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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L755	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Murina beelzebub	Murina beelzebub	Murina beelzebub	Murina beelzebub	Murina beelzebub	Murina beelzebub	Murina beelzebub	Murina beelzebub	Murina beelzebub		[HMW] Murina beelzebub Nguyen Truong Son, Furey & Csorba, 2011 , “ Vietnam , Quang Tri Province , Huong Hoa District, Bac Huong Hoa Nature Reserve, centered at 16°56'15"N , 106°34'52"E , elevation 400m .” See M. feae . Separated from M. tubinaris on genetic data and morphology. Monotypic.; [MDD2022] recently described; [IUCN] <p><span lang="EN-US">The species was described in 2011 based on specimens from Viet Nam.</p>; [MDD2023] recently described; [MDD2025_2.0] recently described; [MDD2025_2.2] recently described														beelzebub	<p><span lang="EN-US">The species was described in 2011 based on specimens from Viet Nam.</p>			beelzebub	beelzebub			beelzebub Nguyen Truong Son, Furey, & Csorba in Csorba, Nguyen Truong Son, Saveng, & Furey, 2011						N/A																																								NA																											4C3D87E8FF6D6AD2FF8F906B1A01B76C	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Vespertilionidae_716.pdf.imf	hash://md5/b004ff90fffb6a44fffc96591e00bb32	911	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/4C/3D/87/4C3D87E8FF6D6AD2FF8F906B1A01B76C.xml	Murina beelzebub	Vespertilionidae	Murina	beelzebub	Nguyen Truong Son, Furey & Csorba	2011	Murine belzébuth @fr | Beelzebub-Rohrennase @de | Ratonero narizudo de belzebu @es | Diabolic Tube-nosed Bat @en	Murina beelzebub Nguyen Truong Son, Furey & Csorba, 2011 , “ Vietnam , Quang Tri Province , Huong Hoa District, Bac Huong Hoa Nature Reserve, centered at 16°56'15"N , 106°34'52"E , elevation 400m .” See M. feae . Separated from M. tubinaris on genetic data and morphology. Monotypic.	Known from four localities in C & SC Vietnam (Bac Huong Hoa and Ngoc Linh nature reserves, Ba To, and Kon Ka Kinh National Park).	Head-body 40-49 mm , tail 33-44- 8 mm , ear 13-2-14- 2 mm , hindfoot 5- 5-8 mm , forearm 34-4-37- 3 mm ; weight 5- 5 g . Pelage is long and silky; dorsally very dark brown (hairs very dark brown, almost black, for basal four-fifths, then light gray fading to darker tip, with longer silvery guard hairs mixed throughout); ventrally grayish white (hairs very dark brown, nearly black, for basal two-thirds, then white to tip). Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet; ventral pelage extends sparsely onto uropatagium and a little onto wings. Faceis sparsely haired except long protuberant nostrils, which are naked. Ears are short and broad with smoothly convex anterior margins, slight notch on posterior margin, and broadly rounded tip; tragus is long, narrow, and tapering toward pointed tip. Wing attaches to base of claw on first toe. Skull has relatively inflated rostrum and uninflated braincase; sagittal crest is lacking, and lambdoidal crests relatively well developed. C' is shorter than P*; P* is relatively short and small at about half the crown area and slightly more than half the height of P*; mesostyle of M' and M”is greatly reduced, both having a flat labial surface; talonid of M, and M, is equal in size to respective trigonid. Chromosomal complement has 2n = 44 and FN = 50.	Collected on forest trails near a stream in disturbed secondary forest, and on a ridgetop in primary montane forest; at elevations of 400-1600 m .	No information.	No information.	No information.	No information.	Not assessed on The IUCN RedLust, as it is included in Scully’s Tube-nosed Bat ( M. tubinaris ), which is listed as Least Concern. Currently known only from four localities in Vietnam .	Csorba et al. (2011) | Nguyen Truong Son et al. (2015) | Soisook (2013)	https://zenodo.org/record/6398578/files/figure.png	338. Beelzebub Tube-nosed Bat Murina beelzebub French: Murine belzébuth / German: Beelzebub-Rohrennase / Spanish: Ratonero narizudo de belzebu Other common names: Diabolic Tube-nosed Bat Taxonomy. Murina beelzebub Nguyen Truong Son, Furey & Csorba, 2011 , “ Vietnam , Quang Tri Province , Huong Hoa District, Bac Huong Hoa Nature Reserve, centered at 16°56'15"N , 106°34'52"E , elevation 400m .” See M. feae . Separated from M. tubinaris on genetic data and morphology. Monotypic. Distribution. Known from four localities in C & SC Vietnam (Bac Huong Hoa and Ngoc Linh nature reserves, Ba To, and Kon Ka Kinh National Park). Descriptive notes. Head-body 40-49 mm , tail 33-44- 8 mm , ear 13-2-14- 2 mm , hindfoot 5- 5-8 mm , forearm 34-4-37- 3 mm ; weight 5- 5 g . Pelage is long and silky; dorsally very dark brown (hairs very dark brown, almost black, for basal four-fifths, then light gray fading to darker tip, with longer silvery guard hairs mixed throughout); ventrally grayish white (hairs very dark brown, nearly black, for basal two-thirds, then white to tip). Dorsal pelage extends sparsely onto wings, uropatagium, thumbs, and feet; ventral pelage extends sparsely onto uropatagium and a little onto wings. Faceis sparsely haired except long protuberant nostrils, which are naked. Ears are short and broad with smoothly convex anterior margins, slight notch on posterior margin, and broadly rounded tip; tragus is long, narrow, and tapering toward pointed tip. Wing attaches to base of claw on first toe. Skull has relatively inflated rostrum and uninflated braincase; sagittal crest is lacking, and lambdoidal crests relatively well developed. C' is shorter than P*; P* is relatively short and small at about half the crown area and slightly more than half the height of P*; mesostyle of M' and M”is greatly reduced, both having a flat labial surface; talonid of M, and M, is equal in size to respective trigonid. Chromosomal complement has 2n = 44 and FN = 50. Habitat. Collected on forest trails near a stream in disturbed secondary forest, and on a ridgetop in primary montane forest; at elevations of 400-1600 m . Food and Feeding. No information. Breeding. No information. Activity patterns. No information. Movements, Home range and Social organization. No information. Status and Conservation. Not assessed on The IUCN RedLust, as it is included in Scully’s Tube-nosed Bat ( M. tubinaris ), which is listed as Least Concern. Currently known only from four localities in Vietnam . Bibliography. Csorba et al. (2011), Nguyen Truong Son et al. (2015), Soisook (2013).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Vespertilionidae	Murina beelzebub	Murina		beelzebub	Son, Furey, & Csorba	2011	0	J. Mammal.	92(4): 899	Diabolic Tube-nosed Bat	None.	Vietnam, Quang Tri Province, Huong Hoa District, Bac Huong Hoa Nature Reserve (16 degrees 56’15”N, 106 degrees34’52”E, elevation 400m)	Central Vietnam	Not listed.	Data Deficient		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Murina beelzebub	23	Beelzebub Tube-nosed Bat	Diabolic Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	VESPERTILIONIDAE	MURININAE	NA	Murina	NA	beelzebub	Nguyen Truong Son, Furey, & Csorba in Csorba, Nguyen Truong Son, Ith Saveng, & Furey	2011	0	Murina_beelzebub	Csorba, G., Son, N. T., Saveng, I., & Furey, N. M. (2011). Revealing cryptic bat diversity: three new Murina and redescription of M. tubinaris from Southeast Asia. Journal of Mammalogy, 92(4), 899.	https://academic.oup.com/jmammal/article/92/4/891/890104	HNHM 2007.50.24		"Vietnam, Quang Tri Province, Huong Hoa District, Bac Huong Hoa Nature Reserve, centered at 16°56'15"N, 106°34'52"E, elevation 400 m."	16.938	106.58	beelzebub Nguyen Truong Son, Furey, & Csorba in Csorba, Nguyen Truong Son, Ith Saveng, & Furey, 2011	recently described	Csorba, G., Son, N. T., Saveng, I., & Furey, N. M. (2011). Revealing cryptic bat diversity: three new Murina and redescription of M. tubinaris from Southeast Asia. Journal of Mammalogy, 92(4), 891-904.	Vietnam	Asia	Indomalaya	DD	0	0	0	Murina_beelzebub	0	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	80000000	Murina beelzebub	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	VESPERTILIONIDAE	Murina	beelzebub	Son, Furey &; Csorba, 2011	<p><span lang="EN-US">The species was described in 2011 based on specimens from Viet Nam.</p>	80000000	Murina beelzebub	Data Deficient		2019	2016-08-21 00:00:00 UTC	3.1	English	<p><span lang="EN-US">This species is listed as Data Deficient as it is known only from a few specimens and very few informations is available on its distribution, abundance, ecology or threats.</p>	<p><span lang="HU">At Bac Huong Hoa Nature Reserve the type series was collected in harp traps set on forest trails near a stream in disturbed secondary forest at an elevation of 400 m asl. At Kon Ka Kinh Nature Reserve <span lang="HU">M. beelzebub <span lang="HU"> <span lang="HU">was collected on a ridge top in primary montane forest at an elevation of 1,600 m (Csorba et al. 2011). </span></span></p>	<p><span lang="EN-US">This species was captured in forests, therefore logging may constitute a threat.<span lang="EN-US"></span></p>	<p><span lang="EN-US">The species is known from only a handful of specimens from Viet Nam.</p>	Unknown	<p><span lang="EN-US">The species was reported only from Viet Nam. Csorba et al . (2011) described the species based on specimens collected in Bac Huong Hoa Nature Reserve (Quang Tri) and Kon Ka Kinh Nature Reserve (Gia Lai). Son et al. (2015) report the species from further two Vietnamese locations: Ngoc Linh Nature Reserve (Kon Tum) and Ba To area (Quang Ngai).</p>		Terrestrial	Research into the species distribution, abundance, ecology and threats is needed.	Indomalayan		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Vespertilionidae	Murina		beelzebub	Son, Furey, & Csorba	2011	0	J. Mammal.	92(4): 899	Diabolic Tube-nosed Bat	None.	Vietnam, Quang Tri Province, Huong Hoa District, Bac Huong Hoa Nature Reserve (16 degrees 56’15”N, 106 degrees34’52”E, elevation 400m)	Central Vietnam	Not listed.	Data Deficient		Murina beelzebub	1005321	23	Beelzebub Tube-nosed Bat	Diabolic Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Vespertilionidae	MURININAE	NA	Murina	NA	beelzebub	Nguyen Truong Son, Furey, & Csorba in Csorba, Nguyen Truong Son, Ith Saveng, & Furey	2011	0	Murina_beelzebub	Csorba, G., Son, N. T., Saveng, I., & Furey, N. M. (2011). Revealing cryptic bat diversity: three new Murina and redescription of M. tubinaris from Southeast Asia. Journal of Mammalogy, 92(4), 899.	https://academic.oup.com/jmammal/article/92/4/891/890104	HNHM 2007.50.24		"Vietnam, Quang Tri Province, Huong Hoa District, Bac Huong Hoa Nature Reserve, centered at 16°56'15"N, 106°34'52"E, elevation 400 m."	16.9375	106.5811	beelzebub Nguyen Truong Son, Furey, & Csorba in Csorba, Nguyen Truong Son, Ith Saveng, & Furey, 2011	recently described	Csorba, G., Son, N. T., Saveng, I., & Furey, N. M. (2011). Revealing cryptic bat diversity: three new Murina and redescription of M. tubinaris from Southeast Asia. Journal of Mammalogy, 92(4), 891-904.				Vietnam	Asia	Indomalaya	DD	0	0	0	Murina_beelzebub	0	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Murina_beelzebub	1005321	23	Beelzebub Tube-nosed Bat	Diabolic Tube-nosed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Vespertilionidae	Murininae	NA	Murina	NA	beelzebub	Nguyen Truong Son, Furey, & Csorba in Csorba, Nguyen Truong Son, Saveng, & Furey	0	Murina beelzebub	Csorba, G., Son, N.T., Saveng, I. and Furey, N.M. 2011-08-16. Revealing cryptic bat diversity: three new _Murina_ and redescription of _M. tubinaris_ from Southeast Asia. Journal of Mammalogy 92(4):891-904.	https://doi.org/10.1644/10-MAMM-A-269.1	HNHM 2007.50.24	holotype		"Vietnam, Quang Tri Province, Huong Hoa District, Bac Huong Hoa Nature Reserve, centered at 16°56'15"N, 106°34'52"E, elevation 400 m."	16.9375	106.5811	recently described	Csorba, G., Son, N. T., Saveng, I., & Furey, N. M. (2011). Revealing cryptic bat diversity: three new Murina and redescription of M. tubinaris from Southeast Asia. Journal of Mammalogy, 92(4), 891-904.				Vietnam	Asia	Indomalaya	DD	0	0	0	Murina_beelzebub	0	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Vespertilionidae	Murina		beelzebub	Son, Furey & Csorba	2011	0	J. Mammal.	92(4): 899	Diabolic Tube-nosed Bat	None.	Vietnam, Quang Tri Province, Huong Hoa District, Bac Huong Hoa Nature Reserve (16 degrees 56’15”N, 106 degrees34’52”E, elevation 400m)	Central Vietnam	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/84488085/84488093/' target='_blank'>Data Deficient</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Murina beelzebub; Murina beelzebub; Murina beelzebub; Murina beelzebub; Murina beelzebub; beelzebub; Murine belzébuth; Beelzebub-Rohrennase; Ratonero narizudo de belzebu; Diabolic Tube-nosed Bat; Beelzebub Tube-nosed Bat; Diabolic Tube-nosed Bat; Diabolic Tube-nosed Bat; M. beelzebub