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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L763	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Murina fionae	Murina fionae	Murina fionae	Murina fionae	Murina fionae	Murina fionae	Murina fionae	Murina fionae	Murina fionae		[HMW] Murina fionae Francis & Eger, 2012 , “Pha Deng, ~ 8 km E of Ban Navang, Khammouan Province , Laos ( 17°57'N , 105°23'E ); altitude 1140 m .” Murina fionae is included in a clade with M. guilleni , M. cf. cyclotis (from India ), M. cyclotis , M. ct. peninsularis (from Borneo), and M. peninsularis based on limited genetic data. Monotypic.; [MDD2022] recently described; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														fionae				fionae	fionae			fionae C. M. Francis & Eger, 2012						N/A																																								NA																											4C3D87E8FF676AD8FF7A943C1DA4B38C	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	917	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF676AD8FF7A943C1DA4B38C.xml	Murina fionae	Vespertilionidae	Murina	fionae	Francis & Eger	2012	Murine de Fiona @fr | Fiona-Rohrennase @de | Ratonero narizudo de Fiona @es	Murina fionae Francis & Eger, 2012 , “Pha Deng, ~ 8 km E of Ban Navang, Khammouan Province , Laos ( 17°57'N , 105°23'E ); altitude 1140 m .” Murina fionae is included in a clade with M. guilleni , M. cf. cyclotis (from India ), M. cyclotis , M. ct. peninsularis (from Borneo), and M. peninsularis based on limited genetic data. Monotypic.	N & C Vietnam , C Laos , and EC Cambodia .	Head-body 42-1-51- 3 mm , tail 37-44- 5 mm , ear 12- 1-16 mm , hindfoot 6:8-9- 3 mm , forearm 34-2-40- 1 mm ; weight 7- 5-12 g . Fur long and silky. Dorsal pelage is orange (hairs with pale buff bases and orange-browntips), with longer guard hairs scattered throughout; venter is pale buff orange and whiter near chin (hairs unicolored buff orange). Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet. Face is sparsely haired except for long protuberant naked nostrils. Ears are short, broad, and rounded, with smoothly convex anterior margins, no notch on posterior margins, and broadly rounded tips; tragusis long and narrow and tapers toward pointed tip. Wing attaches near base of claw on first toe. Baculum has upwardly arched dorsal surface and deeply concave ventral surface, being somewhat heartshaped (small notch at tip and pointed at base) in dorsal view and very short (1- 1 mm long). Skull is large and robust, with well-domed braincase; sagittal crest is very well developed, and lambdoidalcrest is weakly developed; I? is lateral to I; C! is much taller than P*, but basal area is only slightly larger or subequal to that of P*; P? is subequalto P*in height, but its basal area is two-thirds to nearly equal to that of P*; M' and M? lack mesostyles; and talonids of M, and M, are reduced relative to their trigonids.	Wet, hill evergreen forests at elevations of 830-1140 m in the Annamite Mountains and semideciduous forests at 290 m in Cambodia .	No information.	No information.	No information.	No information.	Not assessed on The [UCNRed List. Virtually nothing is known about ecology of Fiona’s Tube-nosed Bat, and additional studies are needed to clarify its conservation status.	Francis & Eger (2012) | Nguyen Truong Son et al. (2015) | Soisook (2013) | Soisook, Karapan, Satasook, Thong Vu Dinh et al. (2013) | Soisook, Thaw Win-Naing et al. (2017)	https://zenodo.org/record/6398610/files/figure.png	354. Fiona’s Tube-nosed Bat Murina fionae French: Murine de Fiona / German: Fiona-Rohrennase / Spanish: Ratonero narizudo de Fiona Taxonomy. Murina fionae Francis & Eger, 2012 , “Pha Deng, ~ 8 km E of Ban Navang, Khammouan Province , Laos ( 17°57'N , 105°23'E ); altitude 1140 m .” Murina fionae is included in a clade with M. guilleni , M. cf. cyclotis (from India ), M. cyclotis , M. ct. peninsularis (from Borneo), and M. peninsularis based on limited genetic data. Monotypic. Distribution. N & C Vietnam , C Laos , and EC Cambodia . Descriptive notes. Head-body 42-1-51- 3 mm , tail 37-44- 5 mm , ear 12- 1-16 mm , hindfoot 6:8-9- 3 mm , forearm 34-2-40- 1 mm ; weight 7- 5-12 g . Fur long and silky. Dorsal pelage is orange (hairs with pale buff bases and orange-browntips), with longer guard hairs scattered throughout; venter is pale buff orange and whiter near chin (hairs unicolored buff orange). Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet. Face is sparsely haired except for long protuberant naked nostrils. Ears are short, broad, and rounded, with smoothly convex anterior margins, no notch on posterior margins, and broadly rounded tips; tragusis long and narrow and tapers toward pointed tip. Wing attaches near base of claw on first toe. Baculum has upwardly arched dorsal surface and deeply concave ventral surface, being somewhat heartshaped (small notch at tip and pointed at base) in dorsal view and very short (1- 1 mm long). Skull is large and robust, with well-domed braincase; sagittal crest is very well developed, and lambdoidalcrest is weakly developed; I? is lateral to I; C! is much taller than P*, but basal area is only slightly larger or subequal to that of P*; P? is subequalto P*in height, but its basal area is two-thirds to nearly equal to that of P*; M' and M? lack mesostyles; and talonids of M, and M, are reduced relative to their trigonids. Habitat. Wet, hill evergreen forests at elevations of 830-1140 m in the Annamite Mountains and semideciduous forests at 290 m in Cambodia . Food and Feeding. No information. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The [UCNRed List. Virtually nothing is known about ecology of Fiona’s Tube-nosed Bat, and additional studies are needed to clarify its conservation status. Bibliography. Francis & Eger (2012), Nguyen Truong Son et al. (2015), Soisook (2013), Soisook, Karapan, Satasook, Thong Vu Dinh et al. (2013), Soisook, Thaw Win-Naing et al. (2017).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Murina fionae	Murina		fionae	Francis & Eger	2012	0	Acta Chiropterol.	14(1): 32	Fiona's Tube-nosed Bat	None.	Laos, Khammouan Province, Pha Deng, 8kn E of Ban Navang	Laos, Vietnam, and Cambodia	Not listed.	Least Concern		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Murina fionae	23	Fiona's Tube-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MURININAE	NA	Murina	NA	fionae	Francis & Eger	2012	0	Murina_fionae	Francis, C. M. & Eger, J. L. (2012). A review of Tube-nosed bats (Murina) from Laos with a description of two new species. Acta Chiropterologica, 14(1), 32.	https://bioone.org/journals/acta-chiropterologica/volume-14/issue-1/150811012X654231/A-Review-of-Tube-Nosed-Bats-Murina-from-Laos-with/10.3161/150811012X654231.full	ROM 106383		"Pha Deng, ≈8 km E of Ban Navang, Khammouan Province, Laos (17°57'N, 105°23'E); altitude 1140 m."	17.95	105.38	fionae Francis & Eger, 2012	recently described	Francis, C. M., & Eger, J. L. (2012). A review of tube-nosed bats (Murina) from Laos with a description of two new species. Acta Chiropterologica, 14(1), 15-38.	Vietnam|Laos|Cambodia	Asia	Indomalaya	LC	0	0	0	Murina_fionae	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Murina fionae	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Murina	fionae	Francis &; Eger, 2012		80000000	Murina fionae	Least Concern		2020	2018-09-01 00:00:00 UTC	3.1	English	This species is listed as Least Concern. Although there is not much information on the population status or ecology of this species, it is unlikely to meet any of the criteria for Vulnerable or higher listing. It has been reported from at least 10 widely scattered localities from Central Laos and Vietnam to south Laos and Cambodia, such that the known extent of occurrence exceeds 20,000 km2. ; There are no data on population size, but there is no reason to believe it is less than 10,000 individuals, although it does appear to be less common than several other Murina in the region. Although the population has undoubtedly declined due to loss of forested habitat, it has been caught in more than one forest type, and there is no evidence that it has declined fast enough to qualify for listing in a threatened category.	Little is known of the ecology of this species, and no roosting sites have been reported. Records from Lao PDR and Vietnam for which habitat information was reported were captured in harp traps set across trails in wet evergreen hill forest in the Annamite mountains at altitudes of 830m to 1,140m. A specimen from Cambodia was caught in semi-deciduous forest at an altitude of 290m. Most sites involved capture of 1 or 2 individuals, suggesting it likely occurs mainly singly or in small family groups.	Populations have likely declined over the past several decades due to clearing of forest for agriculture or settlements, but without reliable information on habitat specificity, including the extent to which it can use disturbed forests, it is not possible to estimate the extent of habitat loss for the species.	No estimates are available on population size. The species has been caught at 10 localities, with one or two individuals at most sites, but likely occurs in many other locations in between.	Unknown	The species has so far been reported from 10 localities (Francis and Eger 2012, Son et al.   2015) from central Laos and central Vietnam south to Cambodia and south Vietnam. The type locality was Lao People’s Democratic Republic, Khammouan Province, Pha Deng, ~8 km E of Ban Navang (17°57’N, 105°23’E), altitude 1,140 m.	No records of trade in this species. Presumably roosts individually or in small groups, like other members of the genus, which would make it unlikely to be readily captured in significant numbers for consumption or trade.	Terrestrial	Has been reported from reserves or biodiversity conservation areas in all three countries.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Murina		fionae	Francis & Eger	2012	0	Acta Chiropterol.	14(1): 32	Fiona's Tube-nosed Bat	None.	Laos, Khammouan Province, Pha Deng, 8kn E of Ban Navang	Laos, Vietnam, and Cambodia	Not listed.	Least Concern		Murina fionae	1005328	23	Fiona's Tube-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MURININAE	NA	Murina	NA	fionae	Francis & Eger	2012	0	Murina_fionae	Francis, C. M. & Eger, J. L. (2012). A review of Tube-nosed bats (Murina) from Laos with a description of two new species. Acta Chiropterologica, 14(1), 32.	https://bioone.org/journals/acta-chiropterologica/volume-14/issue-1/150811012X654231/A-Review-of-Tube-Nosed-Bats-Murina-from-Laos-with/10.3161/150811012X654231.full	ROM 106383		"Pha Deng, ≈8 km E of Ban Navang, Khammouan Province, Laos (17°57'N, 105°23'E); altitude 1140 m."	17.95	105.3833	fionae Francis & Eger, 2012	recently described	Francis, C. M., & Eger, J. L. (2012). A review of tube-nosed bats (Murina) from Laos with a description of two new species. Acta Chiropterologica, 14(1), 15-38.				Vietnam|Laos|Cambodia	Asia	Indomalaya	LC	0	0	0	Murina_fionae	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Murina_fionae	1005328	23	Fiona's Tube-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Murininae	NA	Murina	NA	fionae	C. M. Francis & Eger	0	Murina fionae	Francis, C.M. and Eger, J.L. 2012-06. A review of tube-nosed bats (_Murina_) from Laos with a description of two new species. Acta Chiropterologica 14(1):15-38.	https://doi.org/10.3161/150811012X654231	ROM:MAM:106383	holotype	http://portal.vertnet.org/o/rom/mammals?id=uri-catalog-rom-mammals-106383	"Pha Deng, ≈8 km E of Ban Navang, Khammouan Province, Laos (17°57'N, 105°23'E); altitude 1140 m."	17.95	105.3833	recently described	Francis, C. M., & Eger, J. L. (2012). A review of tube-nosed bats (Murina) from Laos with a description of two new species. Acta Chiropterologica, 14(1), 15-38.				Vietnam|Laos|Cambodia	Asia	Indomalaya	LC	0	0	0	Murina_fionae	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Murina		fionae	Francis & Eger	2012	0	Acta Chiropterol.	14(1): 32	Fiona's Tube-nosed Bat	None.	Laos, Khammouan Province, Pha Deng, 8kn E of Ban Navang	Laos, Vietnam, and Cambodia	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84500852/84500855/' target='_blank'>Least Concern</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Murina fionae; Murina fionae; Murina fionae; Murina fionae; Murina fionae; fionae; Murine de Fiona; Fiona-Rohrennase; Ratonero narizudo de Fiona; Fiona's Tube-nosed Bat; Fiona's Tube-nosed Bat; M. fionae