giving birth to one young per pregnancy. Activity patterns. In semiarid highlands, foraging activity of the Atacama Myotis begins one hour before dusk and lasts for c.3 hours. Tree cavities, crevices in rocky outcrops, caves, abandoned mines, and other human buildings (e.g. house roof,attic and drainage pipeline) are used as roosts. Mean call components are start frequency of 67 kHz, end frequency of 47-7 kHz, peak frequency of 52 kHz, duration of 3-3 milliseconds, and interval between pulses of 134-9 milliseconds. Movements, Home range and Social organization. Colonies of Atacama Myotis can have more than 30 individuals. Status and Conservation. Classified as Endangered on The IUCNRed List. Although the Atacama Myotisis still reasonably widespread,it is very dependent on specific habitats that have become severely fragmented. Surveys along coastal deserts of Peru and Chile found only a few populations that were disjunct from each other. Area of occupancyis less than 200 km *. With predicted loss of habitat, population of the Atacama Myotisis expected to decline more than 30% in the next three generations. Bibliography. Galaz & Yanez (2006), Galaz et al. (2009), Iriarte (2008), LaVal (1973b), Ossa et al. (2017), Rodriguez-San Pedro, Allendes, Castillo et al. (2014), Rodriguez-San Pedro, Allendes & Ossa (2016), Stadelman et al. (2007), Vargas-Rodriguez et al. (2016).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis atacamensis	Myotis	Unassigned-Myotis	atacamensis	Lataste	1892	1	Actes Soc. Sci. Chile	0.0972	Atacaman Myotis	 nicholsoni Sanborn, 1941.	Neotype: Chile, Tarapacá, near Minimini. Original type locality: Chile, Antofogasta, San Pedro de Atacama	S Peru, N Chile.	Not listed.	Endangered	Listed as a subspecies of chiloensis by Cabrera (1958). Includes nicholsoni ; see LaVal (1973 a ). Neotype designated by Novaes et al. (2022), who also presented a new diagnosis.	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis atacamensis	23	Atacama Myotis	Atacaman Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	atacamensis	Lataste	1892	1						San Pedro de "Atacama," Antofagasta, Chile.			atacamensis (Lataste, 1892)|nicholsoni Sanborn, 1941	NA	NA	Peru|Chile	South America	Neotropic	EN	0	0	0	Myotis_atacamensis	0	sciname match	Myotis_atacamensis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14143	Myotis atacamensis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	atacamensis	(Lataste, 1892)	Listed as a subspecies of Myotis ;chiloensis . Includes M. ;nicholsoni (LaVal, 1973) (Simmons 2005).	20000000	Myotis atacamensis	Endangered	B2ab(ii,iii)	2016	2014-12-12 00:00:00 UTC	3.1	English	This bat is listed as Endangered because, although the species is still reasonably widely distributed, it is very dependent on its specific habitat and it has become severely fragmented. Surveys along the coastal deserts of Peru and Chile have not found connecting populations, and the few ones are well apart from each other. Also, i<span lang="EN-AU">ts area of occupancy (AOO) is less than 200 km<sup>2</sup> and the species qualifies as threatened under criterion B2ab<span lang="EN-US">(ii,iii)<span lang="EN-AU">. There is a predicted changes in land use, population is expected to decline more than 30% in the next three generations (18 years, Pacifici et al. 2013) based on the decline of habitat quality, that qualify the species Vulnerable under criterion A3c.</span>	<span lang="ES-CL">The species occupies deserts and thorny arid scrubs of Peru and Chile form the core of its distribution. The area of distribution includes three major areas, wide apart from each other; each area includes several local and smaller populations. Diurnal roosts include rock crevices, small caves, and hollow trees (Mann 1978, Iriarte 2008), but might use ceilings in large edifications (churches, old houses) or small drainage tunnels<span lang="ES-CL"><span lang="ES-CL">. ;Foraging activity begins one hour before dusk and lasts for about three hours (Galaz ;et al.  2009). ;This is the only Neotropical species of Myotis that hibernates.</span>	Threats to this species include ;<span lang="EN-US">land use change, mining, agriculture, urban development, and eolic energy production (wind farms). ;Along its geographic distribution, the agricultural expansion is changing the landscape, including species composition, plus the increase of eolic farms in northern Chile which are recognized as a direct threat to insectivorous bats.<span lang="ES-CL"></span>	There is no information on population size across its geographic range. There are a few occasional records of small colonies in Chile, like the one described by Galaz and Yañez (2006) (with 20 individuals near ;<span lang="ES-CL">El Refresco, Reserva Nacional ; Pampa del Tamarugal), and the one reported by Rodríguez-San Pedro et al.  (2014) (with almost 30 individuals at Reserva Nacional Las Chinchillas). From 1923 to 2014, the average number of annual records is less than one, and some years have passed without any record at all (1925-1938, 1953-1967 and 1995-2012). Across the years there is evidence of a cycle of high fluctuations, going from a maximum peak to a minimum (unpubl. data). Given that the number of records is somehow related to the abundance of individuals, these variations could be related to the population dynamics of the species, but this remains to be verified.	Unknown	Atacama Myotis is known from southern Peru and northern Chile (Simmons 2005). However historical records indicate the presence of the species in northern Peru, and there are records in the central coast, and a recent southern record from Choapa, Coquimbo, Chile (Rodríguez San Pedro et al.   2014).		Terrestrial	Regional efforts in Chile are looking for protection of the habitats where the species is present. These include the designation of protected areas and their inclusion in the National System of Protected Areas. The species occurs at the ;<span lang="ES-CL">Reserva Nacional Las Chinchillas (Coquimbo region) and ;<span lang="ES-CL">Reserva Nacional ; Pampa del Tamarugal, both in Chile. Further studies are necessary to assess the risk of populations nearby the projects for Eolic farms in northern Chile.</span>	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	atacamensis	Lataste	1892	1	Actes Soc. Sci. Chile	0.097222	Atacaman Myotis	 nicholsoni Sanborn, 1941.	Neotype: Chile, Tarapacá, near Minimini. Original type locality: Chile, Antofogasta, San Pedro de Atacama	S Peru, N Chile.	Not listed.	Endangered	Listed as a subspecies of chiloensis by Cabrera (1958). Includes nicholsoni ; see LaVal (1973 a ). Neotype designated by Novaes et al. (2022), who also presented a new diagnosis.	Myotis atacamensis	1005367	23	Atacama Myotis	Atacaman Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	atacamensis	Lataste	1892	1		Lataste F (1892) Etudes sur la faune Chilienne. II. Note sur les chauve-souris. Actes de la Société Scientifique du Chile 1:70−91.		USNM 391786 [neotype]		San Pedro de "Atacama," Antofagasta, Chile. Restricted by R. Novaes and colleagues in 2022 to "Near Minimini, Tarapacá, Chile (19°10’S, 69°41’W; elevation 1,800 m), inside Atacama desert."	-19.1667	-69.683333	atacamensis (Lataste, 1892)|nicholsoni Sanborn, 1941	recently re-described with the designation of a neotype	Novaes, R. L. M., LaVal, R. K., Wilson, D. E., & Moratelli, R. (2022). Redescription of Myotis atacamensis (Chiroptera: Vespertilionidae) with neotype designation. Zoologia (Curitiba), 39, e21026. https://doi.org/10.1590/s1984-4689.v39.e21026				Peru|Chile	South America	Neotropic	EN	0	0	0	Myotis_atacamensis	0	sciname match	Myotis_atacamensis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_atacamensis	1005367	23	Atacama Myotis	Atacaman Myotis	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	atacamensis	Lataste	1	Vespertilio atacamensis	Lataste, F. 1892. Études sur la faune chilienne. II – Note sur les chauves-souris (Ordre des Chiroptères). Actes de la Société Scientifique de Chili 1:70-91.	https://babel.hathitrust.org/cgi/pt?id=coo.31924007304680&seq=164	USNM:MAMM:391786	neotype	http://n2t.net/ark:/65665/3dbc4f22e-2f77-44a7-bc07-de992ba799e1	San Pedro de "Atacama," Antofagasta, Chile. Restricted by R. Novaes and colleagues in 2022 to "Near Minimini, Tarapacá, Chile (19°10’S, 69°41’W; elevation 1,800 m), inside Atacama desert."	-19.1667	-69.6833	recently re-described with the designation of a neotype	Novaes, R. L. M., LaVal, R. K., Wilson, D. E., & Moratelli, R. (2022). Redescription of Myotis atacamensis (Chiroptera: Vespertilionidae) with neotype designation. Zoologia (Curitiba), 39, e21026. https://doi.org/10.1590/s1984-4689.v39.e21026				Peru|Chile	South America	Neotropic	EN	0	0	0	Myotis_atacamensis	0	sciname match	Myotis_atacamensis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	atacamensis	Lataste	1892	1	Actes Soc. Sci. Chile	0.097222	Atacaman Myotis	nicholsoni Sanborn, 1941.	Neotype: Chile, Tarapacá, near Minimini. Original type locality: Chile, Antofogasta, San Pedro de Atacama	S Peru, N Chile.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14143/22050638/' target='_blank'>Endangered</a>	Listed as a subspecies of chiloensis by Cabrera (1958). Includes nicholsoni; see LaVal (1973a). Neotype designated by Novaes et al. (2022), who also presented a new diagnosis.		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis atacamensis; Myotis atacamensis; Myotis atacamensis; Myotis atacamensis; Myotis atacamensis; Myotis atacamensis; nicholsoni; nicholsoni; atacamensis; nicholsoni; Murin d Atacama; Atacama-Mausohr; Ratonero de Atacama; Atacaman Myotis; Atacama Myotis; Atacaman Myotis; Atacaman Myotis; Atacaman Myotis; M. atacamensis