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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L813 application/vnd.openxmlformats-officedocument.spreadsheetml.sheet N/A N/A N/A N/A N/A Myotis alcathoe Myotis alcathoe Myotis alcathoe Myotis alcathoe Myotis alcathoe Myotis alcathoe Myotis alcathoe Myotis alcathoe Myotis alcathoe Myotis alcathoe [MSW3] See Ruedi et al. (2002).; [HMW] Myotis alcathoe von Helversen & Heller in von Helversen et al., 2001 , “netted over a small stream, the Fournikos Potamos, near the village of Kleistos ( 39°05’N , 21°49°E ), Novos Evritanias, Greece . Subgenus Myotis (62 species); alcathoe species group (2 species). Myotis alcathoe and M. hyrcanicus are sister to one another and appear to form a clade basal to the rest of the subgenus Myotis . Two subspecies recognized.; [batnames2022] See Ruedi et al. (2002) and Benda et al. (2016).; [IUCN] Formerly included in Myotis mystacinus (Kuhl, 1817); the species was differentiated (von Helversen et al. 2001) on the base of karyological, genetic and echolocation characters.; [batnames2023] See Ruedi et al. (2002) and Benda et al. (2016).; [batnames2025_1.7] See Ruedi et al. (2002) and Benda et al. (2016). alcathoevon, circassicus alcathoe, circassicus alcathoe, circassicus Formerly included in Myotis mystacinus (Kuhl, 1817); the species was differentiated (von Helversen et al. 2001) on the base of karyological, genetic and echolocation characters. alcathoe, circassicus alcathoe, circassicus alcathoe, circassicus alcathoe, circassicus alcathoe von Helversen & K.-G. Heller in von Helversen, K.-G. Heller, F. Mayer, Nemeth, Volleth, & GombkötÅ‘, 2001|circassicus Benda, Gazaryan, & Vallo, 2016 N/A _M. a. alcathoe_ Helversen & Heller, 2001; _M. a. circassicus_ Benda, ГазарÑн & Vallo, 2016 Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu). CHIROPTERA Vespertilionidae Myotinae Myotis alcathoe Myotis alcathoe von Helversen and Heller 2001 In von Helversen, Heller, Mayer, Nemeth, Volleth, and Gombkötö, Naturwissenschaften 88 217 Alcathoe Myotis Greece, Nomos Evritanias, near the village of Kleistos, over Fournikos Patomos stream, 39°05'N, 21°49'E. Greece, Hungary, France. Specimens from Bulgaria, Romania, and Ukraine previsously reported as ikonnikovi might represent alcathoe (von Helveren et al., 2001). IUCN 2003 – Not evaluated (new species); not considered in IUCN/SSC Action Plan (2001). See Ruedi et al. (2002). 4C3D87E8FF3B6A84FF4F9D621984B0D7 Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions 978-84-16728-19-0 hbmw_9_Vespertilionidae_716.pdf.imf hash://md5/b004ff90fffb6a44fffc96591e00bb32 954 zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF3B6A84FF4F9D621984B0D7.xml Myotis alcathoe Vespertilionidae Myotis alcathoe von Helversen & Heller 2001 Murin alcathoé @fr | Nymphenfledermaus @de | Ratonero bigotudo pequeno @es | Alcathoe Bat @en | Alcathoe Myotis @en | Alcathoe Whiskered Bat @en Myotis alcathoe von Helversen & Heller in von Helversen et al., 2001 , “netted over a small stream, the Fournikos Potamos, near the village of Kleistos ( 39°05’N , 21°49°E ), Novos Evritanias, Greece . Subgenus Myotis (62 species); alcathoe species group (2 species). Myotis alcathoe and M. hyrcanicus are sister to one another and appear to form a clade basal to the rest of the subgenus Myotis . Two subspecies recognized. M.a.alcathoevonHelversen&Heller,2001—patchilythroughalmostallEurope,fromNSpaintoGermany,includingBritain,andEtoSWUkraineandNWTurkey(EThrace);therearealsoecholocationcallsfromSSwedenthatcertainlyrepresentthisspeciesandtheremayalsoberecordsfromLatvia. M. a. circassicus Benda, Gazaryan & Vallo, 2016 — N slopes of Greater Caucasus in Russia and Abkhazia ( Georgia ); possibly also found in North Ossetia — Alania Republic ( Russia ), NE Turkey ( Artvin and Erzurum provinces), and Azerbaijan . Head-body 39-44 mm ,tail 36-37 mm , ear 11:4-14- 3 mm , hindfoot 5-1-5-8, forearm 29-7-34- 6 mm ; weight 3-5-5- 5 g . The Alcathoe Whiskered Myotis is one of the smallest European Myotis , with brownish and reddish upperparts and lighter brown underparts; hair is comparatively long ( 6-8 mm ) dark at base with light brown tips. fur can be entirely grayish or brownish in young bats. Wings are fairly dark, brownish or blackish, although lighter than in the Common Whiskered Myotis ( M. mystacinus ), and always attached to fifth toe. Ears are short and pale; short tragus has clear notch on edge. Nostrils are heart-shaped. Face is mainly furred, but pinkish bare skin is much paler than that of the Common Whiskered Myotis . Feet, legs and thumbs are remarkably short and small; calcar is slender and relatively long; tail membrane is less pigmented on the edge near the calcar. Penis is narrow from base to tip; baculum is short (c. 0-5 mm long) and broad. Skull is similar to that of the Common Whiskered Myotis and Brandt's Myotis ( M. brandtii ) but has higher braincase. P? and P° are tiny and pressed against C1 and P* P* has a clearly developed cusp; upper molars have protoconules. Chromosomal complement has 2n = 44 and FNa = 52 ( Greece ). The Alcathoe Whiskered Myotis is usually found near streams in mature moist forests, which have complex structure with deciduous trees such as Quercus ( Fabaceae ); Tilia ( Malvaceae ); and Alnus and Betula ( Betulaceae ); also riparian forest and montane Quercus , Carpinus ( Betulaceae ) and Fagus ( Fagaceae ) forests. It seem to favor sloping ground. Positively selects old mature forests due to the large numbers of decaying trees for roosting and maternity colonies. Commonly found in habitats also occupied by Bechstein’s Myotis ( M. bechsteinii ) and Brandt's Myotis . The Alcathoe Whiskered Myotis forages at forest edge byrivers, flying mostly in the canopy and hunting above water. Typically catches prey in flight but the presence of spiders and other non-flying animals suggests it may also capture prey from surface of leaves or ground. Usually feeds on small insects such as mosquitoes, small moths,flies, ants and beetles; also spiders. Locally it has been found to specialize on Nematocera, along with some caddisflies, moths, and spiders. Maternity colonies are normally in the canopy, and typically contain few adult females with their youngsters; males normally roost alone. Colonies tend to be small but rarely grow up to ¢.80 bats. Maternity roosts are very ephemeral, as the species switches roosts every few days. Females can give birth up to mid-July. The Alcathoe Whiskered Myotis usually roosts in fissures behind bark, and in holes in broken and rotten wood, 15-22 m aboveground. It moves from one roost to another frequently, even daily in many cases. It emerges 30-60 minutes after sunset and has a bimodal night activity pattern. Echolocation call is one of the highestfrequency types within Myotis , with relatively long (2:5—4 milliseconds) pulses. Pulses start at 120 kHz and end at c.46-32 kHz, with characteristic frequency of c.53 kHz. This is probably an adaptation to cluttered habitats and dense vegetation; the high frequency, combined with broad wings and high maneuverability, allow the species to fly in reduced spaces full of obstacles, and thus occupy a difficult niche typical of old forests. Echolocation calls are quite weak in terms of intensity, and are difficult to detect at distances over 10 m . The Alcathoe Whiskered Myotis usually moves to nearby rivers or streams at dusk, moving not more than 3-6 km each night to hunting areas; hunting zones are frequently changed. In winter, this species moves to caves or other small cavities in forest, although hibernation colonies have also been found in trees. Solitary animals can be found all year in small clusters. During the first half of August, some caves are used as swarming sites; these are often shared with other similar species, such as Bechstein’s Myotis and Brandt's Myotis . Classified as Data Deficient on The IUCN Red List, due to its recent description. The Alcathoe Whiskered Myotis was initially known only from Greece and Hungary , and it was thought to have a very restricted distribution; now known to be more widespread. It occurs at low densities and in scattered populations, with limited suitable habitat available. Main threats appear to include habitat loss and fragmentation, especially due to wood extraction; collisions with vehicles also said to be significant. Agirre-Mendi et al. (2004) | Ahlén (2010) | Alcalde (2009) | Bashta et al. (2011) | Benda & Karatas (2005) | Benda, Gazaryan & Vallo (2016) | Benda, Ruedi & Uhrin (2003) | Boston et al. (2011) | Danko et al. (2010) | De Pasquale & Galimberti (2014) | von Helversen et al. (2001) | Jan et al. (2010) | Luan et al. (2009) | Niermann et al. (2007) | Nyssen et al. (2015) | Pavlini¢ et al. (2012) | Rehék etal. (2008) | Sachanowicz et al. (2012) | Spitzenberger et al. (2008) | Volleth & Heller (2012) https://zenodo.org/record/6398853/files/figure.png 435. Alcathoe Whiskered Myotis Myotis alcathoe French: Murin alcathoé / German: Nymphenfledermaus / Spanish: Ratonero bigotudo pequeno Other common names: Alcathoe Bat , Alcathoe Myotis , Alcathoe Whiskered Bat Taxonomy. Myotis alcathoe von Helversen & Heller in von Helversen et al., 2001 , “netted over a small stream, the Fournikos Potamos, near the village of Kleistos ( 39°05’N , 21°49°E ), Novos Evritanias, Greece . Subgenus Myotis (62 species); alcathoe species group (2 species). Myotis alcathoe and M. hyrcanicus are sister to one another and appear to form a clade basal to the rest of the subgenus Myotis . Two subspecies recognized. Subspecies and Distribution. M.a.alcathoevonHelversen&Heller,2001—patchilythroughalmostallEurope,fromNSpaintoGermany,includingBritain,andEtoSWUkraineandNWTurkey(EThrace);therearealsoecholocationcallsfromSSwedenthatcertainlyrepresentthisspeciesandtheremayalsoberecordsfromLatvia. M. a. circassicus Benda, Gazaryan & Vallo, 2016 — N slopes of Greater Caucasus in Russia and Abkhazia ( Georgia ); possibly also found in North Ossetia — Alania Republic ( Russia ), NE Turkey ( Artvin and Erzurum provinces), and Azerbaijan . Descriptive notes. Head-body 39-44 mm ,tail 36-37 mm , ear 11:4-14- 3 mm , hindfoot 5-1-5-8, forearm 29-7-34- 6 mm ; weight 3-5-5- 5 g . The Alcathoe Whiskered Myotis is one of the smallest European Myotis , with brownish and reddish upperparts and lighter brown underparts; hair is comparatively long ( 6-8 mm ) dark at base with light brown tips. fur can be entirely grayish or brownish in young bats. Wings are fairly dark, brownish or blackish, although lighter than in the Common Whiskered Myotis ( M. mystacinus ), and always attached to fifth toe. Ears are short and pale; short tragus has clear notch on edge. Nostrils are heart-shaped. Face is mainly furred, but pinkish bare skin is much paler than that of the Common Whiskered Myotis . Feet, legs and thumbs are remarkably short and small; calcar is slender and relatively long; tail membrane is less pigmented on the edge near the calcar. Penis is narrow from base to tip; baculum is short (c. 0-5 mm long) and broad. Skull is similar to that of the Common Whiskered Myotis and Brandt's Myotis ( M. brandtii ) but has higher braincase. P? and P° are tiny and pressed against C1 and P* P* has a clearly developed cusp; upper molars have protoconules. Chromosomal complement has 2n = 44 and FNa = 52 ( Greece ). Habitat. The Alcathoe Whiskered Myotis is usually found near streams in mature moist forests, which have complex structure with deciduous trees such as Quercus ( Fabaceae ); Tilia ( Malvaceae ); and Alnus and Betula ( Betulaceae ); also riparian forest and montane Quercus , Carpinus ( Betulaceae ) and Fagus ( Fagaceae ) forests. It seem to favor sloping ground. Positively selects old mature forests due to the large numbers of decaying trees for roosting and maternity colonies. Commonly found in habitats also occupied by Bechstein’s Myotis ( M. bechsteinii ) and Brandt's Myotis . Food and Feeding. The Alcathoe Whiskered Myotis forages at forest edge byrivers, flying mostly in the canopy and hunting above water. Typically catches prey in flight but the presence of spiders and other non-flying animals suggests it may also capture prey from surface of leaves or ground. Usually feeds on small insects such as mosquitoes, small moths,flies, ants and beetles; also spiders. Locally it has been found to specialize on Nematocera, along with some caddisflies, moths, and spiders. Breeding. Maternity colonies are normally in the canopy, and typically contain few adult females with their youngsters; males normally roost alone. Colonies tend to be small but rarely grow up to ¢.80 bats. Maternity roosts are very ephemeral, as the species switches roosts every few days. Females can give birth up to mid-July. Activity patterns. The Alcathoe Whiskered Myotis usually roosts in fissures behind bark, and in holes in broken and rotten wood, 15-22 m aboveground. It moves from one roost to another frequently, even daily in many cases. It emerges 30-60 minutes after sunset and has a bimodal night activity pattern. Echolocation call is one of the highestfrequency types within Myotis , with relatively long (2:5—4 milliseconds) pulses. Pulses start at 120 kHz and end at c.46-32 kHz, with characteristic frequency of c.53 kHz. This is probably an adaptation to cluttered habitats and dense vegetation; the high frequency, combined with broad wings and high maneuverability, allow the species to fly in reduced spaces full of obstacles, and thus occupy a difficult niche typical of old forests. Echolocation calls are quite weak in terms of intensity, and are difficult to detect at distances over 10 m . Movements, Home range and Social organization. The Alcathoe Whiskered Myotis usually moves to nearby rivers or streams at dusk, moving not more than 3-6 km each night to hunting areas; hunting zones are frequently changed. In winter, this species moves to caves or other small cavities in forest, although hibernation colonies have also been found in trees. Solitary animals can be found all year in small clusters. During the first half of August, some caves are used as swarming sites; these are often shared with other similar species, such as Bechstein’s Myotis and Brandt's Myotis . Status and Conservation. Classified as Data Deficient on The IUCN Red List, due to its recent description. The Alcathoe Whiskered Myotis was initially known only from Greece and Hungary , and it was thought to have a very restricted distribution; now known to be more widespread. It occurs at low densities and in scattered populations, with limited suitable habitat available. Main threats appear to include habitat loss and fragmentation, especially due to wood extraction; collisions with vehicles also said to be significant. Bibliography. Agirre-Mendi et al. (2004), Ahlén (2010), Alcalde (2009), Bashta et al. (2011), Benda & Karatas (2005), Benda, Gazaryan & Vallo (2016), Benda, Ruedi & Uhrin (2003), Boston et al. (2011), Danko et al. (2010), De Pasquale & Galimberti (2014), von Helversen et al. (2001), Jan et al. (2010), Luan et al. (2009), Niermann et al. (2007), Nyssen et al. (2015), Pavlini¢ et al. (2012), Rehék etal. (2008), Sachanowicz et al. (2012), Spitzenberger et al. (2008), Volleth & Heller (2012). Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022. Vespertilionidae Myotis alcathoe Myotis Unassigned-Myotis alcathoe von Helversen and Heller 2001 0 In von Helversen, Heller, Mayer, Nemeth, Volleth, and Gombkötö, Naturwissenschaften 91:37:00 Alcathoe Myotis circassicus Benda, Garzaryan, & Vallo 2016 Greece, Nomos Evritanias, near the village of Kleistos, over Fournikos Patomos stream, 39°05'N, 21°49'E. Greece, Hungary, France, Portugal. Specimens from Bulgaria, Romania, and Ukraine previsously reported as ikonnikovi might represent alcathoe (von Helveren et al., 2001). Not listed. Data Deficient See Ruedi et al. (2002) and Benda et al. (2016). Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023 Myotis alcathoe 23 Alcathoe Whiskered Myotis Alcathoe Bat|Alcathoe Myotis|Alcathoe Whiskered Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA VESPERTILIONIFORMES NA NA VESPERTILIONOIDEA VESPERTILIONIDAE MYOTINAE NA Myotis Myotis alcathoe von Helversen & K. G. Heller in von Helversen, K. G. Heller, F. Mayer, Nemeth, Volleth, & Gombkötö 2001 0 Myotis_alcathoe von Helversen, O., Heller, K. G., Mayer, F., Nemeth, A., Volleth, M. & Gombkötö, P. (2001). Cryptic mammalian species: a new species of whiskered bat (Myotis alcathoe n. sp.) in Europe. Naturwissenschaften, 88, 217. https://link.springer.com/content/pdf/10.1007/s001140100225.pdf SMF 90249 "netted over a small stream, the Fournikos Potamos, near the village of Kleistos (39°05'N, 21°49'E), Novos Evritanias, Greece. 39.083 21.817 alcathoe von Helversen & K. G. Heller in von Helversen, K. G. Heller, F. Mayer, Nemeth, Volleth, & Gombkötö, 2001|circassicus Benda, Gazaryan, & Vallo, 2016 NA NA United Kingdom|Spain|France|Belgium|Luxembourg|Germany|Italy|Slovenia|Austria|Czech Republic|Poland|Slovakia|Hungary|Sweden|Ukraine|Croatia|Romania|Bulgaria|Turkey|Russia|Georgia|Azerbaijan? Asia|Europe Palearctic DD 0 0 0 Myotis_alcathoe 0 sciname match Myotis_alcathoe 0 IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022]. 136680 Myotis alcathoe ANIMALIA CHORDATA MAMMALIA CHIROPTERA VESPERTILIONIDAE Myotis alcathoe von Helversen &; Heller, 2001 Formerly included in Myotis mystacinus (Kuhl, 1817); the species was differentiated (von Helversen et al. 2001) on the base of karyological, genetic and echolocation characters. 518740 Myotis alcathoe Data Deficient 2016 2016-04-25 00:00:00 UTC 3.1 English European regional and Global assessment: Data Deficient (DD) EU 28 regional assessment: Data Deficient (DD) The species was only recently described (2001) and is known from few localities. It is not easy to identify morphologically. More localities are being reported, but no information is available on size of range and population or population trend. Consequently it is assessed as Data Deficient. Based on present knowledge the species is endemic to Europe and 90% of known locations are inside the EU. According to present scarce knowledge, the Alcathoe Myotis is a tree dwelling and forest foraging species, feeding primarily on moths and nematoceran flies (LuÄan et al . 2009, Danko et a l. 2010). It seems to prefer old and full-grown oak forests (LuÄan et al . 2009, Danko et al . 2010), but may also occur in rural gardens and urban habitats. It probably exploits underground habitats in winter. Summer colonies may number up to 80 individuals. The only known breeding colony ;was found in a tree hollow. ;
Damage to riparian forest is believed be a threat in parts of range (von Helversen et al. 2001). Wider forest and roost tree loss may also be threats. This species’ population size and trend are unknown. To date, ca. 15 localities are recorded in international publications (von Helversen 2001, Benda ;et al . ;2003, von Helversen 2004). However, new localities continue to be found (EMA Workshop 2006).
;
Unknown The Alcathoe Myotis (Myotis alcathoe ) was recently described and is poorly known (von Helversen ;et al. ;2001), but current information suggests that it is endemic to central and southern Europe. It occurs in Spain, France, ;Switzerland, Germany, Czech Republic, Slovakia, Hungary, Montenegro, Serbia, Bulgaria, and Greece (Ruedi ;et al . ;2002, Benda ;et al . ;2003, Agirre-Mendi ;et al. ;2004, von Helversen 2004, von Helversen ;et al . ;2006, P. Benda ;in litt. ;2006). Recent records revealed this bat’s presence also in Austria, Poland, Belgium, Romania and the United Kingdom (Spitzenberger et al. 2008, Jan et al. 2010, Sachanowicz ;et al. 2012, ;Uhrin ;et al. ; 2014, ;Nyssen et al. 2015).
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Terrestrial It is protected by national legislation in most range states. There are also international legal obligations for the species' protection through the Bonn Convention (Eurobats) and Bern Convention. It is included in Annex IV of the EU Habitats &; Species Directive, and some habitat protection may be provided through Natura 2000. Conservation recommendations include further research on distribution, population status and trends, ecology, habitat requirements, and threats. Measures to increase public awareness of this little-known species are also recommended. Palearctic FALSE FALSE Global & Europe Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 Vespertilionidae Myotis Unassigned - Myotis alcathoe von Helversen and Heller 2001 0 In von Helversen, Heller, Mayer, Nemeth, Volleth, and Gombkötö, Naturwissenschaften 91:37:00 Alcathoe Myotis circassicus Benda, Garzaryan, & Vallo 2016 Greece, Nomos Evritanias, near the village of Kleistos, over Fournikos Patomos stream, 39°05'N, 21°49'E. Greece, Hungary, France, Portugal. Specimens from Bulgaria, Romania, and Ukraine previsously reported as ikonnikovi might represent alcathoe (von Helveren et al., 2001). Not listed. Data Deficient See Ruedi et al. (2002) and Benda et al. (2016). Myotis alcathoe 1005360 23 Alcathoe Whiskered Myotis Alcathoe Bat|Alcathoe Myotis|Alcathoe Whiskered Bat Theria Placentalia Boreoeutheria Laurasiatheria CHIROPTERA VESPERTILIONIFORMES NA NA VESPERTILIONOIDEA Vespertilionidae MYOTINAE NA Myotis Myotis alcathoe von Helversen & K. G. Heller in von Helversen, K. G. Heller, F. Mayer, Nemeth, Volleth, & Gombkötö 2001 0 Myotis_alcathoe von Helversen, O., Heller, K. G., Mayer, F., Nemeth, A., Volleth, M. & Gombkötö, P. (2001). Cryptic mammalian species: a new species of whiskered bat (Myotis alcathoe n. sp.) in Europe. Naturwissenschaften, 88, 217. https://link.springer.com/content/pdf/10.1007/s001140100225.pdf SMF 90249 "netted over a small stream, the Fournikos Potamos, near the village of Kleistos (39°05'N, 21°49'E), Novos Evritanias, Greece. 39.08333 21.81667 alcathoe von Helversen & K. G. Heller in von Helversen, K. G. Heller, F. Mayer, Nemeth, Volleth, & Gombkötö, 2001|circassicus Benda, Gazaryan, & Vallo, 2016 NA NA United Kingdom|Spain|France|Belgium|Luxembourg|Germany|Italy|Slovenia|Austria|Czech Republic|Poland|Slovakia|Hungary|Sweden|Ukraine|Croatia|Romania|Bulgaria|Turkey|Russia|Georgia|Azerbaijan? Asia|Europe Palearctic DD 0 0 0 Myotis_alcathoe 0 sciname match Myotis_alcathoe 0 Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393 Myotis_alcathoe 1005360 23 Alcathoe Whiskered Myotis Alcathoe Bat|Alcathoe Myotis|Alcathoe Whiskered Bat Theria Placentalia Boreoeutheria Laurasiatheria Chiroptera Yangochiroptera NA NA Vespertilionoidea Vespertilionidae Myotinae NA Myotis Myotis alcathoe von Helversen & K.-G. Heller in von Helversen, K.-G. Heller, F. Mayer, Nemeth, Volleth, & GombkötÅ‘ 0 Myotis alcathoe Helversen, O. von, Heller, K.-G., Mayer, F., Nemeth, A., Volleth, M. and GombkötÅ‘, P. 2001-05-01. Cryptic mammalian species: a new species of whiskered bat (_Myotis alcathoe_ n. sp.) in Europe. Naturwissenschaften 88(5):217-223. https://doi.org/10.1007/s001140100225 SMF:MAMM:90249 holotype "netted over a small stream, the Fournikos Potamos, near the village of Kleistos (39°05'N, 21°49'E), Novos Evritanias, Greece. 39.08333 21.81667 NA NA United Kingdom|Spain|France|Belgium|Luxembourg|Germany|Italy|Slovenia|Austria|Czech Republic|Poland|Slovakia|Hungary|Sweden|Ukraine|Croatia|Romania|Bulgaria|Turkey|Russia|Georgia|Azerbaijan? Asia|Europe Palearctic DD 0 0 0 Myotis_alcathoe 0 sciname match Myotis_alcathoe 0 Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586 Vespertilionidae Myotis Myotis alcathoe von Helversen & Heller 2001 0 In von Helversen, Heller, Mayer, Nemeth, Volleth, and Gombkötö, Naturwissenschaften 91:37:00 Alcathoe Myotis circassicus Benda, Garzaryan, & Vallo 2016 Greece, Nomos Evritanias, near the village of Kleistos, over Fournikos Patomos stream, 39°05'N, 21°49'E. Greece, Hungary, France, Portugal. Specimens from Bulgaria, Romania, and Ukraine previsously reported as ikonnikovi might represent alcathoe (von Helveren et al., 2001). Not Listed Data Deficient See Ruedi et al. (2002) and Benda et al. (2016). Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505 NA Myotis alcathoe; Myotis alcathoe; Myotis alcathoe; Myotis alcathoe; Myotis alcathoe; Myotis alcathoe; alcathoevon; circassicus; circassicus; alcathoe; circassicus; Murin alcathoé; Nymphenfledermaus; Ratonero bigotudo pequeno; Alcathoe Bat; Alcathoe Myotis; Alcathoe Whiskered Bat; Alcathoe Whiskered Myotis; Alcathoe Bat; Alcathoe Myotis; Alcathoe Whiskered Bat; Alcathoe Myotis; Alcathoe Myotis; M. alcathoe