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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L783	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	Murina puta	Murina puta	Murina puta	Murina puta	Murina puta	Murina puta	Murina puta	Murina puta	Murina puta	Murina puta	Murina puta	Murina puta	Murina puta	Murina puta		[MSW2] Subgenus Murina. Closely related to and possibly conspecific with M. huttoni, see Yoshiyuki (1989:215).; [MSW3] Subgenus Murina. Closely related to and possibly conspecific with huttoni, see Yoshiyuki (1989).; [HMW] Murina puta Kishida, 1924 , Erh-Shui, Chang Hua, Taiwan . See M. harrisoni . Most closely related to M. huttonii , and some authors have suggested that they may be conspecific. Monotypic.; [batnames2022] Subgenus Murina . Closely related to and possibly conspecific with huttoni , see Yoshiyuki (1989).; [IUCN] The taxonomic status of this species is uncertain. It may be conspecific with M. huttoni , to which it is closely related (Yoshiyuki 1989).; [batnames2023] Subgenus Murina . Closely related to and possibly conspecific with huttoni , see Yoshiyuki (1989).; [batnames2025_1.7] Subgenus Murina. Closely related to and possibly conspecific with huttoni, see Yoshiyuki (1989).														puta	The taxonomic status of this species is uncertain. It may be conspecific with M. huttoni , to which it is closely related (Yoshiyuki 1989).				puta			puta Kishida, 1924					Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Murina puta	Taiwan, Chang Hua, Erh-Shui.	Kishida	1924	Zool. Mag. (Tokyo), 36:130-133.	Distribution: Confined to Taiwan.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		Taiwan; status uncertain	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Kishida	1924	Zool. Mag. (Tokyo), 36:130-133.	Subgenus Murina. Closely related to and possibly conspecific with M. huttoni, see Yoshiyuki (1989:215).	Taiwan.	Taiwan, Chang Hua, Erh-Shui.		KISHIDA	1924	Size medium (forearm length, 35-36mm). Ears rounded. Talonids of anterior and middle lower molars and their cusps well developed. Upper canine relatively long. Teeth considerably enlarged.	Distribution: Confined to Taiwan.	No subspecies, but may be a subspecies of M. huttoni.		132	species	M. puta	KISHIDA	1924	Murina	subgenus	Murina puta				Size medium (forearm length, 35-36mm). Ears rounded. Talonids of anterior and middle lower molars and their cusps well developed. Upper canine relatively long. Teeth considerably enlarged.	No subspecies, but may be a subspecies of M. huttoni.		10. M. puta KISHIDA 1924 [cyclotis group],	10	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Murininae		Murina puta	Murina	Murina	puta	Kishida		1924		Zool. Mag. (Tokyo)	36		127		Taiwanese Tube-nosed Bat	Taiwan, Chang Hua, Erh-Shui.	Taiwan.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		Subgenus Murina. Closely related to and possibly conspecific with huttoni, see Yoshiyuki (1989).	4C3D87E8FF636ADCFA869E3716FFB6BC	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	913	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF636ADCFA869E3716FFB6BC.xml	Murina puta	Vespertilionidae	Murina	puta	Kishida	1924	Murine de Taiwan @fr | Taiwan-Rohrennase @de | Ratonero narizudo de Taiwan @es | Taiwanese Tube-nosed Bat @en	Murina puta Kishida, 1924 , Erh-Shui, Chang Hua, Taiwan . See M. harrisoni . Most closely related to M. huttonii , and some authors have suggested that they may be conspecific. Monotypic.	Taiwan .	Head-body 59-61 mm , tail 32-36 mm , ear 17-19 mm , hindfoot 10-11 mm , forearm 33-39 mm . Pelage is long and flufty; dorsally medium brown or sometimes reddish to grayish brown (hairs with dark bases, tan middle band, and reddish tips); ventrally paler. Dorsal pelage extendssparsely on to wings, uropatagium, thumbs, and feet. Face is sparsely haired except long protuberant nostrils, which are naked. Ears are short, broad, and rounded, with smoothly convex anterior margins, no notch on posterior margin, with a broadly rounded tip; tragus is long, narrow, and tapering toward pointed tip. Wing attachesto base of claw on first toe. Skull is robust and broad. I? is anterior to I% basal area of C! is equal to or larger than that of P*; P* is more than two-thirds the height and basal area of P*.	Restricted to temperate montane forests.	No information.	No information.	No information.	No information.	Classified as Near Threatened on The IUCN Red List. The Taiwan Tube-nosed Bat has a relatively restricted range. It may be affected by deforestation, although this is no longer carried outto a significant degree on the higher elevations of Taiwan . The species has been regionally redlisted in China as endangered.	Csorba et al. (2007) | Kuo Haochih et al. (2009) | Lin Liangkong et al. (1997) | Ruedi, Biswas & Csorba (2012) | Smith & Xie Yan (2008) | Smith, Johnston, Jones & Rossiter (2008e)	https://zenodo.org/record/6398596/files/figure.png	347. Taiwan Tube-nosed Bat Murina puta French: Murine de Taiwan / German: Taiwan-Rohrennase / Spanish: Ratonero narizudo de Taiwan Other common names: Taiwanese Tube-nosed Bat Taxonomy. Murina puta Kishida, 1924 , Erh-Shui, Chang Hua, Taiwan . See M. harrisoni . Most closely related to M. huttonii , and some authors have suggested that they may be conspecific. Monotypic. Distribution. Taiwan . Descriptive notes. Head-body 59-61 mm , tail 32-36 mm , ear 17-19 mm , hindfoot 10-11 mm , forearm 33-39 mm . Pelage is long and flufty; dorsally medium brown or sometimes reddish to grayish brown (hairs with dark bases, tan middle band, and reddish tips); ventrally paler. Dorsal pelage extendssparsely on to wings, uropatagium, thumbs, and feet. Face is sparsely haired except long protuberant nostrils, which are naked. Ears are short, broad, and rounded, with smoothly convex anterior margins, no notch on posterior margin, with a broadly rounded tip; tragus is long, narrow, and tapering toward pointed tip. Wing attachesto base of claw on first toe. Skull is robust and broad. I? is anterior to I% basal area of C! is equal to or larger than that of P*; P* is more than two-thirds the height and basal area of P*. Habitat. Restricted to temperate montane forests. Food and Feeding. No information. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Classified as Near Threatened on The IUCN Red List. The Taiwan Tube-nosed Bat has a relatively restricted range. It may be affected by deforestation, although this is no longer carried outto a significant degree on the higher elevations of Taiwan . The species has been regionally redlisted in China as endangered. Bibliography. Csorba et al. (2007), Kuo Haochih et al. (2009), Lin Liangkong et al. (1997), Ruedi, Biswas & Csorba (2012), Smith & Xie Yan (2008), Smith, Johnston, Jones & Rossiter (2008e).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Murina puta	Murina	Murina	puta	Kishida	1924	0	Zool. Mag. (Tokyo)	38:07:00	Taiwanese Tube-nosed Bat	None.	Taiwan, Chang Hua, Erh-Shui.	Taiwan.	Not listed.	Least Concern	Subgenus Murina . Closely related to and possibly conspecific with huttoni , see Yoshiyuki (1989).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Murina puta	23	Taiwan Tube-nosed Bat	Taiwanese Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MURININAE	NA	Murina	NA	puta	Kishida	1924	0						Erh-Shui, Chang Hua, Taiwan.			puta Kishida, 1924	NA	NA	Taiwan	Asia	Palearctic	LC	0	0	0	Murina_puta	0	sciname match	Murina_puta	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	13944	Murina puta	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Murina	puta	Kishida, 1924	The taxonomic status of this species is uncertain. It may be conspecific with M. huttoni , to which it is closely related (Yoshiyuki 1989).	20000000	Murina puta	Least Concern		2019	2018-08-27 00:00:00 UTC	3.1	English	This species is listed as Least Concerned as it has a stable population and its EOO ;(40,251 km<sup>2</sup>) ;and AOO are too great to qualify under any criteria. It is suspected to be restricted to primary temperate forest in the mountainous interior of Taiwan, and there has been deforestation within its suspected distribution range in the past (Smith et al . 2008). However, all recent studies indicate it is one of the most common species across forested habitats and various plantations in the mountain areas in the island. Road-kill incidence has been reported in the species, suggesting the distribution of the species is fragmented.	This species is commonly found in boreal and mixed forests, temperate forest, bamboo and tree plantations, and abandoned orchard in mountain areas in Taiwan. It flies along trails, small logging road and edge of forests, and above small streams with dense canopy cover. It flies mostly in lower heights from near the ground to less than 1.5 m (Lee et al. 2007, Lu 2013, Y.-F. Lee, unpublished data). It is often found roosting on tree trunk, segments of dead stem of Oldhams' bamboo (Bambusa oldhamii ), dead leaves of shell ginger (Alpinia zerumbet ), Flying Spider-monkey tree fern (Cyathea lepifera ), rice paper plant (Tetrapanax papyrifer ), Japanese Cider (Cryptomeria japonica ), long-leaf Chinkapin (Castanopsis carlesii ), upright elephant ear (Alocasia odora ), plantain (Musa uranoscopos ) within forest, both dead and furled leaves of banana (Musa spp. ) in plantations, and bat boxes (Chen 2011, Cheng et al . 2015, Chou et al . 2008, Chu et al . 2008, Ho et al . 2016). Individuals have been observed switching roosts at a distance of c. 100 m every 1-4 days (Chu et al. 2008, Chen 2011, Cheng et al. 2015). Insectivorous, its diet includes coleopterans, hemipterans, lepidopterans, neuropterans, dipterans, orthopterans, hymenopterans, and spiders (Lee et al. 2005, Lu 2013). A recent study suggests it uses both echolocation and chemical cues to detect prey and is capable of obtaining insects directly from object surface (Liao 2013). Females have been observed in pregnancy in April and May (Cheng et al . 2015), and lactating in May and June (Kuo, H.-C., Chen, S.-F., and Rossiter, S. personal communication).	Deforestation has been suspected as a potential threat to this species since it was suspected to be restricted to temperate forests in high elevation of mountainous interior, where logging was carried out. Recent studies found this species are in nature, disturbed forests and various types of plantation from low to high elevations in the mountain area (Kuo et al . 2014, Cheng et al . 2015) and suggest the species is common in the island. Road-kill incidence of this species has been reported more often than other bat species recently (Taiwan Roadkill Observation Network). Hence, the development of road systems within its distribution range is suspiciously increasing the isolation among subpopulations.	This species is common and wide distributed in throughout the mountain areas with elevation lower than 2,600 m asl in Taiwan (Kuo et al. 2014, Cheng et al. 2015). The population is considered stable, but recent road-kill incidence surveys suggest the subpopulations might be fragmented by the well-developed road systems in some of its distribution range.	Stable	This species is endemic to Taiwan (Kuo 2004) and widely distributed from low to middle high elevation in hill and mountain areas on the island (Kuo et al . 2014).		Terrestrial	There are no known conservation measures in place for this species. This species occurs in several protected areas, including the Kenting National Park, Yangmingshan National park, Shei-Pa National Park, Taroko National Park, and Yushan National Park (Lee et al. 2007, Lee and Huang 2007, Ho et al. 2016, Cheng, H.-S. and Chou, C.-H. unpublished data). Research is needed in the fields of population status, biology and ecology, habitat status and threats.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Murina	Murina	puta	Kishida	1924	0	Zool. Mag. (Tokyo)	38:07:00	Taiwanese Tube-nosed Bat	None.	Taiwan, Chang Hua, Erh-Shui.	Taiwan.	Not listed.	Least Concern	Subgenus Murina . Closely related to and possibly conspecific with huttoni , see Yoshiyuki (1989).	Murina puta																																																0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Murina_puta	1005346	23	Taiwan Tube-nosed Bat	Taiwanese Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Murininae	NA	Murina	NA	puta	Kishida	0	Murina puta	Kishida, K. 1924. [Title in Japanese]. Zoological Magazine, Tokyo 36:127-139.	https://hdl.handle.net/2027/uc1.31210014193187?urlappend=%3Bseq=156%3Bownerid=117225479-453	untraced (number not known)	holotype		Erh-Shui, Chang Hua, Taiwan.			NA	NA				Taiwan	Asia	Palearctic	LC	0	0	0	Murina_puta	0	sciname match	Murina_puta	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Murina	Murina	puta	Kishida	1924	0	Zool. Mag. (Tokyo)	38:07:00	Taiwanese Tube-nosed Bat	None.	Taiwan, Chang Hua, Erh-Shui.	Taiwan.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/13944/22093018/' target='_blank'>Least Concern</a>	Subgenus Murina. Closely related to and possibly conspecific with huttoni, see Yoshiyuki (1989).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Murina puta; Murina puta; Murina puta; Murina puta; Murina puta; Murina puta; puta; Murine de Taiwan; Taiwan-Rohrennase; Ratonero narizudo de Taiwan; Taiwanese Tube-nosed Bat; Taiwan Tube-nosed Bat; Taiwanese Tube-nosed Bat; Taiwanese Tube-nosed Bat; Taiwanese Tube-nosed Bat; M. puta