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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L797	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Murina walstoni	Murina walstoni	Murina walstoni	Murina walstoni	Murina walstoni	Murina walstoni	Murina walstoni	Murina walstoni	Murina walstoni		[HMW] Murina walstoni Furey, Csorba & Nguyen Truong Son, 2011 , “ Cambodia , Ratanakiri Province , Veun Sai District, Veun Sai Protected Forest, 14°01'49”N , 106°45'06"E , elevation 110 m .” Murina walstoni appears to be related to M. swilla and M. aenea based on limited genetic data. Monotypic.; [MDD2022] recently described; [IUCN] <span lang="EN-US">The taxon was described in 2011 based on specimens from Cambodia and Viet Nam.; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														walstoni	<span lang="EN-US">The taxon was described in 2011 based on specimens from Cambodia and Viet Nam.			walstoni	walstoni			walstoni Furey, Csorba, & Nguyen Truong Son in Csorba, Nguyen Truong Son, Saveng, & Furey, 2011						N/A																																								NA																											4C3D87E8FF616ADEFF859E2F1636BF37	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	915	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF616ADEFF859E2F1636BF37.xml	Murina walstoni	Vespertilionidae	Murina	walstoni	Furey, Csorba & Nguyen Truong Son	2011	Murine de Walston @fr | Walston-Réhrennase @de | Ratonero narizudo de Walston @es	Murina walstoni Furey, Csorba & Nguyen Truong Son, 2011 , “ Cambodia , Ratanakiri Province , Veun Sai District, Veun Sai Protected Forest, 14°01'49”N , 106°45'06"E , elevation 110 m .” Murina walstoni appears to be related to M. swilla and M. aenea based on limited genetic data. Monotypic.	EC & WC Thailand , C & S Laos , S Vietnam (including Phu Quoc I), and SE & SW Cambodia .	Head-body 34:8-45- 4 mm , tail 26-6-32- 5 mm , ear 11-9-14- 7 mm , hindfoot 6- 1-7 mm ,forearm 28-1-34- 7 mm ; weight 4-2-4- 6 g . Fursilky. Dorsal pelage is warm brown (hairs with whitish bases and grayish brown or orangish brown tips); venter is pure white (hairs white from base to tip). Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet. Face is sparsely haired except for long protuberant naked nostrils. Ears are short, broad, and rounded with smoothly convex anterior margins, no notch on posterior margins, and broadly rounded tips; tragus is long and narrow and tapes toward pointed tip. Wing attaches to base of claw on first toe. Skull has non-inflated rostrum and domed braincase; sagittal and lambdoidalcrests are well developed; I? is partly anterior to I%, C' is taller than P* but has smaller basal area; P? is much shorter than P* and has one-half the basal area: mesostyles of M' and M? are moderately developed; and talonids of M, and M, are larger in size than their respective trigonids.	Various forest types from open heavily disturbed forests to pristine forests, lowland mixed deciduous forests, semi-evergreen forests, and dry forests at elevations of 100-400 m .	No information.	A lactating Walston’s Tube-nosed Bat was caught in August in Thailand .	One Walston’s Tube-nosed Bat was collected roosting under a dead banana leaf in Thailand . Calls have steep FM sweeps. In Kamphaeng Phet , Thailand , recordings of one male had start frequencies of 140-153 kHz, end frequencies of 52— 65 kHz, peak frequencies of 117-2-128 kHz, and durations of 1-7-2-4 milliseconds. Similar frequencies were recorded from one female in Loei , Thailand , with start frequencies of 145-149 kHz, end frequencies of 46-49 kHz, peak frequencies of 108-8— 113-7 kHz, and durations of 3-1-4 milliseconds.	Walston’s Tube-nosed Bat reportedly roosts alone.	Not assessed on The IUCN Red List. Walston’s Tube-nosed Bat appears to be relatively widespread, but very little is known aboutits ecology and threats. It is unlikely to be at risk because it occurs in heavily disturbed habitats.	Csorba et al. (2011) | Nguyen Truong Son et al. (2015) | Soisook (2013) | Soisook, Karapan, Satasook & Bates (2013) | Soisook, Thaw Win-Naingng et al. (2017)	https://zenodo.org/record/6398600/files/figure.png	349. Walston’s Tube-nosed Bat Murina walstoni French: Murine de Walston / German: Walston-Réhrennase / Spanish: Ratonero narizudo de Walston Taxonomy. Murina walstoni Furey, Csorba & Nguyen Truong Son, 2011 , “ Cambodia , Ratanakiri Province , Veun Sai District, Veun Sai Protected Forest, 14°01'49”N , 106°45'06"E , elevation 110 m .” Murina walstoni appears to be related to M. swilla and M. aenea based on limited genetic data. Monotypic. Distribution. EC & WC Thailand , C & S Laos , S Vietnam (including Phu Quoc I), and SE & SW Cambodia . Descriptive notes. Head-body 34:8-45- 4 mm , tail 26-6-32- 5 mm , ear 11-9-14- 7 mm , hindfoot 6- 1-7 mm ,forearm 28-1-34- 7 mm ; weight 4-2-4- 6 g . Fursilky. Dorsal pelage is warm brown (hairs with whitish bases and grayish brown or orangish brown tips); venter is pure white (hairs white from base to tip). Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet. Face is sparsely haired except for long protuberant naked nostrils. Ears are short, broad, and rounded with smoothly convex anterior margins, no notch on posterior margins, and broadly rounded tips; tragus is long and narrow and tapes toward pointed tip. Wing attaches to base of claw on first toe. Skull has non-inflated rostrum and domed braincase; sagittal and lambdoidalcrests are well developed; I? is partly anterior to I%, C' is taller than P* but has smaller basal area; P? is much shorter than P* and has one-half the basal area: mesostyles of M' and M? are moderately developed; and talonids of M, and M, are larger in size than their respective trigonids. Habitat. Various forest types from open heavily disturbed forests to pristine forests, lowland mixed deciduous forests, semi-evergreen forests, and dry forests at elevations of 100-400 m . Food and Feeding. No information. Breeding. A lactating Walston’s Tube-nosed Bat was caught in August in Thailand . Activity patterns. One Walston’s Tube-nosed Bat was collected roosting under a dead banana leaf in Thailand . Calls have steep FM sweeps. In Kamphaeng Phet , Thailand , recordings of one male had start frequencies of 140-153 kHz, end frequencies of 52— 65 kHz, peak frequencies of 117-2-128 kHz, and durations of 1-7-2-4 milliseconds. Similar frequencies were recorded from one female in Loei , Thailand , with start frequencies of 145-149 kHz, end frequencies of 46-49 kHz, peak frequencies of 108-8— 113-7 kHz, and durations of 3-1-4 milliseconds. Movements, Home range and Social organization. Walston’s Tube-nosed Bat reportedly roosts alone. Status and Conservation. Not assessed on The IUCN Red List. Walston’s Tube-nosed Bat appears to be relatively widespread, but very little is known aboutits ecology and threats. It is unlikely to be at risk because it occurs in heavily disturbed habitats. Bibliography. Csorba et al. (2011), Nguyen Truong Son et al. (2015), Soisook (2013), Soisook, Karapan, Satasook & Bates (2013), Soisook, Thaw Win-Naingng et al. (2017).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Murina walstoni	Murina		walstoni	Furey, Csorba, & Son	2011	0	J. Mammal.	92(4): 900	Walston&apos;s Tube-nosed Bat	None.	Cambodia, Ratanakiri Province, Veun Sai District, Veun Sai Protected Forest	Cambodia, Vietnam.	Not listed.	Data Deficient		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Murina walstoni	23	Walston's Tube-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MURININAE	NA	Murina	NA	walstoni	Furey, Csorba, & Nguyen Truong Son in Csorba, Nguyen Truong Son, Ith Saveng, & Furey	2011	0	Murina_walstoni	Csorba, G., Son, N. T., Saveng, I., & Furey, N. M. (2011). Revealing cryptic bat diversity: three new Murina and redescription of M. tubinaris from Southeast Asia. Journal of Mammalogy, 92(4), 900.	https://academic.oup.com/jmammal/article/92/4/891/890104	HNHM 2010.20.1		"Cambodia, Ratanakiri Province, Veun Sai District, Veun Sai Protected Forest, 14°01'49"N, 106°45'06"E, elevation 110 m."	14.03	106.75	walstoni Furey, Csorba, & Nguyen Truong Son in Csorba, Nguyen Truong Son, Ith Saveng, & Furey, 2011	recently described	Csorba, G., Son, N. T., Saveng, I., & Furey, N. M. (2011). Revealing cryptic bat diversity: three new Murina and redescription of M. tubinaris from Southeast Asia. Journal of Mammalogy, 92(4), 891-904.	Thailand|Laos|Vietnam|Cambodia	Asia	Indomalaya	DD	0	0	0	Murina_walstoni	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Murina walstoni	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Murina	walstoni	Furey, Csorba &; Son, 2011	<span lang="EN-US">The taxon was described in 2011 based on specimens from Cambodia and Viet Nam.	80000000	Murina walstoni	Data Deficient		2020	2016-08-21 00:00:00 UTC	3.1	English	This species is listed as Data Deficient as it is known only from a few specimens and very little information is available on its distribution, abundance, ecology or threats.	The holotype (Veun Sai Protected Forest, Ratanakiri, Cambodia) was caught 1 hour after sunset in a harp trap set on a dry streambed in semievergreen forest at 110 m asl. The specimen from Botum Sakor National Park (Cambodia) was caught in a harp trap in an area of heavily disturbed lowland forest. The specimen from Yok Don National Park (Viet Nam) was similarly caught in a harp trap in deciduous forest at an elevation of 180 m near the Thac Phat-Buddha waterfall (Csorba et al. 2011). Most of the specimens captured in Laos were in the extreme south, in seasonally dry, semi-deciduous forests, or in sparse, disturbed forests in the lowlands. Some individuals were also captured in heavily disturbed, relatively open seasonally dry forest north of Vientiane, Lao PDR. These and other records all suggest M. walstoni uses more open, drier and can tolerate more disturbed habitats than most other SE Asian Murina species (Francis and Eger 2012, Csorba, pers. comm.).	<p><span lang="EN-US">This species was captured in forests, therefore logging of forests may constitute a threat.<span lang="EN-US"></span></p>	The species was found to be the commonest Murina species in a dry dipterocarp forest in Cambodia.	Unknown	The species was described based on specimens from Veun Sai Protected Forest (Ratanakiri, Cambodia), Botum Sakor National Park (Koh Kong, Cambodia) and Yok Don National Park (Dak Lak, Vietnam). Francis and Eger (2012) reported further specimens from Lao PDR (Attapu, Champassak and Vientiane states) and also from Dong Nai, Viet Nam. Kruskop (2013) reported the species from Cat Tien (Dong Nai, Vietnam), whereas Son et al. (2015) published Vietnamese occurrence data from Nui Chua National Park (Ninh Thuan), Vinh Cuu Nature Reserve (Dong Nai) and Phu Quoc National Park (Kien Giang). Unpublished records also exist for the Loi Hai commune area immediately west of Nui Chua National Park (Ninh Thuan, Vietnam) and for Preah Vihear and Kampong Thom province in Cambodia (Furey, pers. comm.).		Terrestrial		Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Murina		walstoni	Furey, Csorba, & Son	2011	0	J. Mammal.	92(4): 900	Walston&apos;s Tube-nosed Bat	None.	Cambodia, Ratanakiri Province, Veun Sai District, Veun Sai Protected Forest	Cambodia, Vietnam.	Not listed.	Data Deficient		Murina walstoni	1005356	23	Walston's Tube-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MURININAE	NA	Murina	NA	walstoni	Furey, Csorba, & Nguyen Truong Son in Csorba, Nguyen Truong Son, Ith Saveng, & Furey	2011	0	Murina_walstoni	Csorba, G., Son, N. T., Saveng, I., & Furey, N. M. (2011). Revealing cryptic bat diversity: three new Murina and redescription of M. tubinaris from Southeast Asia. Journal of Mammalogy, 92(4), 900.	https://academic.oup.com/jmammal/article/92/4/891/890104	HNHM 2010.20.1		"Cambodia, Ratanakiri Province, Veun Sai District, Veun Sai Protected Forest, 14°01'49"N, 106°45'06"E, elevation 110 m."	14.03028	106.7517	walstoni Furey, Csorba, & Nguyen Truong Son in Csorba, Nguyen Truong Son, Ith Saveng, & Furey, 2011	recently described	Csorba, G., Son, N. T., Saveng, I., & Furey, N. M. (2011). Revealing cryptic bat diversity: three new Murina and redescription of M. tubinaris from Southeast Asia. Journal of Mammalogy, 92(4), 891-904.				Thailand|Laos|Vietnam|Cambodia	Asia	Indomalaya	DD	0	0	0	Murina_walstoni	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Murina_walstoni	1005356	23	Walston's Tube-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Murininae	NA	Murina	NA	walstoni	Furey, Csorba, & Nguyen Truong Son in Csorba, Nguyen Truong Son, Saveng, & Furey	0	Murina walstoni	Csorba, G., Son, N.T., Saveng, I. and Furey, N.M. 2011-08-16. Revealing cryptic bat diversity: three new _Murina_ and redescription of _M. tubinaris_ from Southeast Asia. Journal of Mammalogy 92(4):891-904.	https://doi.org/10.1644/10-MAMM-A-269.1	HNHM 2010.20.1	holotype		"Cambodia, Ratanakiri Province, Veun Sai District, Veun Sai Protected Forest, 14°01'49"N, 106°45'06"E, elevation 110 m."	14.03028	106.7517	recently described	Csorba, G., Son, N. T., Saveng, I., & Furey, N. M. (2011). Revealing cryptic bat diversity: three new Murina and redescription of M. tubinaris from Southeast Asia. Journal of Mammalogy, 92(4), 891-904.		Dahal, D. R., GyÅ‘rössy, D., Thapa, S., Csorba, G., Görföl, T., Pandey, K., ... & Singh, N. B. (2025). First record of Murina walstoni (Chiroptera: Vespertilionidae) outside Southeast Asia. Mammalia, (0).		Thailand|Laos|Vietnam|Cambodia|Nepal	Asia	Indomalaya	DD	0	0	0	Murina_walstoni	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Murina		walstoni	Furey, Csorba & Son	2011	0	J. Mammal.	92(4): 900	Walston&apos;s Tube-nosed Bat	None.	Cambodia, Ratanakiri Province, Veun Sai District, Veun Sai Protected Forest	Cambodia, Vietnam.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84562267/84562270/' target='_blank'>Data Deficient</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Murina walstoni; Murina walstoni; Murina walstoni; Murina walstoni; Murina walstoni; walstoni; Murine de Walston; Walston-Réhrennase; Ratonero narizudo de Walston; Walston's Tube-nosed Bat; Walston&apos;s Tube-nosed Bat; M. walstoni