http://www.w3.org/ns/prov#wasDerivedFrom	http://purl.org/dc/elements/1.1/format	name_CH1_1980	name_MSW1_1982	name_CH3_1991	name_MSW2_1993	name_Koopman_1994	name_MSW3_2005	name_HMW_2019	name_BatNames_2022	name_MDD_2022	name_IUCN_2022	name_BatNames_2023	name_MDD_2023	name_MDD_2025_2.0	name_batnames_2025_1.7	name_MDD_2025_2.2	column151	taxonomic_notes_concatenated	column171	synonyms_CH1	subspecies__MSW2	synonyms__MSW1	synonyms_CH3	synonyms_MSW2	subspecies_Koopman94_interpreted	subspecies_MSW3_interpreted	synonym_MSW3_interpreted	subspecies_HMW_interpreted	synonym_HMW_interpreted	subspecies_batnames_interpreted	synonym_batnames_interpreted	synonym_MDD_interpreted	synonym_IUCN_interpreted	subspecies_batnames2023_interpreted	synonym_batnames2023_interpreted	synonym_MDD2023_interpreted	synonym_MDD2025_interpreted	subspecies_batnames2025_interpreted	synonyms_batnames2025_interpreted	nominalNames	column391	docOrigin_CH1	commonName_CH1	distribution_CH1	docOrigin_MSW1	column451	typeLocality_MSW1	authority_MSW1	year_MSW1	citation_MSW1	distribution	comment_MSW1	docOrigin_CH3	commonName_CH3	distribution_CH3	docOrigin_MSW2	authority_MSW2	year_MSW2	citation_MSW2	comments_MSW2	distribution_MSW2	typeLocality_MSW2	docOrigin_Koopman94	authority_Koopman94	year_Koopman94	description_Koopman94	distribution_Koopman94	diversity_Koopman94	subspecies_Koopman94	page	rank	name	authority	year	parent	parent_rank	corrected_name	actual_species_count	claimed_species_count	dental_formula	description	diversity	full_subspecies_text	name_line	species_index	subspecies	synonym	text	docOrigin_MSW3	order_MSW3	family_MSW3	subfamily_MSW3	tribe_MSW3	name_MSW3	genus_MSW3	subgenus_MSW3	species_MSW3	authoritySpeciesAuthor_MSW3	(parentheses (1=author & date in parentheses)_MSW3	authoritySpeciesYear_MSW3	actualDate_MSW3	citation_MSW3	volume_MSW3	issue_MSW3	pages_MSW3	type_species_MSW3	commonName_MSW3	typeLocality_MSW3	distribution_MSW3	status_MSW3	synonym_MSW3	comments_MSW3	docId_HMW	docOrigin_HMW	docISBN_HMW	docName_HMW	docMasterId_HMW	docPageNumber_HMW	derivedFrom_HMW	name_HMW	family_HMW	genus_HMW	species_HMW	authoritySpeciesAuthor_HMW	authoritySpeciesYear	commonNames_HMW	taxonomy_HMW	subspeciesAndDistribution_HMW	descriptiveNotes_HMW	habitat_HMW	foodAndFeeding_HMW	breeding_HMW	activityPatterns_HMW	movementsHomeRangeAndSocialOrganization_HMW	statusAndConservation_HMW	bibliography_HMW	distributionImageURL_HMW	verbatimText_HMW	docOrigin_batnames	family_batnames	name_batnames	genus_batnames	subgenus_batnames	species_batnames	authoritySpeciesAuthor_batnames	date_batnames	parentheses_batnames (1=author & date in parentheses)	citation_batnames	docPageNumber_batnames	common Name_batnames	synonyms_batnames	type_locality_batnames	Distribution_batnames	CITES_batnames	IUCN_batnames	comments_batnames	docOrigin_MDD	name_MDD	phylosort_MDD	mainCommonName_MDD	otherCommonNames_MDD	subclass_MDD	infraclass_MDD	magnorder_MDD	superorder_MDD	order_MDD	suborder_MDD	infraorder_MDD	parvorder_MDD	superfamily_MDD	family_MDD	subfamily_MDD	tribe_MDD	genus_MDD	subgenus_MDD	specificEpithet_MDD	authoritySpeciesAuthor_MDD	authoritySpeciesYear_MDD	authorityParentheses_MDD	originalNameCombination_MDD	authoritySpeciesCitation_MDD	authoritySpeciesLink_MDD	holotypeVoucher_MDD	holotypeVoucherURIs_MDD	typeLocality_MDD	typeLocalityLatitude_MDD	typeLocalityLongitude_MDD	nominalNames_MDD	taxonomyNotes_MDD	taxonomyNotesCitation_MDD	countryDistribution_MDD	continentDistribution_MDD	biogeographicRealm_MDD	iucnStatus_MDD	extinct_MDD	domestic_MDD	flagged_MDD	CMW_sciName_MDD	diffSinceCMW_MDD	MSW3_matchtype_MDD	MSW3_sciName_MDD	diffSinceMSW3_MDD	docOrigin_IUCN	internalTaxonId_IUCN	NAME_IUCN	kingdomName_IUCN	phylumName_IUCN	className_IUCN	orderName_IUCN	familyName_IUCN	genusName_IUCN	speciesName_IUCN	authoritySpeciesAuthorYear_IUCN	taxonomicNotes_IUCN	assessmentId_IUCN	scientificName_IUCN	redlistCategory_IUCN	redlistCriteria_IUCN	yearPublished_IUCN	assessmentDate_IUCN	criteriaVersion_IUCN	language_IUCN	rationale_IUCN	habitat_IUCN	threats_IUCN	population_IUCN	populationTrend_IUCN	range_IUCN	useTrade_IUCN	systems_IUCN	conservationActions_IUCN	realm_IUCN	yearLastSeen_IUCN	possiblyExtinct_IUCN	possiblyExtinctInTheWild_IUCN	scopes_IUCN	docOrigin_batnames2023	FAMILY_batnames2023	GENUS_batnames2023	SUBGENUS_batnames2023	SPECIES_batnames2023	authoritySpeciesAuthor_batnames2023	authoritySpeciesYearbatnames2023	PARENTHESES_batnames2023 (1=AUTHOR & DATE IN PARENTHESES)	CITATION_batnames2023	PAGES_batnames2023	COMMON NAME_batnames2023	SYNONYMS_batnames2023	TYPE LOCALITY_batnames2023	DISTRIBUTION_batnames2023	CITES_batnames2023	IUCN_batnames2023	COMMENTS_batnames2023	name MDD2023	id_MDD2023	phylosort_MDD2023	mainCommonName_MDD2023	otherCommonNames_MDD2023	subclass_MDD2023	infraclass_MDD2023	magnorder_MDD2023	superorder_MDD2023	order_MDD2023	suborder_MDD2023	infraorder_MDD2023	parvorder_MDD2023	superfamily_MDD2023	Family_mdd2023	subfamily_MDD2023	tribe_MDD2023	genus_MDD2023	subgenus_MDD2023	specificEpithet_MDD2023	authoritySpeciesAuthor_MDD2023	authoritySpeciesYear_MDD2023	authorityParentheses_MDD2023	originalNameCombination_MDD2023	authoritySpeciesCitation_MDD2023	authoritySpeciesLink_MDD2023	holotypeVoucher_MDD2023	holotypeVoucherURIs_MDD2023	typeLocality_MDD2023	typeLocalityLatitude_MDD2023	typeLocalityLongitude_MDD2023	nominalNames_MDD2023	taxonomyNotes_MDD2023	taxonomyNotesCitation_MDD2023	distributionNotes_MDD2023	distributionNotesCitation_MDD2023	subregionDistribution_MDD2023	countryDistribution_MDD2023	continentDistribution_MDD2023	biogeographicRealm_MDD2023	iucnStatus_MDD2023	extinct_MDD2023	domestic_MDD2023	flagged_MDD2023	CMW_sciName_MDD2023	diffSinceCMW_MDD2023	MSW3_matchtype_MDD2023	MSW3_sciName_MDD2023	diffSinceMSW3_MDD2023	docOrigin_MDD2025	sciName	id	phylosort	mainCommonName	otherCommonNames	subclass	infraclass	magnorder	superorder	order	suborder	infraorder	parvorder	superfamily	family	subfamily	tribe	genus	subgenus	specificEpithet	authoritySpeciesAuthor	authorityParentheses	originalNameCombination	authoritySpeciesCitation	authoritySpeciesLink	typeVoucher	typeKind	typeVoucherURIs	typeLocality	typeLocalityLatitude	typeLocalityLongitude	taxonomyNotes	taxonomyNotesCitation	distributionNotes	distributionNotesCitation	subregionDistribution	countryDistribution	continentDistribution	biogeographicRealm	iucnStatus	extinct	domestic	flagged	CMW_sciName	diffSinceCMW	MSW3_matchtype	MSW3_sciName	diffSinceMSW3	docOrigin_batnames2025	Family	Genus	Subgenus	Species	Author	Date	Parentheses (1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L760	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Murina cyclotis	Murina cyclotis	Murina cyclotis	Murina cyclotis	Murina cyclotis	Murina cyclotis	Murina cyclotis	Murina cyclotis	Murina cyclotis	Murina cyclotis	Murina cyclotis	Murina cyclotis	Murina cyclotis	Murina cyclotis	Murina cyclotis		[MSW2] Subgenus Murina.; [MSW3] Subgenus Murina. Reviewed in part by Hill (1983), Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b).; [HMW] Murina cyclotis Dobson, 1872 , Darjeeling, India . See M. fionae . There appears to be at least two species included in M. cyclotis based on genetic data, with specimens from southern India sister to M. guilleni and true M. cyclotis sister to M. peninsularis . Name eileenae might be applicable to this taxon, but specimens from Sri Lankan (type locality of eileenae) and northern Indian (type locality of M. cyclotis ) have not been included in any genetic studies. The two taxa are considered subspecies here, but they will likely be elevated to distinct species after additional research. Specimens from the Philippines have not yet been included in any morphological or genetic studies and are tentatively retained under M. cyclotis . There is also extreme morphological variation across South-east Asia, especially in hair color and ear shape, but it has not been investigated taxonomically. Two subspecies recognized.; [batnames2022] Subgenus Murina . Does not include peninsularis ; see Francis and Eger (2012). Reviewed by Soisook et al . (2013) and in part by Hill (1983), Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001 b ). Specimens from the Phillippines have not been allocated to cyclotis / peninsularis (S. Tsang, pers.comm., 2018).; [MDD2022] previously included M. peninsularis; [IUCN] The species, as covered here, actually represents a complex of species, only some of which have been formally recognized or described. Populations in south peninsular Thailand, peninsular Malaysia and Borneo, originally described as a large subspecies of M. cyclotis , were recognized as a distinct species under the name M. peninsularis by Francis and Eger (2012) and Soisook et al. (2013). Specimens from Peninsular Thailand, south of the Isthmus of Kra (and overlapping the range of M. peninsularis ) were recently described as a separate species M. guilleni with a closely related but distinct subspecies M. guilleni nicobarensis from the Nicobar islands (Soisook et al 2013). Populations in the Philippines include several genetically distinct clades that differ from any of the above forms (J. Eger pers. comm.) but these have not yet been formally described. The identity of the specimen from Lombok has not been studied, but it is unlikely to be the same as the mainland forms. The Sri Lankan subspecies M. cyclotis eileenae may represent another separate species, but this has not been examined genetically. Similarly, genetic data are not yet available from India (the type locality of M. cyclotis ) to determine whether these are the same species as populations in northern Thailand and Indochina. The recently described species M. fionae (Francis and Eger 2012) from Laos, Vietnam and Cambodia is morphologically similar to M. peninsularis but is genetically distinct and morphologically readily distinguished from sympatric populations of ;M. cyclotis ; by its larger size.; [batnames2023] Subgenus Murina . Does not include peninsularis ; see Francis and Eger (2012). Reviewed by Soisook et al . (2013) and in part by Hill (1983), Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001 b ). Specimens from the Phillippines have not been allocated to cyclotis / peninsularis (S. Tsang, pers.comm., 2018).; [MDD2023] previously included M. peninsularis; [MDD2025_2.0] previously included M. peninsularis; [batnames2025_1.7] Subgenus Murina. Does not include peninsularis; see Francis and Eger (2012). Reviewed by Soisook et al. (2013) and in part by Hill (1983), Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b). Specimens from the Phillippines have not been allocated to cyclotis/peninsularis (S. Tsang, pers.comm., 2018).; [MDD2025_2.2] previously included M. peninsularis						eileenae, peninsularis.	eileenae, cyclotis, peninsularis	cyclotis, eileenae, peninsularis		cyclotis, eileenae		cyclotis, eileenae		cyclotis, eileenae	The species, as covered here, actually represents a complex of species, only some of which have been formally recognized or described. Populations in south peninsular Thailand, peninsular Malaysia and Borneo, originally described as a large subspecies of M. cyclotis , were recognized as a distinct species under the name M. peninsularis by Francis and Eger (2012) and Soisook et al. (2013). Specimens from Peninsular Thailand, south of the Isthmus of Kra (and overlapping the range of M. peninsularis ) were recently described as a separate species M. guilleni with a closely related but distinct subspecies M. guilleni nicobarensis from the Nicobar islands (Soisook et al 2013). Populations in the Philippines include several genetically distinct clades that differ from any of the above forms (J. Eger pers. comm.) but these have not yet been formally described. The identity of the specimen from Lombok has not been studied, but it is unlikely to be the same as the mainland forms. The Sri Lankan subspecies M. cyclotis eileenae may represent another separate species, but this has not been examined genetically. Similarly, genetic data are not yet available from India (the type locality of M. cyclotis ) to determine whether these are the same species as populations in northern Thailand and Indochina. The recently described species M. fionae (Francis and Eger 2012) from Laos, Vietnam and Cambodia is morphologically similar to M. peninsularis but is genetically distinct and morphologically readily distinguished from sympatric populations of ;M. cyclotis ; by its larger size.	cyclotis, eileenae		cyclotis, eileenae	cyclotis, eileenae	cyclotis, eileenae		cyclotis Dobson, 1872|eileenae W. W. A. Phillips, 1932		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Round-eared tube-nosed bat	N India – S China – Malaya, Hainan, Sri Lanka, Philippines, Borneo; ref. 4.106	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Murina cyclotis	India, Darjeeling.	Dobson	1872	Proc. Asiat. Soc. Bengal, p. 210.	Distribution: Ranging from northeastern India to Hainan in southeastern China, south to Ceylon and Malaya and east through Borneo to Mindanao in the southern Philippines.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Round-eared tube-nosed bat	N India – S China – Malaya, Hainan – Borneo, Sri Lanka, (?) Philippines; ref. 4.12	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Dobson	1872	Proc. Asiat. Soc. Bengal, p. 210.	Subgenus Murina.	Sri Lanka and India to Kwangtung and Hainan (China); Vietnam, south to W Malaysia; Borneo; Philippines; Lesser Sunda Isis.	India, Darjeeling.		DOBSON	1872	Size medium (forearm length, 30-38 mm). Ears rounded. Talonids of anterior and middle lower molars and their cusps reduced. Upper canine relatively long. Teeth not greatly enlarged.	Distribution: Ranging from northeastern India to Hainan in southeastern China, south to Ceylon and Malaya and east through Borneo to Mindanao in the southern Philippines.	Three subspecies:	M. c. eileenae (Ceylon), M. c. cyclotis (northeastern India to Hainan and Vietnam), M. c. peninsularis (Malaya to Mindanao).	132	species	M. cyclotis	DOBSON	1872	Murina	subgenus	Murina cyclotis				Size medium (forearm length, 30-38 mm). Ears rounded. Talonids of anterior and middle lower molars and their cusps reduced. Upper canine relatively long. Teeth not greatly enlarged.	Three subspecies:		11. M. cyclotis DOBSON 1872 [cyclotis group],	11	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Vespertilionidae	Murininae		Murina cyclotis	Murina	Murina	cyclotis	Dobson		1872		Proc. Asiat. Soc. Bengal			210		Round-eared Tube-nosed Bat	India, Darjeeling.	Sri Lanka and India to Kwangtung and Hainan (China); Myanamar, Laos, and Vietnam, south to W Malaysia, Borneo, Sumatra, Philippines, and Lesser Sunda Isls. Records from Cambodia are erroneous (Kock, 2000a).	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	eileenae Phillips, 1932; peninsularis Hill, 1964.	Subgenus Murina. Reviewed in part by Hill (1983), Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b).	4C3D87E8FF676AD9FA8392CC1C41B91E	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	917	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF676AD9FA8392CC1C41B91E.xml	Murina cyclotis	Vespertilionidae	Murina	cyclotis	Dobson	1872	Murine a oreilles rondes @fr | Rundohrréhrennase @de | Ratonero narizudo de orejas redondas @es	Murina cyclotis Dobson, 1872 , Darjeeling, India . See M. fionae . There appears to be at least two species included in M. cyclotis based on genetic data, with specimens from southern India sister to M. guilleni and true M. cyclotis sister to M. peninsularis . Name eileenae might be applicable to this taxon, but specimens from Sri Lankan (type locality of eileenae) and northern Indian (type locality of M. cyclotis ) have not been included in any genetic studies. The two taxa are considered subspecies here, but they will likely be elevated to distinct species after additional research. Specimens from the Philippines have not yet been included in any morphological or genetic studies and are tentatively retained under M. cyclotis . There is also extreme morphological variation across South-east Asia, especially in hair color and ear shape, but it has not been investigated taxonomically. Two subspecies recognized.	M.c.cyclotisDobson,1872—CNepal,NE&ECIndia(Sikkim,WestBengal,Megha-laya,Mizoram,andAndhraPradesh),SEChina(Jiangxi,Guangxi,andSHainan),SEAsia(includingConSonandCatBaIsandtheBaiTuLongArchipelago),andthePhilippines(Luzon,Catanduanes,Sibuyan,Biliran,Cebu,Bohol,Siquijor,Camiguin,Mindanao,andPalawanIs). M. c. eileenae W. W. A Phillips, 1932 — Sri Lanka and tentatively S India ( Tamil Nadu ).	Head—body 38-50 mm ,tail 26-2-43- 5 mm , ear 12-18- 6 mm , hindfoot 6-3-9- 7 mm , forearm 28-9-36- 8 mm ; weight 4-1-8- 2 g . Females average larger in bodysize than males. Fur silky. Dorsal pelage is orangish brown (hairs with dark gray bases and orange-brown tips); venter is grayish white (hairs with gray bases and light gray or whitish brown tips) or with light orange-brown tinge on some individuals. Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet. Faceis sparsely haired except for long protuberant naked nostrils. Ear shape is variable from broadly rounded with convex anterior margins to narrower with somewhat more elliptical and less convex anterior margins, but never with notch on posterior margins; tragusis long and narrow and tapers toward pointed tip. Wing attaches near base of claw on first toe. Baculum is very small ( 0-8 mm long), arched upward, deeply concave ventrally, and almost round, with W-shaped indentation at tip and distinct concavity at base. Skull has relatively long and non-inflated rostrum and slightly domed braincase (more domed in males); sagittal and lambdoidal crests are poorly developed; C! is taller than P*, with basal area that is subequal to or slightly larger than P* P? is two-thirds the basal area of P* but two-thirds to slightly shorter in height; M' and M? lack mesostyles and have Ushaped indentation on labial sides; and talonids of M, and M, are more than one-half to two-thirds the size of their respective trigonids. Chromosomal complement has 2n = 44 and FN = 50 ( Vietnam ).	Various forest habitats such as lowland, wet, and hill evergreen forests; mixed deciduousforest; and dry dipterocarp forest from sea level up to elevations of¢. 1650 m .	Round-eared Tube-nosed Bats reportedly fly low to the ground and likely forage in vegetation clutter.	Young Round-eared Tube-nosed Bats were collected in August in West Bengal and July in Meghalaya , India .	The Round-eared Tube-nosed Bat roosts primarily in foliage, but some roosts have been found in small rock caves. Calls are steep FM sweeps, with start frequencies of 141-163 kHz, end frequencies of 56-72 kHz, peak frequencies of 96-3-109 kHz, and durations of 1-5-2-3 milliseconds in Thailand .	The Round-eared Tube-nosed Bat roosts alone or in groups up to only three individuals.	Classified as Least Concern on The [UCN Red List. The Roundeared Tube-nosed Batis widespread and does not seem to face any major threats.	Bates & Harrison (1997) | Bates, Francis, Rosell-Ambal, Tabaranza et al. (2008) | Corbet & Hill (1992) | Francis (2008a) | Heaney, Balete, Dolar et al. (1998) | Heaney, Balete & Rickart (2016) | Heaney, Gonzales et al. (1991) | Hill (1964) | Lepiten (1995) | Nguyen Truong Son et al. (2015) | Rickart et al. (1991) | Smith & Xie Yan (2008) | Soisook (2013) | Soisook, Karapan, Satasook, Thong Vu Dinh et al. (2013) | Soisook, Thaw Win-Naing et al. (2017) | Thong Vu Dinh et al. (2010)	https://zenodo.org/record/6398614/files/figure.png	356. Round-eared Tube-nosed Bat Murina cyclotis French: Murine a oreilles rondes / German: Rundohrréhrennase / Spanish: Ratonero narizudo de orejas redondas Taxonomy. Murina cyclotis Dobson, 1872 , Darjeeling, India . See M. fionae . There appears to be at least two species included in M. cyclotis based on genetic data, with specimens from southern India sister to M. guilleni and true M. cyclotis sister to M. peninsularis . Name eileenae might be applicable to this taxon, but specimens from Sri Lankan (type locality of eileenae) and northern Indian (type locality of M. cyclotis ) have not been included in any genetic studies. The two taxa are considered subspecies here, but they will likely be elevated to distinct species after additional research. Specimens from the Philippines have not yet been included in any morphological or genetic studies and are tentatively retained under M. cyclotis . There is also extreme morphological variation across South-east Asia, especially in hair color and ear shape, but it has not been investigated taxonomically. Two subspecies recognized. Subspecies and Distribution. M.c.cyclotisDobson,1872—CNepal,NE&ECIndia(Sikkim,WestBengal,Megha-laya,Mizoram,andAndhraPradesh),SEChina(Jiangxi,Guangxi,andSHainan),SEAsia(includingConSonandCatBaIsandtheBaiTuLongArchipelago),andthePhilippines(Luzon,Catanduanes,Sibuyan,Biliran,Cebu,Bohol,Siquijor,Camiguin,Mindanao,andPalawanIs). M. c. eileenae W. W. A Phillips, 1932 — Sri Lanka and tentatively S India ( Tamil Nadu ). Descriptive notes. Head—body 38-50 mm ,tail 26-2-43- 5 mm , ear 12-18- 6 mm , hindfoot 6-3-9- 7 mm , forearm 28-9-36- 8 mm ; weight 4-1-8- 2 g . Females average larger in bodysize than males. Fur silky. Dorsal pelage is orangish brown (hairs with dark gray bases and orange-brown tips); venter is grayish white (hairs with gray bases and light gray or whitish brown tips) or with light orange-brown tinge on some individuals. Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet. Faceis sparsely haired except for long protuberant naked nostrils. Ear shape is variable from broadly rounded with convex anterior margins to narrower with somewhat more elliptical and less convex anterior margins, but never with notch on posterior margins; tragusis long and narrow and tapers toward pointed tip. Wing attaches near base of claw on first toe. Baculum is very small ( 0-8 mm long), arched upward, deeply concave ventrally, and almost round, with W-shaped indentation at tip and distinct concavity at base. Skull has relatively long and non-inflated rostrum and slightly domed braincase (more domed in males); sagittal and lambdoidal crests are poorly developed; C! is taller than P*, with basal area that is subequal to or slightly larger than P* P? is two-thirds the basal area of P* but two-thirds to slightly shorter in height; M' and M? lack mesostyles and have Ushaped indentation on labial sides; and talonids of M, and M, are more than one-half to two-thirds the size of their respective trigonids. Chromosomal complement has 2n = 44 and FN = 50 ( Vietnam ). Habitat. Various forest habitats such as lowland, wet, and hill evergreen forests; mixed deciduousforest; and dry dipterocarp forest from sea level up to elevations of¢. 1650 m . Food and Feeding. Round-eared Tube-nosed Bats reportedly fly low to the ground and likely forage in vegetation clutter. Breeding. Young Round-eared Tube-nosed Bats were collected in August in West Bengal and July in Meghalaya , India . Activity patterns. The Round-eared Tube-nosed Bat roosts primarily in foliage, but some roosts have been found in small rock caves. Calls are steep FM sweeps, with start frequencies of 141-163 kHz, end frequencies of 56-72 kHz, peak frequencies of 96-3-109 kHz, and durations of 1-5-2-3 milliseconds in Thailand . Movements, Home range and Social organization. The Round-eared Tube-nosed Bat roosts alone or in groups up to only three individuals. Status and Conservation. Classified as Least Concern on The [UCN Red List. The Roundeared Tube-nosed Batis widespread and does not seem to face any major threats. Bibliography. Bates & Harrison (1997), Bates, Francis, Rosell-Ambal, Tabaranza et al. (2008), Corbet & Hill (1992), Francis (2008a), Heaney, Balete, Dolar et al. (1998), Heaney, Balete & Rickart (2016), Heaney, Gonzales et al. (1991), Hill (1964), Lepiten (1995), Nguyen Truong Son et al. (2015), Rickart et al. (1991), Smith & Xie Yan (2008), Soisook (2013), Soisook, Karapan, Satasook, Thong Vu Dinh et al. (2013), Soisook, Thaw Win-Naing et al. (2017), Thong Vu Dinh et al. (2010).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Murina cyclotis	Murina	Murina	cyclotis	Dobson	1872	0	Proc. Asiat. Soc. Bengal	p. 210	Round-eared Tube-nosed Bat	<b> eileenae </b>Phillips, 1932	India, Darjeeling.	Sri Lanka and India to Kwangtung and Hainan (China); Myanamar, Laos, and Vietnam, south to W Malaysia, Borneo, Sumatra, Philippines, and Lesser Sunda Isls. Records from Cambodia are erroneous (Kock, 2000a).	Not listed.	Least Concern	Subgenus Murina . Does not include peninsularis ; see Francis and Eger (2012). Reviewed by Soisook et al . (2013) and in part by Hill (1983), Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001 b ). Specimens from the Phillippines have not been allocated to cyclotis / peninsularis (S. Tsang, pers.comm., 2018).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Murina cyclotis	23	Round-eared Tube-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MURININAE	NA	Murina	NA	cyclotis	Dobson	1872	0	Murina_cyclotis	Dobson, G. E. (1872). Notes on some bats collected by Captain W. G. Murray, in the North-western Himalaya, with description of new species. Proceedings of the Asiatic Society of Bengal, 1872, 210.	https://www.biodiversitylibrary.org/item/111587#page/224/mode/1up	BM 1909.4.4.4 [syntype]		Darjeeling, India.			cyclotis Dobson, 1872|eileenae W. W. A. Phillips, 1932	previously included M. peninsularis	Francis, C. M., & Eger, J. L. (2012). A review of tube-nosed bats (Murina) from Laos with a description of two new species. Acta Chiropterologica, 14(1), 15-38.	India|Nepal|Sri Lanka|China|Myanmar|Vietnam|Cambodia|Thailand|Laos|Philippines	Asia	Indomalaya	LC	0	0	0	Murina_cyclotis	0	sciname match	Murina_cyclotis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	200000000	Murina cyclotis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Murina	cyclotis	Dobson, 1872	The species, as covered here, actually represents a complex of species, only some of which have been formally recognized or described. Populations in south peninsular Thailand, peninsular Malaysia and Borneo, originally described as a large subspecies of M. cyclotis , were recognized as a distinct species under the name M. peninsularis by Francis and Eger (2012) and Soisook et al. (2013). Specimens from Peninsular Thailand, south of the Isthmus of Kra (and overlapping the range of M. peninsularis ) were recently described as a separate species M. guilleni with a closely related but distinct subspecies M. guilleni nicobarensis from the Nicobar islands (Soisook et al 2013). Populations in the Philippines include several genetically distinct clades that differ from any of the above forms (J. Eger pers. comm.) but these have not yet been formally described. The identity of the specimen from Lombok has not been studied, but it is unlikely to be the same as the mainland forms. The Sri Lankan subspecies M. cyclotis eileenae may represent another separate species, but this has not been examined genetically. Similarly, genetic data are not yet available from India (the type locality of M. cyclotis ) to determine whether these are the same species as populations in northern Thailand and Indochina. The recently described species M. fionae (Francis and Eger 2012) from Laos, Vietnam and Cambodia is morphologically similar to M. peninsularis but is genetically distinct and morphologically readily distinguished from sympatric populations of ;M. cyclotis ; by its larger size.	20000000	Murina cyclotis	Least Concern		2020	2019-06-06 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, it occurs in a number of protected areas, has a tolerance of a degree of habitat modification, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	This species is a forest dweller. In South India it roosts among leaves of cardamom plantations in forested tracts and in caves. It roosts in small colonies of two to five individuals (Molur et al. 2002). It hunts for small insects and is a low and agile flyer (Bates and Harrison 1997, Aul and Vijayakumar 2003). In China, it is found in agroforestry areas at intermediate elevations. They roost in foliage, small caves or rock shelters (Smith and Xie 2008). In the Philippines, this species is found in primary lowland forest, lightly disturbed lowland and lower montane forest (Heaney et al. 1991, Lepiten 1995, Rickart et al. 1991, Ruedas et al. 1994). It has also been recorded from secondary forest elsewhere (G. Csorba pers. comm.).	The threats to this species are not well known, although it is unlikely to be threatened throughout its range. Although it is definitely threatened by deforestation (generally resulting from logging operations and the conversion of land to agricultural and other uses), it has been recorded from some modified habitats and seems to be somewhat adaptable in parts of its range.	In South Asia, the abundance, population size and trends for this species are not known (Molur et al. 2002). It is moderately common in the Philippines (Heaney et al. 1998) and is one of the most commonly caught species of tube-nosed bats in the Indo-Burma subregion.	Unknown	This widespread species ranges from southern and northern South Asia, into southern China, most of mainland Southeast Asia, and parts of insular Southeast Asia. In South Asia the species is presently known form India (Andhra Pradesh, Meghalaya, Mizoram, Nicobar Islands, Sikkim, Tamil Nadu and West Bengal), Nepal (Western Nepal) and Sri Lanka (Central Province) in South Asia (Aul and Vijaykumar 2003, Molur et al. 2002, Vanitharani et al. 2005). In South Asia, it has been recorded from 250 up to an elevation of 1,500 m asl (Molur et al. 2002). In China, it ranges in Jiangxi and Hainan, extending to Zhongnan peninsula (Smith and Xie 2008). It is present through nearly all of mainland Southeast Asia and the Philippines. In the Philippines, it has been recorded from Biliran, Camiguin, Catanduanes, Luzon (Camarines Sur province), Mindanao (Bukidnon Province), Sibuyan, and Siqujor (Heaney et al. 1998), Balbalasang, Kalinga Province (Heaney et al. 2004) and has an elevation range from 250 to 1,500 m asl (Heaney et al. 1998). It has also been recorded from Samar using a tunnel trap at 100 m asl (Gonzales unpublished data 2004), from Negros (Cariño unpublished data 2004), Polillo, and on Panay along a river at 1,140 m asl (Ramayla and Garcia pers. comm. 2006).		Terrestrial	It is present in a number of protected areas throughout its range.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Murina	Murina	cyclotis	Dobson	1872	0	Proc. Asiat. Soc. Bengal	p. 210	Round-eared Tube-nosed Bat	<b> eileenae </b>Phillips, 1932	India, Darjeeling.	Sri Lanka and India to Kwangtung and Hainan (China); Myanamar, Laos, and Vietnam, south to W Malaysia, Borneo, Sumatra, Philippines, and Lesser Sunda Isls. Records from Cambodia are erroneous (Kock, 2000a).	Not listed.	Least Concern	Subgenus Murina . Does not include peninsularis ; see Francis and Eger (2012). Reviewed by Soisook et al . (2013) and in part by Hill (1983), Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001 b ). Specimens from the Phillippines have not been allocated to cyclotis / peninsularis (S. Tsang, pers.comm., 2018).	Murina cyclotis	1005324	23	Round-eared Tube-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MURININAE	NA	Murina	NA	cyclotis	Dobson	1872	0	Murina_cyclotis	Dobson, G. E. (1872). Notes on some bats collected by Captain W. G. Murray, in the North-western Himalaya, with description of new species. Proceedings of the Asiatic Society of Bengal, 1872, 210.	https://www.biodiversitylibrary.org/item/111587#page/224/mode/1up	BM 1909.4.4.4 [syntype]		Darjeeling, India.			cyclotis Dobson, 1872|eileenae W. W. A. Phillips, 1932	previously included M. peninsularis	Francis, C. M., & Eger, J. L. (2012). A review of tube-nosed bats (Murina) from Laos with a description of two new species. Acta Chiropterologica, 14(1), 15-38.				India|Nepal|Sri Lanka|China|Myanmar|Vietnam|Cambodia|Thailand|Laos|Philippines	Asia	Indomalaya	LC	0	0	0	Murina_cyclotis	0	sciname match	Murina_cyclotis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Murina_cyclotis	1005324	23	Round-eared Tube-nosed Bat		Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Murininae	NA	Murina	NA	cyclotis	Dobson	0	Murina cyclotis	Dobson, G.E. 1872. Notes on some bats collected by Captain W. G. Murray, in the North-Western Himalaya, with description of new species. Proceedings of the Asiatic Society of Bengal 1872(10):208-210.	https://www.biodiversitylibrary.org/page/36180073	BMNH:Mamm:1909.4.4.4, ZSI 15605	syntypes	https://data.nhm.ac.uk/object/b3dce104-e878-4af0-800f-12e4c727531f	Darjeeling, India.			previously included M. peninsularis	Francis, C. M., & Eger, J. L. (2012). A review of tube-nosed bats (Murina) from Laos with a description of two new species. Acta Chiropterologica, 14(1), 15-38.				India|Nepal|Sri Lanka|China|Myanmar|Vietnam|Cambodia|Thailand|Laos|Philippines	Asia	Indomalaya	LC	0	0	0	Murina_cyclotis	0	sciname match	Murina_cyclotis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Murina	Murina	cyclotis	Dobson	1872	0	Proc. Asiat. Soc. Bengal	p. 210	Round-eared Tube-nosed Bat	eileenae Phillips, 1932	India, Darjeeling.	Sri Lanka and India to Kwangtung and Hainan (China); Myanamar, Laos, and Vietnam, south to W Malaysia, Borneo, Sumatra, Philippines, and Lesser Sunda Isls. Records from Cambodia are erroneous (Kock, 2000a).	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/154196798/22094685/' target='_blank'>Least Concern</a>	Subgenus Murina. Does not include peninsularis; see Francis and Eger (2012). Reviewed by Soisook et al. (2013) and in part by Hill (1983), Bates and Harrison (1997), Sinha (1999), and Hendrichsen et al. (2001b). Specimens from the Phillippines have not been allocated to cyclotis/peninsularis (S. Tsang, pers.comm., 2018).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Murina cyclotis; Murina cyclotis; Murina cyclotis; Murina cyclotis; Murina cyclotis; Murina cyclotis; cyclotis; eileenae; peninsularis; cyclotis; eileenae; eileenae; cyclotis; eileenae; Murine a oreilles rondes; Rundohrréhrennase; Ratonero narizudo de orejas redondas; Round-eared Tube-nosed Bat; Round-eared Tube-nosed Bat; Round-eared Tube-nosed Bat; M. cyclotis