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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L794	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Murina tubinaris	Murina tubinaris	Murina tubinaris	Murina tubinaris	Murina tubinaris	Murina tubinaris	Murina tubinaris	Murina tubinaris	Murina tubinaris	Murina tubinaris	Murina tubinaris	Murina tubinaris	Murina tubinaris	Murina tubinaris	Murina tubinaris		[MSW2] Subgenus Murina. Listed as a subspecies of huttoni by Ellerman and Morrison-Scott (1951:186); but see Hill (1963o:49, 50).; [MSW3] Subgenus Murina. Listed as a subspecies of huttoni by Ellerman and Morrison-Scott (1951), but apparently distinct, see Hill (1963a, 1983), Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b). Koopman and Danforth (1989) suggested that tubinaris may be conspecific with suilla, but this has not been supported by recent authors.; [HMW] Harpiocephalus tubinaris Scully, 1881 , “Gilgit,” Kashmir, Pakistan . Murina tubinaris previously included populations that are now attributed to M. feae (including M. cineracea as a synonym) in Indochina, M. beelzebub in central Annamite Mountains, and M. jaintianainJaintia Hills of India and Chin Hills of Myanmar . Genetic and morphological data restricted M. tubinaris to north-western India and Pakistan . Specific status of specimens from West Bengal, Arunachal Pradesh, Mizoram, Sikkim, and some areas of northern Myanmar that were previously included under M. tubinaris and distributional limits between cryptic speciessplit from M. tubinaris remain uncertain. Monotypic.; [batnames2022] Subgenus Murina . Reviewed by Csorba et al. (2011).; [IUCN] Earlier treated either as a subspecies of Murina huttoni (Peters, 1872) (Tate 1941, Ellerman and Morrison-Scott 1951), or a distinct species (Hill 1963, 1964, Corbet and Hill 1992, Koopman 1993, Bates and Harrison 1997, Simmons 2005, Srinivasulu and Srinivasulu 2012). This taxon until recently ranged from Pakistan to Thailand and Vietnam. The populations from Southeast Asia and northeast India have been assigned to M. faea  (Son et al. 2015). M. tubinaris sensu stricto is presently restricted to Pakistan and northwest India (Csorba et al. 2011).; [batnames2023] Subgenus Murina . Reviewed by Csorba et al. (2011).; [batnames2025_1.7] Subgenus Murina. Reviewed by Csorba et al. (2011).														tubinaris	Earlier treated either as a subspecies of Murina huttoni (Peters, 1872) (Tate 1941, Ellerman and Morrison-Scott 1951), or a distinct species (Hill 1963, 1964, Corbet and Hill 1992, Koopman 1993, Bates and Harrison 1997, Simmons 2005, Srinivasulu and Srinivasulu 2012). This taxon until recently ranged from Pakistan to Thailand and Vietnam. The populations from Southeast Asia and northeast India have been assigned to M. faea  (Son et al. 2015). M. tubinaris sensu stricto is presently restricted to Pakistan and northwest India (Csorba et al. 2011).			tubinaris	tubinaris			tubinaris (Scully, 1881)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		Kashmir – Vietnam; ref. 4.106	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Murina tubinaris	Pakistan, Gilgit.	Scully	1881	Proc. Zool. Soc. Lond., 1881:200.	Distribution: Ranging from Pakistan to Vietnam.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		N Pakistan – Vietnam	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Scully	1881	Proc. Zool. Soc. Lond., 1881:200.	Subgenus Murina. Listed as a subspecies of huttoni by Ellerman and Morrison-Scott (1951:186); but see Hill (1963o:49, 50).	Pakistan, N India, Burma, Thailand, Laos, Vietnam.	Pakistan, Gilgit.		SCULLY	1881	Rostrum not particularly massive. Size relatively small (forearm length, 29-35 mm). Canines relatively long.	Distribution: Ranging from Pakistan to Vietnam.	No subspecies. Probably a subspecies of M. suilla.		132	species	M. tubinaris	SCULLY	1881	Murina	subgenus	Murina tubinaris				Rostrum not particularly massive. Size relatively small (forearm length, 29-35 mm). Canines relatively long.	No subspecies. Probably a subspecies of M. suilla.		6. M. tubinaris (SCULLY 1881) [suilla group].	6	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Murininae		Murina tubinaris	Murina	Murina	tubinaris	Scully	y	1881		Proc. Zool. Soc. Lond.	1881		200		Scully's Tube-nosed Bat	Pakistan, Kasmir, Gilgit.	Pakistan, N India, Burma, Thailand, Laos, Vietnam.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).		Subgenus Murina. Listed as a subspecies of huttoni by Ellerman and Morrison-Scott (1951), but apparently distinct, see Hill (1963a, 1983), Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b). Koopman and Danforth (1989) suggested that tubinaris may be conspecific with suilla, but this has not been supported by recent authors.	4C3D87E8FF606AD8FA50918F1DBFB92F	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	916	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF606AD8FA50918F1DBFB92F.xml	Murina tubinaris	Vespertilionidae	Murina	tubinaris		1881	Murine de Scully @fr | Scully-Rohrennase @de | Ratonero narizudo de Scully @es	Harpiocephalus tubinaris Scully, 1881 , “Gilgit,” Kashmir, Pakistan . Murina tubinaris previously included populations that are now attributed to M. feae (including M. cineracea as a synonym) in Indochina, M. beelzebub in central Annamite Mountains, and M. jaintianainJaintia Hills of India and Chin Hills of Myanmar . Genetic and morphological data restricted M. tubinaris to north-western India and Pakistan . Specific status of specimens from West Bengal, Arunachal Pradesh, Mizoram, Sikkim, and some areas of northern Myanmar that were previously included under M. tubinaris and distributional limits between cryptic speciessplit from M. tubinaris remain uncertain. Monotypic.	N Pakistan and NW India ( Jammu and Kashmir , and Himachal Pradesh ).	Head-body 45.7 mm, tail 35- 6 mm , ear 14 mm , hindfoot 9- 4 mm (type specimen), forearm 31-35- 6 mm . Dorsal pelage is light grayish brown (hairs with medium brown bases and light gray band that progressesto light brown before darker brown tips); venter is whitish (hairs with medium brown bases and off-white to tips). Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet. Face is sparsely haired except for long protuberant naked nostrils. Ears are short, broad, and rounded, with smoothly convex anterior margins, small notch on posterior margins, and broadly rounded tips; tragusis long and narrow and tapers toward pointed tip. Wing attaches near base of first toe. Skull is small, with relatively flat braincase and non-inflated rostrum; sagittal crest is absent, and lambdoidal crests are relatively weak; I” is anterior to I’, and they are equal in height; basal area of C! is larger than that of P*, and it is also taller; basal area and height of P? are less than one-half those of P*; mesostyles of M' and M? are reduced; and talonids of M, and M,are equalin size to their respective trigonids.	Montane forests at elevations of ¢. 3000 m in Himachal Pradesh, ¢. 2615 m in Pakistan , and ¢. 2154 m in Kashmir.	No information.	Young Scully’s Tube-nosed Bats were caught in July in Kashmir.	One Scully’s Tube-nosed Bat was found roosting in a tree c¢.1-: 2 m aboveground in Pakistan .	Scully’s Tube-nosed Bat was recorded roosting alone once.	Classified as Least Concern on The IUCN Red List. Distribution of Scully’s Tube-nosed Bat has decreased, and its conservation status needs reassessment.	Bates & Harrison (1997) | Csorba, Bates, Molur & Srinivasulu (2008b) | Csorba, Nguyen Truong Son et al. (2011) | Kruskop & Eger (2008) | Saikia et al. (2011) | Soisook (2013)	https://zenodo.org/record/6398608/files/figure.png	353. Scully’s Tube-nosed Bat Murina tubinaris French: Murine de Scully / German: Scully-Rohrennase / Spanish: Ratonero narizudo de Scully Taxonomy. Harpiocephalus tubinaris Scully, 1881 , “Gilgit,” Kashmir, Pakistan . Murina tubinaris previously included populations that are now attributed to M. feae (including M. cineracea as a synonym) in Indochina, M. beelzebub in central Annamite Mountains, and M. jaintianainJaintia Hills of India and Chin Hills of Myanmar . Genetic and morphological data restricted M. tubinaris to north-western India and Pakistan . Specific status of specimens from West Bengal, Arunachal Pradesh, Mizoram, Sikkim, and some areas of northern Myanmar that were previously included under M. tubinaris and distributional limits between cryptic speciessplit from M. tubinaris remain uncertain. Monotypic. Distribution. N Pakistan and NW India ( Jammu and Kashmir , and Himachal Pradesh ). Descriptive notes. Head-body 45.7 mm, tail 35- 6 mm , ear 14 mm , hindfoot 9- 4 mm (type specimen), forearm 31-35- 6 mm . Dorsal pelage is light grayish brown (hairs with medium brown bases and light gray band that progressesto light brown before darker brown tips); venter is whitish (hairs with medium brown bases and off-white to tips). Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet. Face is sparsely haired except for long protuberant naked nostrils. Ears are short, broad, and rounded, with smoothly convex anterior margins, small notch on posterior margins, and broadly rounded tips; tragusis long and narrow and tapers toward pointed tip. Wing attaches near base of first toe. Skull is small, with relatively flat braincase and non-inflated rostrum; sagittal crest is absent, and lambdoidal crests are relatively weak; I” is anterior to I’, and they are equal in height; basal area of C! is larger than that of P*, and it is also taller; basal area and height of P? are less than one-half those of P*; mesostyles of M' and M? are reduced; and talonids of M, and M,are equalin size to their respective trigonids. Habitat. Montane forests at elevations of ¢. 3000 m in Himachal Pradesh, ¢. 2615 m in Pakistan , and ¢. 2154 m in Kashmir. Food and Feeding. No information. Breeding. Young Scully’s Tube-nosed Bats were caught in July in Kashmir. Activity patterns. One Scully’s Tube-nosed Bat was found roosting in a tree c¢.1-: 2 m aboveground in Pakistan . Movements, Home range and Social organization. Scully’s Tube-nosed Bat was recorded roosting alone once. Status and Conservation. Classified as Least Concern on The IUCN Red List. Distribution of Scully’s Tube-nosed Bat has decreased, and its conservation status needs reassessment. Bibliography. Bates & Harrison (1997), Csorba, Bates, Molur & Srinivasulu (2008b), Csorba, Nguyen Truong Son et al. (2011), Kruskop & Eger (2008), Saikia et al. (2011), Soisook (2013).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Murina tubinaris	Murina	Murina	tubinaris	Scully	1881	1	Proc. Zool. Soc. Lond.	1884:20:00	Scully's Tube-nosed Bat	None.	Pakistan, Kasmir, Gilgit.	NE Pakistan through NW India (Himchal Pradesh)	Not listed.	Data Deficient	Subgenus Murina . Reviewed by Csorba et al. (2011).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Murina tubinaris	23	Scully's Tube-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MURININAE	NA	Murina	NA	tubinaris	Scully	1881	1	Harpiocephalus_tubinaris	Scully, J. (1881). On the mammals of Gilgit. Proceedings of the Zoological Society of London, 1881, 200.	https://www.biodiversitylibrary.org/item/96698#page/268/mode/1up	NA		"Gilgit," Kashmir, Pakistan.			tubinaris (Sully, 1881)	NA	NA	Pakistan|India	Asia	Indomalaya	DD	0	0	0	Murina_tubinaris	0	sciname match	Murina_tubinaris	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Murina tubinaris	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Murina	tubinaris	(Scully, 1881)	Earlier treated either as a subspecies of Murina huttoni (Peters, 1872) (Tate 1941, Ellerman and Morrison-Scott 1951), or a distinct species (Hill 1963, 1964, Corbet and Hill 1992, Koopman 1993, Bates and Harrison 1997, Simmons 2005, Srinivasulu and Srinivasulu 2012). This taxon until recently ranged from Pakistan to Thailand and Vietnam. The populations from Southeast Asia and northeast India have been assigned to M. faea  (Son et al. 2015). M. tubinaris sensu stricto is presently restricted to Pakistan and northwest India (Csorba et al. 2011).	20000000	Murina tubinaris	Data Deficient		2019	2018-08-31 00:00:00 UTC	3.1	English	Listed as Data Deficient as no information on its population, ecology, threats and current distribution is known.	In South Asia, little is known about the habitat or ecology of this species except that this species is found in montane forests on the mountain slopes and roosts among trees (Molur et al. 2002). One or two young are born (Bates and Harrison 1997).	Nothing is known about threats to this species.	Nothing is known about its population.	Unknown	This species is present in northern South Asia, and is known historically from India (Jammu and Kashmir) and Pakistan (North West Frontier Province) (Molur et al. 2002).		Terrestrial	In South Asia, there are no conservation measures in place and this species has not been recorded form any protected areas. Ecological and population studies recommended (Molur et al. 2002). In Southeast Asia, it has been recorded from some protected areas.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Murina	Murina	tubinaris	Scully	1881	1	Proc. Zool. Soc. Lond.	1884:20:00	Scully's Tube-nosed Bat	None.	Pakistan, Kasmir, Gilgit.	NE Pakistan through NW India (Himchal Pradesh)	Not listed.	Data Deficient	Subgenus Murina . Reviewed by Csorba et al. (2011).	Murina tubinaris	1005354	23	Scully's Tube-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MURININAE	NA	Murina	NA	tubinaris	Scully	1881	1	Harpiocephalus_tubinaris	Scully, J. (1881). On the mammals of Gilgit. Proceedings of the Zoological Society of London, 1881, 200.	https://www.biodiversitylibrary.org/item/96698#page/268/mode/1up	NA		"Gilgit," Kashmir, Pakistan.			tubinaris (Sully, 1881)	NA	NA				Pakistan|India	Asia	Indomalaya	DD	0	0	0	Murina_tubinaris	0	sciname match	Murina_tubinaris	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Murina_tubinaris	1005354	23	Scully's Tube-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Murininae	NA	Murina	NA	tubinaris	Scully	1	Harpiocephalus tubinaris	Scully, J. 1881-06. On the mammals of Gilgit. Proceedings of the Zoological Society of London 1881(1):197-209.	https://www.biodiversitylibrary.org/page/30759333	lost (number not known)	holotype		"Gilgit," Kashmir, Pakistan.			NA	NA				Pakistan|India	Asia	Indomalaya	DD	0	0	0	Murina_tubinaris	0	sciname match	Murina_tubinaris	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Murina	Murina	tubinaris	Scully	1881	1	Proc. Zool. Soc. Lond.	1884:20:00	Scully's Tube-nosed Bat	None.	Pakistan, Kasmir, Gilgit.	NE Pakistan through NW India (Himchal Pradesh)	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84560827/22096188/' target='_blank'>Data Deficient</a>	Subgenus Murina. Reviewed by Csorba et al. (2011).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Murina tubinaris; Murina tubinaris; Murina tubinaris; Murina tubinaris; Murina tubinaris; Murina tubinaris; tubinaris; Murine de Scully; Scully-Rohrennase; Ratonero narizudo de Scully; Scully's Tube-nosed Bat; Scully's Tube-nosed Bat; Scully's Tube-nosed Bat; M. tubinaris