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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L769	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Murina harrisoni	Murina harrisoni	Murina harrisoni	Murina harrisoni	Murina harrisoni	Murina harrisoni	Murina harrisoni	Murina harrisoni	Murina harrisoni		[HMW] Murina harrisoni Csorba & Bates, 2005 , “O TukChehn, Kirirom National Park, Kompong Speu Province, Cambodia , 11º 29.611'N , 104°12.746'E .” Based on limited genetic data, Murina harrisoni appears to be related to M. ussuriensis , M. huttonii , and M. puta . Includes tiensa from Vietnam ,first thoughtto be a “large form” of M. huttonii , then named as a new species in 2007, but subsequently shown to be a synonym of harrisoni ; there are two genetic clades, but reported morphological differences proved to be due to strong sexual dimorphism. Monotypic.; [batnames2022] Includes tiensa ; see Francis and Eger (2012), Wu et al. (2010), and Son et al. (2015).; [MDD2022] recently described; includes the recently described tiensa; [IUCN] Vietnamese populations of the species were formerly regarded as represent a separate species, ;Murina tiensa , but recent genetic (Francis and Eger 2012), morphological and morphometric studies (Son ;et al. ;2015) showed that – although two distinct genetic clades are present – they are conspecific. The observed morphological differences which were once thought to be diagnostic to separate the two species are intraspecific variations and are due to the strong sexual dimorphism.; [batnames2023] Includes tiensa ; see Francis and Eger (2012), Wu et al. (2010), and Son et al. (2015).; [MDD2023] recently described; includes the recently described tiensa; [MDD2025_2.0] recently described; includes the recently described tiensa; [batnames2025_1.7] Includes tiensa; see Francis and Eger (2012), Wu et al. (2010), and Son et al. (2015).; [MDD2025_2.2] recently described; includes the recently described tiensa												harrisoni	harrisoni - tiensa	harrisoni, tiensa	Vietnamese populations of the species were formerly regarded as represent a separate species, ;Murina tiensa , but recent genetic (Francis and Eger 2012), morphological and morphometric studies (Son ;et al. ;2015) showed that – although two distinct genetic clades are present – they are conspecific. The observed morphological differences which were once thought to be diagnostic to separate the two species are intraspecific variations and are due to the strong sexual dimorphism.	harrisoni	harrisoni - tiensa	harrisoni, tiensa 	harrisoni, tiensa 	harrisoni 	harrisoni - tiensa	harrisoni Csorba & P. J. J. Bates, 2005|tiensa Csorba, Vu Dinh Thong, P. J. J. Bates, & Furey, 2007						N/A																																								NA																											4C3D87E8FF6C6AD3FF4091451447BAA0	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	912	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF6C6AD3FF4091451447BAA0.xml	Murina harrisoni	Vespertilionidae	Murina	harrisoni	Csorba & Bates	2005	Murine de Harrison @fr | Harrison-Rohrennase @de | Ratonero narizudo de Harrison @es | Harrison's Murine Bat @en	Murina harrisoni Csorba & Bates, 2005 , “O TukChehn, Kirirom National Park, Kompong Speu Province, Cambodia , 11º 29.611'N , 104°12.746'E .” Based on limited genetic data, Murina harrisoni appears to be related to M. ussuriensis , M. huttonii , and M. puta . Includes tiensa from Vietnam ,first thoughtto be a “large form” of M. huttonii , then named as a new species in 2007, but subsequently shown to be a synonym of harrisoni ; there are two genetic clades, but reported morphological differences proved to be due to strong sexual dimorphism. Monotypic.	C Myanmar , N & W Thailand , N Laos , Vietnam , S Cambodia , and SE China ( Guangxi and Hainan ).	Head-body 38-9-55- 1 mm ,tail 33-8-45- 5 mm , ear 12-2-18- 8 mm , hindfoot 7-4-10- 2 mm , forearm 34-40- 1 mm ; weight 8:4-10- 5 g . Females are larger than males. Fur is long and silky; dorsally orangish brown (hairs with pale base, the rest being yellowish red, gradually darkening toward tip; no shiny guard hairs); ventrally grayish brown (hairs uncolored whitish or light gray). Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet. Face is sparsely haired except long protuberant nostrils, which are naked. Ears are short, broad, and rounded, with smoothly convex anterior margins, no notch on posterior margin, with a broadly rounded tip; tragus is long, narrow, and tapering toward pointed tip. Wing attaches to base of claw on first toe. Skull has uninflated rostrum and flat braincase; sagittal and lambdoidal crests are well developed. I? is lateral to I’; C' is much taller than P* and is equal or slightly larger in basal area; P* is subequal in height to P* mesostyles of M' and M? are weakly developed; talonids of M, and M, are two-thirds the size of their respective trigonids. Chromosomal complement has 2n = 44 and FN = 50 (Hainan and Thailand ).	Subtropical montane secondary forest in Guangxi, undisturbed tropical rainforest in Hainan, degraded to nearly pristine lowland and montane forests in Vietnam , and disturbed semi-evergreen gallery forest and dry lowland dipterocarp forest in Cambodia .	No information.	No information.	In Vietnam ,in flight, calls were a steep FM sweep with an average start frequency of 145-1 kHz (122-2-164-8 kHz), end frequency 50-1 kHz (45-8-55-8 kHz), and duration 1-9 milliseconds (1-3-3-1 milliseconds).	No information.	Classified as Least Concern on The IUCN Red List.	Csorba & Bates (2005) | Csorba, Bates & Francis (2016) | Csorba, Thong Vu Dinh et al. (2007) | Francis & Eger (2012) | Nguyen Truong Son etal. (2015) | Soisook (2013) | Thong Vu Dinh et al. (2011) | Wu Yi et al. (2010)	https://zenodo.org/record/6398586/files/figure.png	342. Harrison's Tube-nosed Bat Murina harrisoni French: Murine de Harrison / German: Harrison-Rohrennase / Spanish: Ratonero narizudo de Harrison Other common names: Harrison's Murine Bat Taxonomy. Murina harrisoni Csorba & Bates, 2005 , “O TukChehn, Kirirom National Park, Kompong Speu Province, Cambodia , 11º 29.611'N , 104°12.746'E .” Based on limited genetic data, Murina harrisoni appears to be related to M. ussuriensis , M. huttonii , and M. puta . Includes tiensa from Vietnam ,first thoughtto be a “large form” of M. huttonii , then named as a new species in 2007, but subsequently shown to be a synonym of harrisoni ; there are two genetic clades, but reported morphological differences proved to be due to strong sexual dimorphism. Monotypic. Distribution. C Myanmar , N & W Thailand , N Laos , Vietnam , S Cambodia , and SE China ( Guangxi and Hainan ). Descriptive notes. Head-body 38-9-55- 1 mm ,tail 33-8-45- 5 mm , ear 12-2-18- 8 mm , hindfoot 7-4-10- 2 mm , forearm 34-40- 1 mm ; weight 8:4-10- 5 g . Females are larger than males. Fur is long and silky; dorsally orangish brown (hairs with pale base, the rest being yellowish red, gradually darkening toward tip; no shiny guard hairs); ventrally grayish brown (hairs uncolored whitish or light gray). Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet. Face is sparsely haired except long protuberant nostrils, which are naked. Ears are short, broad, and rounded, with smoothly convex anterior margins, no notch on posterior margin, with a broadly rounded tip; tragus is long, narrow, and tapering toward pointed tip. Wing attaches to base of claw on first toe. Skull has uninflated rostrum and flat braincase; sagittal and lambdoidal crests are well developed. I? is lateral to I’; C' is much taller than P* and is equal or slightly larger in basal area; P* is subequal in height to P* mesostyles of M' and M? are weakly developed; talonids of M, and M, are two-thirds the size of their respective trigonids. Chromosomal complement has 2n = 44 and FN = 50 (Hainan and Thailand ). Habitat. Subtropical montane secondary forest in Guangxi, undisturbed tropical rainforest in Hainan, degraded to nearly pristine lowland and montane forests in Vietnam , and disturbed semi-evergreen gallery forest and dry lowland dipterocarp forest in Cambodia . Food and Feeding. No information. Breeding. No information. Activity patterns. In Vietnam ,in flight, calls were a steep FM sweep with an average start frequency of 145-1 kHz (122-2-164-8 kHz), end frequency 50-1 kHz (45-8-55-8 kHz), and duration 1-9 milliseconds (1-3-3-1 milliseconds). Movements, Home range and Social organization. No information. Status and Conservation. Classified as Least Concern on The IUCN Red List. Bibliography. Csorba & Bates (2005), Csorba, Bates & Francis (2016), Csorba, Thong Vu Dinh et al. (2007), Francis & Eger (2012), Nguyen Truong Son etal. (2015), Soisook (2013), Thong Vu Dinh et al. (2011), Wu Yi et al. (2010).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Murina harrisoni	Murina		harrisoni	Csorba & Bates	2005	0	Acta Chiropterol.	7(1): 2	Harrison&apos;s Tube-nosed Bat	 tiensa Csorba, Thong, Bates, & Furey, 2007	Cambodia, Kompong Speu Province,, Kirirom National Park, O Tuk Chehn	Known only from the type locality.	Not listed.	Least Concern	Includes tiensa ; see Francis and Eger (2012), Wu et al. (2010), and Son et al. (2015).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Murina harrisoni	23	Harrison's Tube-nosed Bat	Harrison's Murine Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MURININAE	NA	Murina	NA	harrisoni	Csorba & P. J. J. Bates	2005	0	Murina_harrisoni	Csorba, G., & Bates, P. J. (2005). Description of a new species of Murina from Cambodia (Chiroptera: Vespertilionidae: Murininae). Acta Chiropterologica, 7(1), 2.	https://bioone.org/journals/acta-chiropterologica/volume-7/issue-1/1733-5329(2005)7[1:DOANSO]2.0.CO;2/Description-of-a-new-species-of-Murina-from-Cambodia-Chiroptera/10.3161/1733-5329(2005)7[1:DOANSO]2.0.CO;2.full	HZM 1.36316		"O TukChehn, Kirirom National Park, Kompong Speu Province, Cambodia, 11°29.611'N, 104°12.746'E."	11.494	104.21	harrisoni Csorba & P. J. J. Bates, 2005|tiensa Csorba, Vu Dinh Thong, P. J. J. Bates, & Furey, 2007	recently described; includes the recently described tiensa	Csorba, G., & Bates, P. J. (2005). Description of a new species of Murina from Cambodia (Chiroptera: Vespertilionidae: Murininae). Acta Chiropterologica, 7(1), 1-7.|Csorba, G., Thong, V. D., Bates, P. J., & Furey, N. M. (2007). Description of a new species of Murina from Vietnam (Chiroptera: Vespertilionidae: Murininae). Occasional Papers, Museum of Texas Tech University, 268, 1-12.|Francis, C. M., & Eger, J. L. (2012). A review of tube-nosed bats (Murina) from Laos with a description of two new species. Acta Chiropterologica, 14(1), 15-38.	China|Myanmar|Thailand|Laos|Vietnam|Cambodia	Asia	Indomalaya	LC	0	0	0	Murina_harrisoni	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	100000000	Murina harrisoni	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Murina	harrisoni	Csorba &; Bates, 2005	Vietnamese populations of the species were formerly regarded as represent a separate species, ;Murina tiensa , but recent genetic (Francis and Eger 2012), morphological and morphometric studies (Son ;et al. ;2015) showed that – although two distinct genetic clades are present – they are conspecific. The observed morphological differences which were once thought to be diagnostic to separate the two species are intraspecific variations and are due to the strong sexual dimorphism.	20000000	Murina harrisoni	Least Concern		2016	2016-02-21 00:00:00 UTC	3.1	English	<p><span lang="HU">This species has a wide distribution, a presumed large population and a wide habitat tolerance, so it is listed as Least Concern.</p>	<p><span lang="HU">In Cambodia the species has been collected in disturbed semi evergreen gallery forest and in dry lowland dipterocarp forests (Csorba and Bates 2005, Csorba and Furey pers. comm.), in Thailand and Myanmar in hill areas, in Vietnam from degraded forests to nearly pristine habitats (including lowlands and mountain peaks), in Guangxi in subtropical, montane secondary forest (Francis and Eger 2012, Thong et al. 2011), and in Hainan in an undisturbed tropical rainforest (Wu et al. 2010).</p>	The threats to this species are not known.	The abundance and population size of this species are not known.	Unknown	The Harrison Murine Bat (Murina harrisoni ) has been found in ;Cambodia (Csorba and Bates 2005), Laos, Thailand, Myanmar, China (Francis and Eger 2012, Wu et al . 2010), Vietnam (Csorba et al. 2007, Thong et al. 2011, Son et al. ; 2016).<p></p>		Terrestrial	<p><span lang="HU">The species has been collected from several protected areas in Cambodia, Vietnam and China (Csorba and Bates 2005, Son et al. 2015, Francis and Eger 2012).</p>	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Murina		harrisoni	Csorba & Bates	2005	0	Acta Chiropterol.	7(1): 2	Harrison&apos;s Tube-nosed Bat	 tiensa Csorba, Thong, Bates, & Furey, 2007	Cambodia, Kompong Speu Province,, Kirirom National Park, O Tuk Chehn	Known only from the type locality.	Not listed.	Least Concern	Includes tiensa ; see Francis and Eger (2012), Wu et al. (2010), and Son et al. (2015).	Murina harrisoni	1005334	23	Harrison's Tube-nosed Bat	Harrison's Murine Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MURININAE	NA	Murina	NA	harrisoni	Csorba & P. J. J. Bates	2005	0	Murina_harrisoni	Csorba, G., & Bates, P. J. (2005). Description of a new species of Murina from Cambodia (Chiroptera: Vespertilionidae: Murininae). Acta Chiropterologica, 7(1), 2.	https://bioone.org/journals/acta-chiropterologica/volume-7/issue-1/1733-5329(2005)7[1:DOANSO]2.0.CO;2/Description-of-a-new-species-of-Murina-from-Cambodia-Chiroptera/10.3161/1733-5329(2005)7[1:DOANSO]2.0.CO;2.full	HZM 1.36316		"O TukChehn, Kirirom National Park, Kompong Speu Province, Cambodia, 11°29.611'N, 104°12.746'E."	11.49352	104.2124	harrisoni Csorba & P. J. J. Bates, 2005|tiensa Csorba, Vu Dinh Thong, P. J. J. Bates, & Furey, 2007	recently described; includes the recently described tiensa	Csorba, G., & Bates, P. J. (2005). Description of a new species of Murina from Cambodia (Chiroptera: Vespertilionidae: Murininae). Acta Chiropterologica, 7(1), 1-7.|Csorba, G., Thong, V. D., Bates, P. J., & Furey, N. M. (2007). Description of a new species of Murina from Vietnam (Chiroptera: Vespertilionidae: Murininae). Occasional Papers, Museum of Texas Tech University, 268, 1-12.|Francis, C. M., & Eger, J. L. (2012). A review of tube-nosed bats (Murina) from Laos with a description of two new species. Acta Chiropterologica, 14(1), 15-38.				China|Myanmar|Thailand|Laos|Vietnam|Cambodia	Asia	Indomalaya	LC	0	0	0	Murina_harrisoni	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Murina_harrisoni	1005334	23	Harrison's Tube-nosed Bat	Harrison's Murine Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Murininae	NA	Murina	NA	harrisoni	Csorba & P. J. J. Bates	0	Murina harrisoni	Csorba, G. and Bates, P.J.J. 2005-06. Description of a new species of _Murina_ from Cambodia (Chiroptera: Vespertilionidae: Murininae). Acta Chiropterologica 7(1):1-7.	https://doi.org/10.3161/1733-5329(2005)7[1:DOANSO]2.0.CO;2	HZM 1.36316	holotype		"O TukChehn, Kirirom National Park, Kompong Speu Province, Cambodia, 11°29.611'N, 104°12.746'E."	11.49352	104.2124	recently described; includes the recently described tiensa	Csorba, G., & Bates, P. J. (2005). Description of a new species of Murina from Cambodia (Chiroptera: Vespertilionidae: Murininae). Acta Chiropterologica, 7(1), 1-7.|Csorba, G., Thong, V. D., Bates, P. J., & Furey, N. M. (2007). Description of a new species of Murina from Vietnam (Chiroptera: Vespertilionidae: Murininae). Occasional Papers, Museum of Texas Tech University, 268, 1-12.|Francis, C. M., & Eger, J. L. (2012). A review of tube-nosed bats (Murina) from Laos with a description of two new species. Acta Chiropterologica, 14(1), 15-38.				China|Myanmar|Thailand|Laos|Vietnam|Cambodia	Asia	Indomalaya	LC	0	0	0	Murina_harrisoni	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Murina		harrisoni	Csorba & Bates	2005	0	Acta Chiropterol.	7(1): 2	Harrison&apos;s Tube-nosed Bat	tiensa Csorba, Thong, Bates, & Furey, 2007	Cambodia, Kompong Speu Province,, Kirirom National Park, O Tuk Chehn	Known only from the type locality.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/99712630/21995130/' target='_blank'>Least Concern</a>	Includes tiensa; see Francis and Eger (2012), Wu et al. (2010), and Son et al. (2015).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Murina harrisoni; Murina harrisoni; Murina harrisoni; Murina harrisoni; Murina harrisoni; tiensa; harrisoni; tiensa; Murine de Harrison; Harrison-Rohrennase; Ratonero narizudo de Harrison; Harrison's Murine Bat; Harrison's Tube-nosed Bat; Harrison's Murine Bat; Harrison&apos;s Tube-nosed Bat; M. harrisoni