http://www.w3.org/ns/prov#wasDerivedFrom	http://purl.org/dc/elements/1.1/format	name_CH1_1980	name_MSW1_1982	name_CH3_1991	name_MSW2_1993	name_Koopman_1994	name_MSW3_2005	name_HMW_2019	name_BatNames_2022	name_MDD_2022	name_IUCN_2022	name_BatNames_2023	name_MDD_2023	name_MDD_2025_2.0	name_batnames_2025_1.7	name_MDD_2025_2.2	column151	taxonomic_notes_concatenated	column171	synonyms_CH1	subspecies__MSW2	synonyms__MSW1	synonyms_CH3	synonyms_MSW2	subspecies_Koopman94_interpreted	subspecies_MSW3_interpreted	synonym_MSW3_interpreted	subspecies_HMW_interpreted	synonym_HMW_interpreted	subspecies_batnames_interpreted	synonym_batnames_interpreted	synonym_MDD_interpreted	synonym_IUCN_interpreted	subspecies_batnames2023_interpreted	synonym_batnames2023_interpreted	synonym_MDD2023_interpreted	synonym_MDD2025_interpreted	subspecies_batnames2025_interpreted	synonyms_batnames2025_interpreted	nominalNames	column391	docOrigin_CH1	commonName_CH1	distribution_CH1	docOrigin_MSW1	column451	typeLocality_MSW1	authority_MSW1	year_MSW1	citation_MSW1	distribution	comment_MSW1	docOrigin_CH3	commonName_CH3	distribution_CH3	docOrigin_MSW2	authority_MSW2	year_MSW2	citation_MSW2	comments_MSW2	distribution_MSW2	typeLocality_MSW2	docOrigin_Koopman94	authority_Koopman94	year_Koopman94	description_Koopman94	distribution_Koopman94	diversity_Koopman94	subspecies_Koopman94	page	rank	name	authority	year	parent	parent_rank	corrected_name	actual_species_count	claimed_species_count	dental_formula	description	diversity	full_subspecies_text	name_line	species_index	subspecies	synonym	text	docOrigin_MSW3	order_MSW3	family_MSW3	subfamily_MSW3	tribe_MSW3	name_MSW3	genus_MSW3	subgenus_MSW3	species_MSW3	authoritySpeciesAuthor_MSW3	(parentheses (1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common Name_batnames	synonyms_batnames	type_locality_batnames	Distribution_batnames	CITES_batnames	IUCN_batnames	comments_batnames	docOrigin_MDD	name_MDD	phylosort_MDD	mainCommonName_MDD	otherCommonNames_MDD	subclass_MDD	infraclass_MDD	magnorder_MDD	superorder_MDD	order_MDD	suborder_MDD	infraorder_MDD	parvorder_MDD	superfamily_MDD	family_MDD	subfamily_MDD	tribe_MDD	genus_MDD	subgenus_MDD	specificEpithet_MDD	authoritySpeciesAuthor_MDD	authoritySpeciesYear_MDD	authorityParentheses_MDD	originalNameCombination_MDD	authoritySpeciesCitation_MDD	authoritySpeciesLink_MDD	holotypeVoucher_MDD	holotypeVoucherURIs_MDD	typeLocality_MDD	typeLocalityLatitude_MDD	typeLocalityLongitude_MDD	nominalNames_MDD	taxonomyNotes_MDD	taxonomyNotesCitation_MDD	countryDistribution_MDD	continentDistribution_MDD	biogeographicRealm_MDD	iucnStatus_MDD	extinct_MDD	domestic_MDD	flagged_MDD	CMW_sciName_MDD	diffSinceCMW_MDD	MSW3_matchtype_MDD	MSW3_sciName_MDD	diffSinceMSW3_MDD	docOrigin_IUCN	internalTaxonId_IUCN	NAME_IUCN	kingdomName_IUCN	phylumName_IUCN	className_IUCN	orderName_IUCN	familyName_IUCN	genusName_IUCN	speciesName_IUCN	authoritySpeciesAuthorYear_IUCN	taxonomicNotes_IUCN	assessmentId_IUCN	scientificName_IUCN	redlistCategory_IUCN	redlistCriteria_IUCN	yearPublished_IUCN	assessmentDate_IUCN	criteriaVersion_IUCN	language_IUCN	rationale_IUCN	habitat_IUCN	threats_IUCN	population_IUCN	populationTrend_IUCN	range_IUCN	useTrade_IUCN	systems_IUCN	conservationActions_IUCN	realm_IUCN	yearLastSeen_IUCN	possiblyExtinct_IUCN	possiblyExtinctInTheWild_IUCN	scopes_IUCN	docOrigin_batnames2023	FAMILY_batnames2023	GENUS_batnames2023	SUBGENUS_batnames2023	SPECIES_batnames2023	authoritySpeciesAuthor_batnames2023	authoritySpeciesYearbatnames2023	PARENTHESES_batnames2023 (1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name MDD2023	id_MDD2023	phylosort_MDD2023	mainCommonName_MDD2023	otherCommonNames_MDD2023	subclass_MDD2023	infraclass_MDD2023	magnorder_MDD2023	superorder_MDD2023	order_MDD2023	suborder_MDD2023	infraorder_MDD2023	parvorder_MDD2023	superfamily_MDD2023	Family_mdd2023	subfamily_MDD2023	tribe_MDD2023	genus_MDD2023	subgenus_MDD2023	specificEpithet_MDD2023	authoritySpeciesAuthor_MDD2023	authoritySpeciesYear_MDD2023	authorityParentheses_MDD2023	originalNameCombination_MDD2023	authoritySpeciesCitation_MDD2023	authoritySpeciesLink_MDD2023	holotypeVoucher_MDD2023	holotypeVoucherURIs_MDD2023	typeLocality_MDD2023	typeLocalityLatitude_MDD2023	typeLocalityLongitude_MDD2023	nominalNames_MDD2023	taxonomyNotes_MDD2023	taxonomyNotesCitation_MDD2023	distributionNotes_MDD2023	distributionNotesCitation_MDD2023	subregionDistribution_MDD2023	countryDistribution_MDD2023	continentDistribution_MDD2023	biogeographicRealm_MDD2023	iucnStatus_MDD2023	extinct_MDD2023	domestic_MDD2023	flagged_MDD2023	CMW_sciName_MDD2023	diffSinceCMW_MDD2023	MSW3_matchtype_MDD2023	MSW3_sciName_MDD2023	diffSinceMSW3_MDD2023	docOrigin_MDD2025	sciName	id	phylosort	mainCommonName	otherCommonNames	subclass	infraclass	magnorder	superorder	order	suborder	infraorder	parvorder	superfamily	family	subfamily	tribe	genus	subgenus	specificEpithet	authoritySpeciesAuthor	authorityParentheses	originalNameCombination	authoritySpeciesCitation	authoritySpeciesLink	typeVoucher	typeKind	typeVoucherURIs	typeLocality	typeLocalityLatitude	typeLocalityLongitude	taxonomyNotes	taxonomyNotesCitation	distributionNotes	distributionNotesCitation	subregionDistribution	countryDistribution	continentDistribution	biogeographicRealm	iucnStatus	extinct	domestic	flagged	CMW_sciName	diffSinceCMW	MSW3_matchtype	MSW3_sciName	diffSinceMSW3	docOrigin_batnames2025	Family	Genus	Subgenus	Species	Author	Date	Parentheses (1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L770	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	Murina leucogaster [synonym of]	Murina leucogaster hilgendorfi	Murina hilgendorfi	Murina hilgendorfi	Murina hilgendorfi	Murina hilgendorfi	Murina hilgendorfi	Murina hilgendorfi	Murina hilgendorfi	Murina hilgendorfi	Murina hilgendorfi	Murina hilgendorfi		[MSW3] Subgenus Murina. Formerly included in leucogaster, but apparently distinct. May include more than one species; see Yoshiyuki (1989). Also see Wang (1959).; [HMW] Harpyiocephalus hilgendorfi Peters, 1880 , Yedo, near Tokyo , Japan . Relationships uncertain but limited genetic data place this species as sister to the M. leucogaster clade (see above). Often considered a race of M. leucogaster but numerous genetic and morphological studies support their treatment at separate species. Race sibirica has occasionally been treated as a distinct species, but morphological data do not support this. The name intermedia is here considered to be a synonym of ognevi , although further morphological and genetic comparison is needed. Three subspecies recognized.; [batnames2022] Subgenus Murina . Formerly included in leucogaster , but apparently distinct. May include more than one species; see Yoshiyuki(1989). Also see Wang (1959).; [IUCN] Muirna hilgendorfi was once synonymized with the related M. leucogaster (Bobrinskyi et al. 1965), and this point ov view was accepted until recently (e.g. Koopman 1994, Botvinkin 2002). Yoshiyuki considered M. hilgendorfi from Japan a distinct species (Yoshiyuki 1989), but erroneously divided it from all mainland populations. Currently species status of M. hilgendorfi is accepted (Simmons 2005) and got some support from molecular data (Kruskop et al. 2012). Relationship with poorly known M. fusca from North-East China require further studies.; [batnames2023] Subgenus Murina . Formerly included in leucogaster , but apparently distinct. May include more than one species; see Yoshiyuki(1989). Also see Wang (1959).; [batnames2025_1.7] Subgenus Murina. Formerly included in leucogaster, but apparently distinct. May include more than one species; see Yoshiyuki(1989). Also see Wang (1959).									intermedia, ognevi, sibirica	hilgendorfi, ognevi, sibirica	ognevi - intermedia?	hilgendorfi	hilgendorfi - intermedia, ognevi, sibirica	hilgendorfi, sibirca, ognevi, intermedia	Muirna hilgendorfi was once synonymized with the related M. leucogaster (Bobrinskyi et al. 1965), and this point ov view was accepted until recently (e.g. Koopman 1994, Botvinkin 2002). Yoshiyuki considered M. hilgendorfi from Japan a distinct species (Yoshiyuki 1989), but erroneously divided it from all mainland populations. Currently species status of M. hilgendorfi is accepted (Simmons 2005) and got some support from molecular data (Kruskop et al. 2012). Relationship with poorly known M. fusca from North-East China require further studies.	hilgendorfi	hilgendorfi - intermedia, ognevi, sibirica	hilgendorfi, sibirca, ognevi, intermedia	hilgendorfi, sibirca, ognevi, intermedia	hilgendorfi 	hilgendorfi - intermedia, ognevi, sibirica	hilgendorfi (W. C. H. Peters, 1880)|sibirica (Kastschenko, 1905)|ognevi Bianchi, 1916|intermedia Mori, 1933						N/A																																								NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Murininae		Murina hilgendorfi	Murina	Murina	hilgendorfi	Peters		1880		Monatsb. K. Preuss. Akad. Wiss. Berlin	1880		24		Hilgendorf's Tube-nosed Bat	Japan, near Tokyo, Yedo.	N China; Upper Yenisei River (Russia); Altai Mtns (Russia, Kazakhstan and Mongolia); Korea; Ussur region (Russia); Sakhalin Isl (Russia); Honshu, Kyushu and Shikiku (Japan).	IUCN 2003 – Not evaluated; not considered in IUCN/SSC Action Plan (2001).	intermedia Mori, 1933; ognevi Bianchi, 1916; sibirica Kastschenko, 1905.	Subgenus Murina. Formerly included in leucogaster, but apparently distinct. May include more than one species; see Yoshiyuki (1989). Also see Wang (1959).	4C3D87E8FF6F6AD1FA91909E1B63BDB7	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	909	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF6F6AD1FA91909E1B63BDB7.xml	Murina hilgendorfi	Vespertilionidae	Murina	hilgendorfi	Peters	1880	Murine de Hilgendorf @fr | Hilgendorf-Rohrennase @de | Ratonero narizudo de Hilgendorf @es | Greater Tube-nosed Bat @en	Harpyiocephalus hilgendorfi Peters, 1880 , Yedo, near Tokyo , Japan . Relationships uncertain but limited genetic data place this species as sister to the M. leucogaster clade (see above). Often considered a race of M. leucogaster but numerous genetic and morphological studies support their treatment at separate species. Race sibirica has occasionally been treated as a distinct species, but morphological data do not support this. The name intermedia is here considered to be a synonym of ognevi , although further morphological and genetic comparison is needed. Three subspecies recognized.	M.h.hilgendorfiPeters,1880—muchofJapan,alsoKunashir(SKurilIs). M.h.ogneviKishida&Mori,1931—RussianFarEast(includingSakhalin),NEChina,andKorea(includingJejuI). M.h. sibirica Kastschenko, 1905 — S Russia , in S Siberia, including Republic of Khakassia , Sayan Mts, Lake Baikal area, and Altai Mts, along with NE Kazakhstan and NW Mongolia.	Head-body 46-5-69- 5 mm , tail 3347 mm , ear 15-20 mm , hindfoot 9-5-13- 8 mm , forearm 39-5—46- 2 mm ; weight 9-11 g . Fur is soft, woolly, and glossy, being composed ofsilky straight guard hairs and soft curly hairs; dorsally silvery brownish gray (hairs have four bands, with dark grayish-brown base, followed by a paler grayish-brown band, an olive- to orange-brown band, and a golden buff tip; guard hairs are silvery and glossy); ventrally paler (hairs with dark brown bases and silvery tips). Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet. Face is sparsely haired except long protuberant nostrils, which are naked. Ears are short and broad (more rounded in ognevt), with smoothly convex anterior margins, distinct notch on posterior margin (less defined in ognevi ), and broadly roundedtip; tragus is long, narrow, and tapering toward pointed tip, which is either bent outwards or straight. Wing attaches to base offirst toe. Baculum is short (1-8- 2 mm ) and saddle-shaped, with bluntly rounded tip, open base, and deep, wide ventral groove. Skull is robust, with short, stout rostrum; braincase 1s low and narrow; lambdoidal crests are well developed, forming occipital helmet; sagittal crest is present but low (absent or barely developed posteriorly). P* is more than half the size of P% M,is highly reduced. Chromosomal complement has 2n = 44 and FNa = 50, 56, or 58.	A variety of temperate hilly or montane habitats, including coniferous-broadleaf and broadleaf forests. It can be found in taiga forest in Mongolia and Russia . It forages in forested and open areas. Recorded at elevations up to 4000 m .	Hilgendorf’s Tube-nosed Bat apparently feeds on flying insects, foraging close to the ground and in the understory offorests. It may perform ground gleaning or quadrupedal ground feeding, and is regularly caughtin pitfall traps. It can hover and has powerful and maneuverable flight. In captivity, it will take mealworm larvae ( 4-10 g /day) but refuses basket worms.	Births have been reported in July in Japan , with a litter size of 1-3. After birth, the ears erect after 3—4 days, eyes open at 8-10 days, and young are completely furred by 12-14 days. Replacement of deciduous teeth with permanent teeth is completed at 23-24 days. Maximum recorded life span is 16 years, although the species typically only lives up to 5-9 years.	Hilgendorf’s Tube-nosed Bat has been reported roosting in a variety of sites, including foliage, tree branches, flying-squirrel nestboxes, various structural objects, houses, abandoned mines, caves, tunnels, and karst caves. They hibernate through the winter from December to March,for ¢.100 days. While hibernating, their body temperature is relatively low at an average of 3-4°C. Calls are a steep FM sweep with start frequency of 104-9 kHz (82-8-140 kHz), end frequency 43-6 kHz (44-9-58-4 kHz), peak frequency 51-2 kHz (46-56-5 kHz), and average duration 1-8 milliseconds (1:1-2-7 milliseconds) in Hokkaido , Japan .	Hilgendorf’s Tube-nosed Bat roosts in small groups generally of 1-10 bats during summer, although colonies 10-100 or more have been reported aggregated in Japan , between early February and late June. Colony size reaches a maximum from late April to earlyJune, although the bats disperse by early June. These colonies have an almost 1:1 sex ratio, and the role of these colonies during this period is still unclear; they may be connected with reproduction. Bats are known to switch roosts often, probably every 1-2 days. A study in Yamanashi , Japan , found that bats seemed to prefer open over hidden roosts throughout the year. They used abandoned mines and tunnels from May to November, particularly in May-July; dead summer cypress thickets were used in June—-October; and outer walls of buildings were used in June and October. During hibernation, most bats have been found roosting alone in crevices and secluded areas, but in Japan there are reports of colonies of several hundred roosting together, although this appears to be uncommon.	Classified as Least Concern on The IUCN Red List. The species 1s widespread but naturally rare throughout its range.	Abe (2004) | Csorba & Bates (2005) | Fukui et al. (2004) | Harada (2011) | Harada etal. (1987a) | Hashimoto & Matsumura (2008) | Ishida et al. (2012) | Jo Yeong-Seok et al. (2018) | Kawai et al. (2014) | Kazakov et al. (2016) | Kruskop (2005) | Kuo Haochih et al. (2009) | Kuramoto (1964, 1981) | Ohdachi et al. (2009) | Smith & Xie Yan (2008) | Takada et al. (2017) | Tsytsulina (2008d) | Urano (2011) | Watanabe & Funakoshi (2017) | Yoshiyuki (1989) | Zeng Xiang et al. (2018)	https://zenodo.org/record/6398567/files/figure.png	333. Hilgendorf’s Tube-nosed Bat Murina hilgendorfi French: Murine de Hilgendorf / German: Hilgendorf-Rohrennase / Spanish: Ratonero narizudo de Hilgendorf Other common names: Greater Tube-nosed Bat Taxonomy. Harpyiocephalus hilgendorfi Peters, 1880 , Yedo, near Tokyo , Japan . Relationships uncertain but limited genetic data place this species as sister to the M. leucogaster clade (see above). Often considered a race of M. leucogaster but numerous genetic and morphological studies support their treatment at separate species. Race sibirica has occasionally been treated as a distinct species, but morphological data do not support this. The name intermedia is here considered to be a synonym of ognevi , although further morphological and genetic comparison is needed. Three subspecies recognized. Subspecies and Distribution. M.h.hilgendorfiPeters,1880—muchofJapan,alsoKunashir(SKurilIs). M.h.ogneviKishida&Mori,1931—RussianFarEast(includingSakhalin),NEChina,andKorea(includingJejuI). M.h. sibirica Kastschenko, 1905 — S Russia , in S Siberia, including Republic of Khakassia , Sayan Mts, Lake Baikal area, and Altai Mts, along with NE Kazakhstan and NW Mongolia. Descriptive notes. Head-body 46-5-69- 5 mm , tail 3347 mm , ear 15-20 mm , hindfoot 9-5-13- 8 mm , forearm 39-5—46- 2 mm ; weight 9-11 g . Fur is soft, woolly, and glossy, being composed ofsilky straight guard hairs and soft curly hairs; dorsally silvery brownish gray (hairs have four bands, with dark grayish-brown base, followed by a paler grayish-brown band, an olive- to orange-brown band, and a golden buff tip; guard hairs are silvery and glossy); ventrally paler (hairs with dark brown bases and silvery tips). Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet. Face is sparsely haired except long protuberant nostrils, which are naked. Ears are short and broad (more rounded in ognevt), with smoothly convex anterior margins, distinct notch on posterior margin (less defined in ognevi ), and broadly roundedtip; tragus is long, narrow, and tapering toward pointed tip, which is either bent outwards or straight. Wing attaches to base offirst toe. Baculum is short (1-8- 2 mm ) and saddle-shaped, with bluntly rounded tip, open base, and deep, wide ventral groove. Skull is robust, with short, stout rostrum; braincase 1s low and narrow; lambdoidal crests are well developed, forming occipital helmet; sagittal crest is present but low (absent or barely developed posteriorly). P* is more than half the size of P% M,is highly reduced. Chromosomal complement has 2n = 44 and FNa = 50, 56, or 58. Habitat. A variety of temperate hilly or montane habitats, including coniferous-broadleaf and broadleaf forests. It can be found in taiga forest in Mongolia and Russia . It forages in forested and open areas. Recorded at elevations up to 4000 m . Food and Feeding. Hilgendorf’s Tube-nosed Bat apparently feeds on flying insects, foraging close to the ground and in the understory offorests. It may perform ground gleaning or quadrupedal ground feeding, and is regularly caughtin pitfall traps. It can hover and has powerful and maneuverable flight. In captivity, it will take mealworm larvae ( 4-10 g /day) but refuses basket worms. Breeding. Births have been reported in July in Japan , with a litter size of 1-3. After birth, the ears erect after 3—4 days, eyes open at 8-10 days, and young are completely furred by 12-14 days. Replacement of deciduous teeth with permanent teeth is completed at 23-24 days. Maximum recorded life span is 16 years, although the species typically only lives up to 5-9 years. Activity patterns. Hilgendorf’s Tube-nosed Bat has been reported roosting in a variety of sites, including foliage, tree branches, flying-squirrel nestboxes, various structural objects, houses, abandoned mines, caves, tunnels, and karst caves. They hibernate through the winter from December to March,for ¢.100 days. While hibernating, their body temperature is relatively low at an average of 3-4°C. Calls are a steep FM sweep with start frequency of 104-9 kHz (82-8-140 kHz), end frequency 43-6 kHz (44-9-58-4 kHz), peak frequency 51-2 kHz (46-56-5 kHz), and average duration 1-8 milliseconds (1:1-2-7 milliseconds) in Hokkaido , Japan . Movements, Home range and Social organization. Hilgendorf’s Tube-nosed Bat roosts in small groups generally of 1-10 bats during summer, although colonies 10-100 or more have been reported aggregated in Japan , between early February and late June. Colony size reaches a maximum from late April to earlyJune, although the bats disperse by early June. These colonies have an almost 1:1 sex ratio, and the role of these colonies during this period is still unclear; they may be connected with reproduction. Bats are known to switch roosts often, probably every 1-2 days. A study in Yamanashi , Japan , found that bats seemed to prefer open over hidden roosts throughout the year. They used abandoned mines and tunnels from May to November, particularly in May-July; dead summer cypress thickets were used in June—-October; and outer walls of buildings were used in June and October. During hibernation, most bats have been found roosting alone in crevices and secluded areas, but in Japan there are reports of colonies of several hundred roosting together, although this appears to be uncommon. Status and Conservation. Classified as Least Concern on The IUCN Red List. The species 1s widespread but naturally rare throughout its range. Bibliography. Abe (2004), Csorba & Bates (2005), Fukui et al. (2004), Harada (2011), Harada etal. (1987a), Hashimoto & Matsumura (2008), Ishida et al. (2012), Jo Yeong-Seok et al. (2018), Kawai et al. (2014), Kazakov et al. (2016), Kruskop (2005), Kuo Haochih et al. (2009), Kuramoto (1964, 1981), Ohdachi et al. (2009), Smith & Xie Yan (2008), Takada et al. (2017), Tsytsulina (2008d), Urano (2011), Watanabe & Funakoshi (2017), Yoshiyuki (1989), Zeng Xiang et al. (2018).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Murina hilgendorfi	Murina	Murina	hilgendorfi	Peters	1880	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	######	Hilgendorf's Tube-nosed Bat	 intermedia Mori, 1933; ognevi Bianchi, 1916; sibirica  Kastschenko, 1905	Japan, near Tokyo, Yedo.	N China; Upper Yenisei River (Russia); Altai Mtns (Russia, Kazakhstan and Mongolia); Korea; Ussur region (Russia); Sakhalin Isl (Russia); Honshu, Kyushu and Shikiku (Japan).	Not listed.	Least Concern	Subgenus Murina . Formerly included in leucogaster , but apparently distinct. May include more than one species; see Yoshiyuki(1989). Also see Wang (1959).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Murina hilgendorfi	23	Hilgendorf's Tube-nosed Bat	Greater Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MURININAE	NA	Murina	NA	hilgendorfi	W. Peters	1880	1	Harpiocephalus_hilgendorfi	Peters, W. C. H. (1880). Mittheilung Ã¼ber die von Hrn. Dr. F. Hilgendorf in Japan gesammelten Chiropteren. Monatsberichte der KÃ¶niglich Preussischen Akademie der Wissenschaften zu Berlin, 1880, 24.	https://www.biodiversitylibrary.org/item/111870#page/32/mode/1up	ZMB 5580		Yedo, near Tokyo, Japan.			hilgendorfi (W. Peters, 1880)|sibirca (Kastschenko, 1905)|ognevi Bianchi, 1916|intermedia Mori, 1933	NA	NA	Kazakhstan?|Mongolia|Russia|China|North Korea|South Korea|Japan	Asia	Palearctic	LC	0	0	0	Murina_hilgendorfi	0	sciname match	Murina_hilgendorfi	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	136409	Murina hilgendorfi	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Murina	hilgendorfi	Peters, 1880	Muirna hilgendorfi was once synonymized with the related M. leucogaster (Bobrinskyi et al. 1965), and this point ov view was accepted until recently (e.g. Koopman 1994, Botvinkin 2002). Yoshiyuki considered M. hilgendorfi from Japan a distinct species (Yoshiyuki 1989), but erroneously divided it from all mainland populations. Currently species status of M. hilgendorfi is accepted (Simmons 2005) and got some support from molecular data (Kruskop et al. 2012). Relationship with poorly known M. fusca from North-East China require further studies.	20000000	Murina hilgendorfi	Least Concern		2019	2018-08-31 00:00:00 UTC	3.1	English	This species has relatively wide range, and spread across its range. It is not thought to be declining fast enough to be listed in a more threatened category. However, as cave disturbance and logging activity increases, habitat loss may pose an increasingly great threat in the future. Monitoring is needed and the species should be reassessed if new data becomes available. The species il listed Least Concern.	Inhabits hilly and mountainous areas, up to 4000 m ASL, with mixed coniferous-broadleaved and broadleaved forests and usually with underground cavities. In northern Mongolia, it is found in taiga zone. Day roosts were found in tree canopies and other tree- or timber-associated shelters (under loose bark, in firewood piles, etc.), and also in caves. In China, this species is known to roost in caves, in trees, and in houses. In Russia, summer roosts are almost unknown, hibernation roosts are in natural caves, where up to 1,000 bat? Could be found simultaneously (Botvinkin 2002). They forage in both forested and open areas. In Japan, this species roosts in various objects: foliage, tree branches, bat boxes, houses, caves, abandoned tunnels (Kawai 2015). From early summer to early autumn, both sexes roosts solitarily or few individuals except maternity colonies. Between early February and late June, 10 to more than 100 individuals aggregate in a cave. Parturition occurs in July. Litter size varies from 1 to 3. Insectivorous, probably a ground-gleaner (at least in part), picking its prey from soil or vegetation surface. The foraging flight is powerful and maneuverable. This species is unusual because it is a ground feeder, searching for beetles. This information is verified by the regularity of them being caught in different rodent traps (Botvinkin 2002), and never being caught in traps higher than ground level (S. Dorjderem pers. comm.).  110 4060 (Fukui et al. 2004) Maximum recorded longevity is up to 16 years (usually ca. 5-9).	No available data across much of the range, although as logging activity and cave disturbance increase, habitat loss may constitute a threat in the future. ; Large part of the species range in Russia occurs in low populated areas, which decrease potential treats for the Russian populations. Large wintering aggregations in caves could be potentially affected by increasing of speleotourism.	There is no population information available on this species at present. It is considered rare in Japan. It was thought to be rare in Russia, based on summer records, but it is quite numerous in cave hibernation sites, showing that its real population is relatively large (Krivosheev 1984, Botvinkin 2002).	Unknown	This species is known from northern China, Upper Yenisei River, Altai mountains, northern Mongolia), Republic of Tyva, Baikal area, Korea, Russian Far East (Primoriye and Khabarovsk region), Sakhalin, Kunashir Island, and Japan. In China, it is known from Great Xingâ€™an Mountain, Nei Mongol and Heilongjiang (Smith and Xie 2008). In Japan, it is known from Hokkaido, Honshu, Shikoku and Kyushu (Kawai 2015).	This species has no commercial value and does not used in commercial trade. It is not known to be used in local cuisine or local medicine anywhere across the range. It could be a potential vector of lyssaviruses (Botvinkin et al. 2003).	Terrestrial	In Mongolia, approximately 8% of the speciesâ€™ range occurs within protected areas (Dulamtseren et al. 2006). It was considered as Vulnerable on previous version of the Japanese Red List (Ministry of the Environment 2007), whereas not listed in the current version due to increasing of distributional records (Ministry of the Environment 2014). It is listed as Least Concern (LC) in the Chinese Red List (Jiang et al. 2016). It is not listed in the Russian Red Data list, but is included in few regional Red Lists. Further research on all aspects of ecology and population trends is strongly recommended.	Palearctic		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Murina	Murina	hilgendorfi	Peters	1880	0	Monatsb. K. Preuss. Akad. Wiss. Berlin	########	Hilgendorf's Tube-nosed Bat	 intermedia Mori, 1933; ognevi Bianchi, 1916; sibirica  Kastschenko, 1905	Japan, near Tokyo, Yedo.	N China; Upper Yenisei River (Russia); Altai Mtns (Russia, Kazakhstan and Mongolia); Korea; Ussur region (Russia); Sakhalin Isl (Russia); Honshu, Kyushu and Shikiku (Japan).	Not listed.	Least Concern	Subgenus Murina . Formerly included in leucogaster , but apparently distinct. May include more than one species; see Yoshiyuki(1989). Also see Wang (1959).	Murina hilgendorfi	1005335	23	Hilgendorf's Tube-nosed Bat	Greater Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MURININAE	NA	Murina	NA	hilgendorfi	W. Peters	1880	1	Harpiocephalus_hilgendorfi	Peters, W. C. H. (1880). Mittheilung Ã¼ber die von Hrn. Dr. F. Hilgendorf in Japan gesammelten Chiropteren. Monatsberichte der KÃ¶niglich Preussischen Akademie der Wissenschaften zu Berlin, 1880, 24.	https://www.biodiversitylibrary.org/item/111870#page/32/mode/1up	ZMB 5580		Yedo, near Tokyo, Japan.			hilgendorfi (W. Peters, 1880)|sibirca (Kastschenko, 1905)|ognevi Bianchi, 1916|intermedia Mori, 1933	NA	NA				Kazakhstan?|Mongolia|Russia|China|North Korea|South Korea|Japan	Asia	Palearctic	LC	0	0	0	Murina_hilgendorfi	0	sciname match	Murina_hilgendorfi	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Murina_hilgendorfi	1005335	23	Hilgendorf's Tube-nosed Bat	Greater Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Murininae	NA	Murina	NA	hilgendorfi	W. C. H. Peters	1	Harpyiocephalus Hilgendorfi	Peters, W.C.H. 1880. Mittheilung Ã¼ber die von Hrn. Dr. F. Hilgendorf in Japan gesammelten Chiropteren. Monatsberichte der KÃ¶niglichen Preussischen Akademie der Wissenschaften zu Berlin 1880:23-25.	https://www.biodiversitylibrary.org/page/36277225	ZMB 5580	holotype		Yedo, near Tokyo, Japan.			NA	NA				Kazakhstan?|Mongolia|Russia|China|North Korea|South Korea|Japan	Asia	Palearctic	LC	0	0	0	Murina_hilgendorfi	0	sciname match	Murina_hilgendorfi	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Murina	Murina	hilgendorfi	Peters	1880	1	Monatsb. K. Preuss. Akad. Wiss. Berlin	########	Hilgendorf's Tube-nosed Bat	intermedia Mori, 1933; ognevi Bianchi, 1916; sibirica  Kastschenko, 1905	Japan, near Tokyo, Yedo.	N China; Upper Yenisei River (Russia); Altai Mtns (Russia, Kazakhstan and Mongolia); Korea; Ussur region (Russia); Sakhalin Isl (Russia); Honshu, Kyushu and Shikiku (Japan).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/136409/22017193/' target='_blank'>Least Concern</a>	Subgenus Murina. Formerly included in leucogaster, but apparently distinct. May include more than one species; see Yoshiyuki(1989). Also see Wang (1959).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Murina hilgendorfi; Murina hilgendorfi; Murina hilgendorfi; Murina hilgendorfi; Murina hilgendorfi; Murina hilgendorfi; intermedia; ognevi; sibirica; hilgendorfi; ognevi; sibirica; ognevi - intermedia?; intermedia; ognevi; sibirica; hilgendorfi; sibirca; ognevi; intermedia; Murine de Hilgendorf; Hilgendorf-Rohrennase; Ratonero narizudo de Hilgendorf; Greater Tube-nosed Bat; Hilgendorf's Tube-nosed Bat; Greater Tube-nosed Bat; Hilgendorf's Tube-nosed Bat; Hilgendorf's Tube-nosed Bat; M. hilgendorfi
