http://www.w3.org/ns/prov#wasDerivedFrom	http://purl.org/dc/elements/1.1/format	name_CH1_1980	name_MSW1_1982	name_CH3_1991	name_MSW2_1993	name_Koopman_1994	name_MSW3_2005	name_HMW_2019	name_BatNames_2022	name_MDD_2022	name_IUCN_2022	name_BatNames_2023	name_MDD_2023	name_MDD_2025_2.0	name_batnames_2025_1.7	name_MDD_2025_2.2	column151	taxonomic_notes_concatenated	column171	synonyms_CH1	subspecies__MSW2	synonyms__MSW1	synonyms_CH3	synonyms_MSW2	subspecies_Koopman94_interpreted	subspecies_MSW3_interpreted	synonym_MSW3_interpreted	subspecies_HMW_interpreted	synonym_HMW_interpreted	subspecies_batnames_interpreted	synonym_batnames_interpreted	synonym_MDD_interpreted	synonym_IUCN_interpreted	subspecies_batnames2023_interpreted	synonym_batnames2023_interpreted	synonym_MDD2023_interpreted	synonym_MDD2025_interpreted	subspecies_batnames2025_interpreted	synonyms_batnames2025_interpreted	nominalNames	column391	docOrigin_CH1	commonName_CH1	distribution_CH1	docOrigin_MSW1	column451	typeLocality_MSW1	authority_MSW1	year_MSW1	citation_MSW1	distribution	comment_MSW1	docOrigin_CH3	commonName_CH3	distribution_CH3	docOrigin_MSW2	authority_MSW2	year_MSW2	citation_MSW2	comments_MSW2	distribution_MSW2	typeLocality_MSW2	docOrigin_Koopman94	authority_Koopman94	year_Koopman94	description_Koopman94	distribution_Koopman94	diversity_Koopman94	subspecies_Koopman94	page	rank	name	authority	year	parent	parent_rank	corrected_name	actual_species_count	claimed_species_count	dental_formula	description	diversity	full_subspecies_text	name_line	species_index	subspecies	synonym	text	docOrigin_MSW3	order_MSW3	family_MSW3	subfamily_MSW3	tribe_MSW3	name_MSW3	genus_MSW3	subgenus_MSW3	species_MSW3	authoritySpeciesAuthor_MSW3	(parentheses (1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common Name_batnames	synonyms_batnames	type_locality_batnames	Distribution_batnames	CITES_batnames	IUCN_batnames	comments_batnames	docOrigin_MDD	name_MDD	phylosort_MDD	mainCommonName_MDD	otherCommonNames_MDD	subclass_MDD	infraclass_MDD	magnorder_MDD	superorder_MDD	order_MDD	suborder_MDD	infraorder_MDD	parvorder_MDD	superfamily_MDD	family_MDD	subfamily_MDD	tribe_MDD	genus_MDD	subgenus_MDD	specificEpithet_MDD	authoritySpeciesAuthor_MDD	authoritySpeciesYear_MDD	authorityParentheses_MDD	originalNameCombination_MDD	authoritySpeciesCitation_MDD	authoritySpeciesLink_MDD	holotypeVoucher_MDD	holotypeVoucherURIs_MDD	typeLocality_MDD	typeLocalityLatitude_MDD	typeLocalityLongitude_MDD	nominalNames_MDD	taxonomyNotes_MDD	taxonomyNotesCitation_MDD	countryDistribution_MDD	continentDistribution_MDD	biogeographicRealm_MDD	iucnStatus_MDD	extinct_MDD	domestic_MDD	flagged_MDD	CMW_sciName_MDD	diffSinceCMW_MDD	MSW3_matchtype_MDD	MSW3_sciName_MDD	diffSinceMSW3_MDD	docOrigin_IUCN	internalTaxonId_IUCN	NAME_IUCN	kingdomName_IUCN	phylumName_IUCN	className_IUCN	orderName_IUCN	familyName_IUCN	genusName_IUCN	speciesName_IUCN	authoritySpeciesAuthorYear_IUCN	taxonomicNotes_IUCN	assessmentId_IUCN	scientificName_IUCN	redlistCategory_IUCN	redlistCriteria_IUCN	yearPublished_IUCN	assessmentDate_IUCN	criteriaVersion_IUCN	language_IUCN	rationale_IUCN	habitat_IUCN	threats_IUCN	population_IUCN	populationTrend_IUCN	range_IUCN	useTrade_IUCN	systems_IUCN	conservationActions_IUCN	realm_IUCN	yearLastSeen_IUCN	possiblyExtinct_IUCN	possiblyExtinctInTheWild_IUCN	scopes_IUCN	docOrigin_batnames2023	FAMILY_batnames2023	GENUS_batnames2023	SUBGENUS_batnames2023	SPECIES_batnames2023	authoritySpeciesAuthor_batnames2023	authoritySpeciesYearbatnames2023	PARENTHESES_batnames2023 (1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name MDD2023	id_MDD2023	phylosort_MDD2023	mainCommonName_MDD2023	otherCommonNames_MDD2023	subclass_MDD2023	infraclass_MDD2023	magnorder_MDD2023	superorder_MDD2023	order_MDD2023	suborder_MDD2023	infraorder_MDD2023	parvorder_MDD2023	superfamily_MDD2023	Family_mdd2023	subfamily_MDD2023	tribe_MDD2023	genus_MDD2023	subgenus_MDD2023	specificEpithet_MDD2023	authoritySpeciesAuthor_MDD2023	authoritySpeciesYear_MDD2023	authorityParentheses_MDD2023	originalNameCombination_MDD2023	authoritySpeciesCitation_MDD2023	authoritySpeciesLink_MDD2023	holotypeVoucher_MDD2023	holotypeVoucherURIs_MDD2023	typeLocality_MDD2023	typeLocalityLatitude_MDD2023	typeLocalityLongitude_MDD2023	nominalNames_MDD2023	taxonomyNotes_MDD2023	taxonomyNotesCitation_MDD2023	distributionNotes_MDD2023	distributionNotesCitation_MDD2023	subregionDistribution_MDD2023	countryDistribution_MDD2023	continentDistribution_MDD2023	biogeographicRealm_MDD2023	iucnStatus_MDD2023	extinct_MDD2023	domestic_MDD2023	flagged_MDD2023	CMW_sciName_MDD2023	diffSinceCMW_MDD2023	MSW3_matchtype_MDD2023	MSW3_sciName_MDD2023	diffSinceMSW3_MDD2023	docOrigin_MDD2025	sciName	id	phylosort	mainCommonName	otherCommonNames	subclass	infraclass	magnorder	superorder	order	suborder	infraorder	parvorder	superfamily	family	subfamily	tribe	genus	subgenus	specificEpithet	authoritySpeciesAuthor	authorityParentheses	originalNameCombination	authoritySpeciesCitation	authoritySpeciesLink	typeVoucher	typeKind	typeVoucherURIs	typeLocality	typeLocalityLatitude	typeLocalityLongitude	taxonomyNotes	taxonomyNotesCitation	distributionNotes	distributionNotesCitation	subregionDistribution	countryDistribution	continentDistribution	biogeographicRealm	iucnStatus	extinct	domestic	flagged	CMW_sciName	diffSinceCMW	MSW3_matchtype	MSW3_sciName	diffSinceMSW3	docOrigin_batnames2025	Family	Genus	Subgenus	Species	Author	Date	Parentheses (1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L811	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Myotis albescens	Myotis albescens	Myotis albescens	Myotis albescens	Myotis albescens	Myotis albescens	Myotis albescens	Myotis albescens	Myotis albescens	Myotis albescens	Myotis albescens	Myotis albescens	Myotis albescens	Myotis albescens	Myotis albescens		[MSW2] Subgenus Leuconoe. Includes argentatus; see LaVal (1973fl:25).; [MSW3] Includes argentatus; see LaVal (1973a). Reviewed in part by López-González et al. (2001). Does not include aenobarbus, which is here placed in Nycticeius following Carter and Dolan (1978); also see Husson (1962). Although I follow Koopman (1993) in listing isidori as a synonym, there are serious problems with idenfication of the holotype; see Carter and Dolan (1978), who suggested that this name might actually belong in Pipistrellus. Does not include mundus; see LaVal (1973a). Apparently closely related to nigricans, levis, and oxyotus; see Ruedi and Mayer (2001).; [HMW] Vespertilio albescens E. Geoffroy Saint-Hilaire, 1806 , “ Paraguay .” Based on neotype selection,restricted by R. K. LaVal in 1973 to “Yaguaron, Paraguari , Paraguay .” Subgenus Pizonyx; albescens species group. Monotypic.; [batnames2022] Includes argentatus ; see LaVal (1973 a ). Includes punensis ; see Moratelli and Wilson (2011).  Reviewed inpart by LÃ³pez-GonzÃ¡lez et al. (2001). Does not include aenobarbus , which is here placed in Nycticeius following Carter andDolan (1978); also see Husson (1962). Although I follow Koopman (1993) in listing isidori as a synonym, there are serious problemswith idenfication of the holotype; see Carter and Dolan (1978), who suggested that this name might actually belong in Pipistrellus .Does not include mundus ; see LaVal (1973 a ). Apparently closely related to nigricans , levis , and oxyotus ; see Ruedi and Mayer (2001).; [IUCN] Apparently closely related to nigricans , levis , and oxyotus .; [batnames2023] Includes argentatus ; see LaVal (1973 a ). Includes punensis ; see Moratelli and Wilson (2011).  Reviewed inpart by LÃ³pez-GonzÃ¡lez et al. (2001). Does not include aenobarbus , which is here placed in Nycticeius following Carter andDolan (1978); also see Husson (1962). Although I follow Koopman (1993) in listing isidori as a synonym, there are serious problemswith idenfication of the holotype; see Carter and Dolan (1978), who suggested that this name might actually belong in Pipistrellus .Does not include mundus ; see LaVal (1973 a ). Apparently closely related to nigricans , levis , and oxyotus ; see Ruedi and Mayer (2001).; [batnames2025_1.7] Includes argentatus; see LaVal (1973a). Includes punensis; see Moratelli and Wilson (2011).  Reviewed inpart by LÃ³pez-GonzÃ¡lez et al. (2001). Does not include aenobarbus, which is here placed in Nycticeius following Carter andDolan (1978); also see Husson (1962). Although I follow Koopman (1993) in listing isidori as a synonym, there are serious problemswith idenfication of the holotype; see Carter and Dolan (1978), who suggested that this name might actually belong in Pipistrellus.Does not include mundus; see LaVal (1973a). Apparently closely related to nigricans, levis, and oxyotus; see Ruedi and Mayer (2001).				argentatus		aenobarbus, argentatus, arsinoe, isidori, leucogaster, mundus.			argentatus, isidori, leucogaster			albescens 	albescens - argentatus, isidori, leucogaster	albescens, leucogaster, punensis	Apparently closely related to nigricans , levis , and oxyotus .	albescens 	albescens - argentatus, isidori, leucogaster, punensis	albescens, leucogaster, punensis 	albescens, leucogaster, punensis, argentatus	albescens 	albescens - argentatus, isidori, leucogaster	albescens (Ã‰. Geoffroy Saint-Hilaire, 1806)|leucogaster (H. R. Schinz, 1821)|punensis J. A. Allen, 1914|argentatus Dalquest & E. R. Hall, 1947		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Paraguay myotis	S Mexico – Paraguay, Uruguay	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Myotis albescens	Paraguay, Paraguari, Yaguaron (of neotype).	E. Geoffroy	1806	Ann. Mus. Hist. Nat. Paris, 8:204.	Distribution: Ranging from southern Mexico to central Argentina but west of the Andes not south of Ecuador.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Silver-tipped myotis	S Mexico – N Argentina, Uruguay	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	E. Geoffrey	1806	Ann. Mus. Hist. Nat. Paris, 8:204.	Subgenus Leuconoe. Includes argentatus; see LaVal (1973fl:25).	S Veracruz (Mexico) to Uruguay and N Argentina.	Paraguay, Paraguari, Yaguaron (of neotype).		E. GEOFFROY	1806	Size fairly small (forearm length 33-39 mm; condylobasal length, 12-14 mm). Margin of uropatagium usually with a poorly developed fringe of hair. Rostrum fairly slender. Middle upper premolar in toothrow. Fur whitetipped.	Distribution: Ranging from southern Mexico to central Argentina but west of the Andes not south of Ecuador.	No subspecies are currently recognized.		108	species	M. albescens	E. GEOFFROY	1806	Leuconoe	subgenus	Myotis albescens				Size fairly small (forearm length 33-39 mm; condylobasal length, 12-14 mm). Margin of uropatagium usually with a poorly developed fringe of hair. Rostrum fairly slender. Middle upper premolar in toothrow. Fur whitetipped.	No subspecies are currently recognized.		76. M. albescens (E. GEOFFROY 1806) [albescens group].	76	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Myotinae		Myotis albescens	Myotis		albescens	E. Geoffroy	y	1806		Ann. Mus. Natn. Hist. Nat. Paris	8		204		Silver-tipped Myotis	Paraguay, Paraguari, Yaguaron (of neotype).	S Veracruz (Mexico), Guatemala, Honduras, Nicaragua, Panama, Colombia, Venezuela, Guyana, Surinam, Equador, Peru, Brazil, Uruguay, N Argentina, Paraguay, and Bolivia.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	argentatus Dalquest and Hall, 1947; isidori D’Orbigny and Gervais, 1847; leucogaster Schinz, 1821.	Includes argentatus; see LaVal (1973a). Reviewed in part by López-González et al. (2001). Does not include aenobarbus, which is here placed in Nycticeius following Carter and Dolan (1978); also see Husson (1962). Although I follow Koopman (1993) in listing isidori as a synonym, there are serious problems with idenfication of the holotype; see Carter and Dolan (1978), who suggested that this name might actually belong in Pipistrellus. Does not include mundus; see LaVal (1973a). Apparently closely related to nigricans, levis, and oxyotus; see Ruedi and Mayer (2001).	4C3D87E8FF496AF7FF3C9ACC1C93B9F5	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	940	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF496AF7FF3C9ACC1C93B9F5.xml	Myotis albescens	Vespertilionidae	Myotis	albescens		1806	Murin poudré @fr | Silberspitzen-Mausohr @de | Ratonero cenizo @es	Vespertilio albescens E. Geoffroy Saint-Hilaire, 1806 , “ Paraguay .” Based on neotype selection,restricted by R. K. LaVal in 1973 to “Yaguaron, Paraguari , Paraguay .” Subgenus Pizonyx; albescens species group. Monotypic.	Widely distributed in Neotropics, from S Veracruz in Mexico, S through Central and South America to Peru , Bolivia , S Brazil , Paraguay , Uruguay , and N Argentina .	Head-body c¢. 46— 49 mm , tall 35-46 mm , ear 9-14 mm , hindfoot 6-9 mm , forearm 32-2-39- 5 mm ; weight 4-8 g . South American populations show a trend toward increasing size southward. Fur is silky and long (dorsal fur 4-9 mm ; ventral fur 3-8 mm ). Dorsal hairs are bicolored, with dark brown bases and antique brown tips (one-fifth of total length), giving golden appearance; yellowish tips are less evident in a few specimens, in which dorsal fur can be nearly unicolored. Ventral hairs are strongly bicolored, with dark brown bases and generally white tips (one-third the length), giving frosted appearance; ventral fur often becomes progressively paler (whiter) from upper thorax to pelvic region. Ears are comparatively short, extending forward halfway from eye to nostril. Antitragal notch is barely evident. Tragus is long and slender, tapering slightly at tip. Membranes vary from cinnamon-brown to mummy brown; in some individuals, wing membrane has translucent brown appearance. Plagiopatagium is attached to feet by a broad band of membrane; fringe of hairs along trailing edge of uropatagium is present; and upper and lower surfaces of uropatagium are barely covered with hairs. Skull is moderate in size (greatest length of skull 13-1-15- 2 mm ); parietal is normally inclined forward; occipital region is generally rounded posteriorly; braincase is globose; sagittal crest is generally absent, but when present,it is very low; lambdoidal crests are generally present, with low to medium development; forehead slopeslittle; rostrum is narrow relative to braincase; postorbital constriction is great relative to width of braincase. P° is generally aligned with P* and is visible in profile; hypocone on M' and M? is absent or poorly developed. Bacula is large, and individual variation is mostly in depth, not shape. Bacular measurements are length 0-85-0-90, depth 0-30-0-41, and width 0-37-0-50. Chromosomal complement has 2n = 44 and FN = 50, with four metacentric pairs and 17 acrocentric pairs of autosomes. X- and Y-chromosomes are submetacentric.	Tropical rainforests, savannas, Chacoan scrublands, thorn scrub lowlands, evergreen forests, open habitats (e.g. wetlands and grasslands), and also secondary forests, agricultural landscapes, and semi-urban areas from sea level up to elevations of ¢. 1500 m (most commonly below 500 m ). The Silver-tipped Myotis is apparently more abundant in areas that are flooded at least part of the year.	The Silver-tipped Myotis forages in forested areas and over water. Diets include various insects, especially Coleoptera , Diptera , Hemiptera, Homoptera , Hymenoptera , Lepidoptera , Neuroptera, Trichoptera , and other arthropods, such as spiders ( Araneae ). Fish scales also have been reported in stomach contents. It has a trawler foraging style,flying a few centimeters above water orflat surfaces, using highintensity echolocation calls to detect, track, and assess targets, and capturing prey with its feet. In the Brazilian Amazonia, it predominantly forages in forest canopies and subcanopies, but in riparian forest or near water bodies, it mostly uses space closest to the ground or water surface.	The Silver-tipped Myotis is polyestrous, and mating occurs 2-3 times annually. In Paraguay , spermatozoa were present in reproductive tracts of females at the beginning of the reproductive season in May; ovulation,fertilization, and implantation occurs c¢.1-3 months after copulation, indicating possible sperm storage and delayed fertilization; first parturition peak occurs in October, followed by copulation and second pregnancy, and possible third pregnancy in January-March. First gestation lasts c.3 months, and second and third gestations might be shorter. Following third peak, reproductive activity declines until beginning of a new annual cycle. Lactation lasts c.1 month, and lactating females were collected in Paraguay in October-December but also May-June during second and third breeding periods. Pregnant females have been reported in January in Mexico, January and July in Central America, June-September in northern South America, and December—February in south-eastern Brazil and northern South America. In Peru , parturition occurred during dry and wet seasons in eastern lowlands and only during dry season on the Pacific slope. Females have one young per pregnancy. Lactating females also were reported in February-April and November in northern South America; October in south-eastern Brazil ; and September— November in southern South America. Individuals reach sexual maturity at c.1 year of age, but females can mate at c¢.2 months.	Silver-tipped Myotis are nocturnal, emerging soon after sunset. Activity peaks occur immediately after dusk and just before dawn, periods that correspond with greatest abundance offlying diurnal and nocturnal insects. Natural roosts include hollow trees, under bark of trees, trunks, cracks, and crevices in rocks, and rocky cliffs. It is frequently found in human buildings, including attics, under roofs, under palm logs and roofs of palm-thatched huts, and in locations with some natural light and exposure, such as outer walls of buildings or open-sided attics. Captures and roosts are associated with areas near or around water. Wing morphology and echolocation frequency are typical of aerial insectivore that uses using cluttered spaces. Echolocation calls show FM initial component terminating with short CF component. Mean call parameters in South America are start frequency of 92.7 kHz (84-8-100-8 kHz), end frequency of 51-4 kHz (47-4-56-5 kHz), peak frequency of 64 kHz (57-3-72-6 kHz), bandwidth of 41-3 kHz (34-1-48 kHz), and call duration of 2-4 milliseconds (1-9-3 milliseconds).	The Silver-tipped Myotis forms small or large groups in roosts; solitary individuals are uncommon. Colonies are mixed-sex. It can cohabit the same roost with Argentine Serotines ( Eptesicus furinalis ), Common Black Myotis ( M. nigricans ), Riparian Myotis (M. riparius ), Sinaloan Mastiff Bats (Molossus sinaloae), Dwarf Dog-faced Bats (Molossops temminckii), and Black Bonneted Bats (Eumops auripendulus), among others.	Classified as Least Concern on The IUCN Red List. Silvertipped Myotis is widespread and presumably has a large population.	Arias-Aguilar et al. (2018) | Ascorra et al. (1996) | Barquez, Diaz, Samudio & Arroyo-Cabrales (2016) | Braun et al. (2009) | Fenton & Bogdanowicz (2002) | Gamboa & Diaz (2019) | Kalko et al. (1996) | LaVal (1973b) | Lopez-Gonzélez et al. (2001) | Marques et al. (2016) | Moratelli & Morielle-Versute (2007) | Moratelli & Oliveira (2011) | Moratelli, Dewynter et al. (2015) | Moratelli, Gardner et al. (2013) | Myers (1977) | Simmons (2005) | Whitaker & Findley (1980) | Wilson (2008b)	https://zenodo.org/record/6398773/files/figure.png	401. Silver-upped Myotis Myotis albescens French: Murin poudré / German: Silberspitzen-Mausohr / Spanish: Ratonero cenizo Taxonomy. Vespertilio albescens E. Geoffroy Saint-Hilaire, 1806 , “ Paraguay .” Based on neotype selection,restricted by R. K. LaVal in 1973 to “Yaguaron, Paraguari , Paraguay .” Subgenus Pizonyx; albescens species group. Monotypic. Distribution. Widely distributed in Neotropics, from S Veracruz in Mexico, S through Central and South America to Peru , Bolivia , S Brazil , Paraguay , Uruguay , and N Argentina . Descriptive notes. Head-body c¢. 46— 49 mm , tall 35-46 mm , ear 9-14 mm , hindfoot 6-9 mm , forearm 32-2-39- 5 mm ; weight 4-8 g . South American populations show a trend toward increasing size southward. Fur is silky and long (dorsal fur 4-9 mm ; ventral fur 3-8 mm ). Dorsal hairs are bicolored, with dark brown bases and antique brown tips (one-fifth of total length), giving golden appearance; yellowish tips are less evident in a few specimens, in which dorsal fur can be nearly unicolored. Ventral hairs are strongly bicolored, with dark brown bases and generally white tips (one-third the length), giving frosted appearance; ventral fur often becomes progressively paler (whiter) from upper thorax to pelvic region. Ears are comparatively short, extending forward halfway from eye to nostril. Antitragal notch is barely evident. Tragus is long and slender, tapering slightly at tip. Membranes vary from cinnamon-brown to mummy brown; in some individuals, wing membrane has translucent brown appearance. Plagiopatagium is attached to feet by a broad band of membrane; fringe of hairs along trailing edge of uropatagium is present; and upper and lower surfaces of uropatagium are barely covered with hairs. Skull is moderate in size (greatest length of skull 13-1-15- 2 mm ); parietal is normally inclined forward; occipital region is generally rounded posteriorly; braincase is globose; sagittal crest is generally absent, but when present,it is very low; lambdoidal crests are generally present, with low to medium development; forehead slopeslittle; rostrum is narrow relative to braincase; postorbital constriction is great relative to width of braincase. P° is generally aligned with P* and is visible in profile; hypocone on M' and M? is absent or poorly developed. Bacula is large, and individual variation is mostly in depth, not shape. Bacular measurements are length 0-85-0-90, depth 0-30-0-41, and width 0-37-0-50. Chromosomal complement has 2n = 44 and FN = 50, with four metacentric pairs and 17 acrocentric pairs of autosomes. X- and Y-chromosomes are submetacentric. Habitat. Tropical rainforests, savannas, Chacoan scrublands, thorn scrub lowlands, evergreen forests, open habitats (e.g. wetlands and grasslands), and also secondary forests, agricultural landscapes, and semi-urban areas from sea level up to elevations of ¢. 1500 m (most commonly below 500 m ). The Silver-tipped Myotis is apparently more abundant in areas that are flooded at least part of the year. Food and Feeding. The Silver-tipped Myotis forages in forested areas and over water. Diets include various insects, especially Coleoptera , Diptera , Hemiptera, Homoptera , Hymenoptera , Lepidoptera , Neuroptera, Trichoptera , and other arthropods, such as spiders ( Araneae ). Fish scales also have been reported in stomach contents. It has a trawler foraging style,flying a few centimeters above water orflat surfaces, using highintensity echolocation calls to detect, track, and assess targets, and capturing prey with its feet. In the Brazilian Amazonia, it predominantly forages in forest canopies and subcanopies, but in riparian forest or near water bodies, it mostly uses space closest to the ground or water surface. Breeding. The Silver-tipped Myotis is polyestrous, and mating occurs 2-3 times annually. In Paraguay , spermatozoa were present in reproductive tracts of females at the beginning of the reproductive season in May; ovulation,fertilization, and implantation occurs c¢.1-3 months after copulation, indicating possible sperm storage and delayed fertilization; first parturition peak occurs in October, followed by copulation and second pregnancy, and possible third pregnancy in January-March. First gestation lasts c.3 months, and second and third gestations might be shorter. Following third peak, reproductive activity declines until beginning of a new annual cycle. Lactation lasts c.1 month, and lactating females were collected in Paraguay in October-December but also May-June during second and third breeding periods. Pregnant females have been reported in January in Mexico, January and July in Central America, June-September in northern South America, and December—February in south-eastern Brazil and northern South America. In Peru , parturition occurred during dry and wet seasons in eastern lowlands and only during dry season on the Pacific slope. Females have one young per pregnancy. Lactating females also were reported in February-April and November in northern South America; October in south-eastern Brazil ; and September— November in southern South America. Individuals reach sexual maturity at c.1 year of age, but females can mate at c¢.2 months. Activity patterns. Silver-tipped Myotis are nocturnal, emerging soon after sunset. Activity peaks occur immediately after dusk and just before dawn, periods that correspond with greatest abundance offlying diurnal and nocturnal insects. Natural roosts include hollow trees, under bark of trees, trunks, cracks, and crevices in rocks, and rocky cliffs. It is frequently found in human buildings, including attics, under roofs, under palm logs and roofs of palm-thatched huts, and in locations with some natural light and exposure, such as outer walls of buildings or open-sided attics. Captures and roosts are associated with areas near or around water. Wing morphology and echolocation frequency are typical of aerial insectivore that uses using cluttered spaces. Echolocation calls show FM initial component terminating with short CF component. Mean call parameters in South America are start frequency of 92.7 kHz (84-8-100-8 kHz), end frequency of 51-4 kHz (47-4-56-5 kHz), peak frequency of 64 kHz (57-3-72-6 kHz), bandwidth of 41-3 kHz (34-1-48 kHz), and call duration of 2-4 milliseconds (1-9-3 milliseconds). Movements, Home range and Social organization. The Silver-tipped Myotis forms small or large groups in roosts; solitary individuals are uncommon. Colonies are mixed-sex. It can cohabit the same roost with Argentine Serotines ( Eptesicus furinalis ), Common Black Myotis ( M. nigricans ), Riparian Myotis (M. riparius ), Sinaloan Mastiff Bats (Molossus sinaloae), Dwarf Dog-faced Bats (Molossops temminckii), and Black Bonneted Bats (Eumops auripendulus), among others. Status and Conservation. Classified as Least Concern on The IUCN Red List. Silvertipped Myotis is widespread and presumably has a large population. Bibliography. Arias-Aguilar et al. (2018), Ascorra et al. (1996), Barquez, Diaz, Samudio & Arroyo-Cabrales (2016), Braun et al. (2009), Fenton & Bogdanowicz (2002), Gamboa & Diaz (2019), Kalko et al. (1996), LaVal (1973b), Lopez-Gonzélez et al. (2001), Marques et al. (2016), Moratelli & Morielle-Versute (2007), Moratelli & Oliveira (2011), Moratelli, Dewynter et al. (2015), Moratelli, Gardner et al. (2013), Myers (1977), Simmons (2005), Whitaker & Findley (1980), Wilson (2008b).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Myotis albescens	Myotis	Unassigned-Myotis	albescens	E. Geoffroy	1806	1	Ann. Mus. Natn. Hist. Nat. Paris	0.475	Silver-tipped Myotis	 argentatus Dalquest and Hall, 1947; isidori D&#39;Orbigny and Gervais, 1847; leucogaster Schinz, 1821.	Paraguay, Paraguari, Yaguaron (of neotype).	S Veracruz (Mexico), Guatemala, Honduras, Nicaragua, Panama, Colombia, Venezuela, Guyana, Surinam, Equador, Peru, Brazil, Uruguay, N Argentina, Paraguay, and Bolivia.	Not listed.	Least Concern	Includes argentatus ; see LaVal (1973 a ). Includes punensis ; see Moratelli and Wilson (2011).  Reviewed inpart by LÃ³pez-GonzÃ¡lez et al. (2001). Does not include aenobarbus , which is here placed in Nycticeius following Carter andDolan (1978); also see Husson (1962). Although I follow Koopman (1993) in listing isidori as a synonym, there are serious problemswith idenfication of the holotype; see Carter and Dolan (1978), who suggested that this name might actually belong in Pipistrellus .Does not include mundus ; see LaVal (1973 a ). Apparently closely related to nigricans , levis , and oxyotus ; see Ruedi and Mayer (2001).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Myotis albescens	23	Silver-tipped Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MYOTINAE	NA	Myotis	Pizonyx	albescens	Ã‰. Geoffroy Saint-Hilaire	1806	1	Vespertilio_albescens	Geoffroy Saint-Hilaire, Ã‰. (1806). MÃ©moire sur le genre et les espÃ¨ces de Vespertilion, l'un des genres de la famille des Chauve-souris. Annales du MusÃ©um d'histoire naturelle, 8, 204.	https://www.biodiversitylibrary.org/item/51185#page/246/mode/1up	AMNH 205195 [neotype]		"Paraguay." Based on neotype selection, restricted by R. K. LaVal in 1973 to "Yaguaron, Paraguari, Paraguay."			albescens (Ã‰. Geoffroy Saint-Hilaire, 1806)|leucogaster (Schinz, 1821)|punensis J. A. Allen, 1914	NA	NA	Mexico|Guatemala|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Guyana|Suriname|French Guiana|Ecuador|Peru|Brazil|Bolivia|Paraguay|Uruguay|Argentina	North America|South America	Nearctic|Neotropic	LC	0	0	0	Myotis_albescens	0	sciname match	Myotis_albescens	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	14140	Myotis albescens	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Myotis	albescens	(Ã‰. Geoffroy, 1806)	Apparently closely related to nigricans , levis , and oxyotus .	20000000	Myotis albescens	Least Concern		2016	2015-07-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern in because of its wide distribution, presumed large population, occurrence in a number of protected areas, tolerance to some degree of habitat modification, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	Silver-tipped Myotis is found in forests, urban habitats, and parks. It is an insectivorous species.	There are no major threats to this species throughout its range.	This species is abundant through out its range, although it is not common in Mexico (Arroyo-Cabrales pers. comm).	Stable	The species occurs in Southern Veracruz (Mexico), Guatemala, Honduras, Nicaragua, Panama, Colombia, Venezuela, Guyana, Suriname, Ecuador, Peru, Brazil, Uruguay, Northern Argentina, Paraguay, and Bolivia (Simmons 2005).		Terrestrial	The species needs taxonomic review (Braquez pers. comm). It is found in protected areas, and in Mexico it is listed as subject to special protection under NOM - 059 - SEMARNAT - 2001 (Arroyo-Cabrales pers. comm).	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Myotis	Unassigned - Myotis	albescens	Ã‰. Geoffroy Saint-Hilaire	1806	1	Ann. Mus. Natn. Hist. Nat. Paris	0.475	Silver-tipped Myotis	 argentatus Dalquest and Hall, 1947; isidori D&#39;Orbigny and Gervais, 1847; leucogaster Schinz, 1821.	Paraguay, Paraguari, Yaguaron (of neotype).	S Veracruz (Mexico), Guatemala, Honduras, Nicaragua, Panama, Colombia, Venezuela, Guyana, Surinam, Equador, Peru, Brazil, Uruguay, N Argentina, Paraguay, and Bolivia.	Not listed.	Least Concern	Includes argentatus ; see LaVal (1973 a ). Includes punensis ; see Moratelli and Wilson (2011).  Reviewed inpart by LÃ³pez-GonzÃ¡lez et al. (2001). Does not include aenobarbus , which is here placed in Nycticeius following Carter andDolan (1978); also see Husson (1962). Although I follow Koopman (1993) in listing isidori as a synonym, there are serious problemswith idenfication of the holotype; see Carter and Dolan (1978), who suggested that this name might actually belong in Pipistrellus .Does not include mundus ; see LaVal (1973 a ). Apparently closely related to nigricans , levis , and oxyotus ; see Ruedi and Mayer (2001).	Myotis albescens	1005359	23	Silver-tipped Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MYOTINAE	NA	Myotis	Pizonyx	albescens	Ã‰. Geoffroy Saint-Hilaire	1806	1	Vespertilio_albescens	Geoffroy Saint-Hilaire, Ã‰. (1806). MÃ©moire sur le genre et les espÃ¨ces de Vespertilion, l'un des genres de la famille des Chauve-souris. Annales du MusÃ©um d'histoire naturelle, 8, 204.	https://www.biodiversitylibrary.org/item/51185#page/246/mode/1up	AMNH 205195 [neotype]		"Paraguay." Based on neotype selection, restricted by R. K. LaVal in 1973 to "Yaguaron, Paraguari, Paraguay."			albescens (Ã‰. Geoffroy Saint-Hilaire, 1806)|leucogaster (Schinz, 1821)|punensis J. A. Allen, 1914	NA	NA				Mexico|Guatemala|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Guyana|Suriname|French Guiana|Ecuador|Peru|Brazil|Bolivia|Paraguay|Uruguay|Argentina	North America|South America	Nearctic|Neotropic	LC	0	0	0	Myotis_albescens	0	sciname match	Myotis_albescens	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Myotis_albescens	1005359	23	Silver-tipped Myotis		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Myotinae	NA	Myotis	Pizonyx	albescens	Ã‰. Geoffroy Saint-Hilaire	1	Vespertilio albescens	Geoffroy Saint-Hilaire, Ã‰. 1806. MÃ©moire sur le genre et les espÃ¨ces de Vespertilion, l'un des genres de la famille des chauve-souris. Annales du MusÃ©um d'histoire naturelle 8:187-205.	https://www.biodiversitylibrary.org/page/29407874	AMNH M-205195	neotype	http://portal.vertnet.org/o/amnh/mammals?id=urn-catalog-amnh-mammals-m-205195	"Paraguay." Based on neotype selection, restricted by R. K. LaVal in 1973 to "Yaguaron, Paraguari, Paraguay."	-25.55	-57.2833	NA	NA				Mexico|Guatemala|Honduras|Nicaragua|Costa Rica|Panama|Colombia|Venezuela|Guyana|Suriname|French Guiana|Ecuador|Peru|Brazil|Bolivia|Paraguay|Uruguay|Argentina	North America|South America	Nearctic|Neotropic	LC	0	0	0	Myotis_albescens	0	sciname match	Myotis_albescens	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Myotis	Pizonyx	albescens	Ã‰. Geoffroy Saint-Hilaire	1806	1	Ann. Mus. Natn. Hist. Nat. Paris	0.475	Silver-tipped Myotis	argentatus Dalquest and Hall, 1947; isidori D&#39;Orbigny and Gervais, 1847; leucogaster Schinz, 1821.	Paraguay, Paraguari, Yaguaron (of neotype).	S Veracruz (Mexico), Guatemala, Honduras, Nicaragua, Panama, Colombia, Venezuela, Guyana, Surinam, Equador, Peru, Brazil, Uruguay, N Argentina, Paraguay, and Bolivia.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/14140/22049892/' target='_blank'>Least Concern</a>	Includes argentatus; see LaVal (1973a). Includes punensis; see Moratelli and Wilson (2011).  Reviewed inpart by LÃ³pez-GonzÃ¡lez et al. (2001). Does not include aenobarbus, which is here placed in Nycticeius following Carter andDolan (1978); also see Husson (1962). Although I follow Koopman (1993) in listing isidori as a synonym, there are serious problemswith idenfication of the holotype; see Carter and Dolan (1978), who suggested that this name might actually belong in Pipistrellus.Does not include mundus; see LaVal (1973a). Apparently closely related to nigricans, levis, and oxyotus; see Ruedi and Mayer (2001).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Myotis albescens; Myotis albescens; Myotis albescens; Myotis albescens; Myotis albescens; Myotis albescens; argentatus; isidori; leucogaster; argentatus; isidori; leucogaster; albescens; leucogaster; punensis; Murin poudré; Silberspitzen-Mausohr; Ratonero cenizo; Silver-tipped Myotis; Silver-tipped Myotis; Silver-tipped Myotis; M. albescens
