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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L746	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Tadarida jugularis	Mormopterus jugularis	Mormopterus jugularis	Mormopterus jugularis	Mormopterus jugularis	Mormopterus jugularis	Mormopterus jugularis	Mormopterus jugularis	Mormopterus jugularis	Mormopterus jugularis	Mormopterus jugularis	Mormopterus jugularis	Mormopterus jugularis	Mormopterus jugularis	Mormopterus jugularis		[MSW2] Subgenus Mormopterus.; [MSW3] acetabulosus species group. Reviewed by Peterson et al. (1995).; [HMW] Nyctinomus (Mormopterus) jugularis Peters in Sclater, 1865, “ Antananarivo ,” Madagascar . This species is monotypic.; [batnames2022]  acetabulosus species group. Reviewed by Peterson et al. (1995).; [batnames2023]  acetabulosus species group. Reviewed by Peterson et al. (1995).; [batnames2025_1.7] acetabulosusspecies group. Reviewed by Peterson et al. (1995).						albiventer.			albiventer			jugularis 	jugularis - albiventer	jugularis, albiventer		jugularis 	jugularis - albiventer	jugularis, albiventer	jugularis, albiventer	jugularis 	jugularis - albiventer	jugularis (W. C. H. Peters in P. L. Sclater, 1865)|albiventer (Dobson, 1877)		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Peters' wrinkle-lipped	Madagascar bat	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Mormopterus jugularis	Madagascar, Antananarivo.	Peters	1865	In Sclater, Proc. Zool. Soc. Lond., 1865:468.	Distribution: Confined to Madagascar.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Peters' wrinkle-lipped bat	Madagascar	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Peters	1865	In Sclater, Proc. Zool. Soc. Lond., 1865:468.	Subgenus Mormopterus.	Madagascar.	Madagascar, Antananarivo.		PETERS	1865	Size fairly large (forearm length, 36-39 mm). No emargination in anterior margin of ear pinna, which is well separated from the ear of the opposite side. Rostrum medium in width with only moderately developed lacrimal tubercles.	Distribution: Confined to Madagascar.	No subspecies.		136	species	M. jugularis	PETERS	1865	Mormopterus	subgenus	Mormopterus jugularis				Size fairly large (forearm length, 36-39 mm). No emargination in anterior margin of ear pinna, which is well separated from the ear of the opposite side. Rostrum medium in width with only moderately developed lacrimal tubercles.	No subspecies.		8. M.jugularis (PETERS 1865 [acetabulosus group].	8	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Molossidae	Molossinae		Mormopterus jugularis	Mormopterus		jugularis	Peters	y	1865		In Sclater, Proc. Zool. Soc. Lond.	1865		468		Peters's Wrinkle-lipped Bat	Madagascar, Tananarive (= Antananarivo).	Madagascar.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.	albiventer Dobson, 1877.	acetabulosus species group. Reviewed by Peterson et al. (1995).	194287C9FFBFBA13B4BEF12ABBC9FE44	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Molossidae_598.pdf.imf	hash://md5/e57bffb1ffbcba10b412f760b226ffce	622	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/19/42/87/194287C9FFBFBA13B4BEF12ABBC9FE44.xml	Mormopterus jugularis	Molossidae	Mormopterus	jugularis		1865	Molosse de Madagascar @fr | Peters-Mastino-Fledermaus @de | Morméptero de Madagascar @es | Peters’s Goblin Bat @en | Peters’s Little Free-tailed Bat @en | Peters’s Wrinkle-lipped Bat @en	Nyctinomus (Mormopterus) jugularis Peters in Sclater, 1865, “ Antananarivo ,” Madagascar . This species is monotypic.	Madagascar except most W region.	Head—body 60-70 mm, tail 20-38 mm, ear 14-17 mm, hindfoot 5-6 mm, forearm 30-39 mm; weight 8: 5—17 g . Males are notably larger than females in some external and cranial measurements. Peters’s Little Mastiff Bat has flattened head, no facial ornamentation, and tail protruding beyond hind margin of uropatagium. Fur is short, soft, and uniformly brown to reddish brown above, becoming pale brown to dirty white on abdomen and throat. Wings and uropatagium are dark brown. Ears are small, subtriangular, and separated at bases, without complex folds or sickle-shaped emargination below tip. Latter feature allows distinction of Peters’s Little Mastiff Bat from the Reunion Little Mastiff Bat (M. francoismoutour) and the Mauritian Little Mastiff Bat ( M. acetabulosus ). Tragus is small but not concealed by antitragus. Adult males have distinct throat gland that is absent in females. Upper lips have several wrinkles, a few spoon-hairs, and many fine hairs. Hindclaws have fine hairs and many stout bristles. It has greater skull lengths (15-4=17-2 mm) than the Reunion Little Mastiff Bat (14-1-15-3 mm). Braincase is low but not particularly flattened, and anterior palatal emargination is wide. M* has third ridge equal or longer than second ridge. Dental formulais11/3,C1/1,P1/2,M 3/3 (x2) = 30. Chromosomal complement has 2n = 48 and FN = 54.	Wide variety of bioclimatic zones from humid to arid forests, open woodlands, and degraded habitats from sea levelto elevations of ¢. 1750 m .	In eastern Madagascar , Peters’s Little Mastiff Bats ate Coleoptera (c.61% by volume), Hemiptera (c.20%), Lepidoptera (c.11%), and Diptera (c.3%).	In late January 2001, a nursery colony with more than 1000 young Peters’s Little Mastiff Bats was observed in Ankarana Special Reserve, northern Madagascar .	Peters’s Little Mastiff Bats are nocturnal. They roost in association with human habitations but also in natural settings such as crevices, roofs of caves, and cliff faces. They have narrowband,relatively long duration echolocation calls with average peak frequency of 32 kHz. Predators include common barn-owls (7yto alba).	Peters’s Little Mastiff Bats are colonial, living in groups of usually less than 100 individuals, but up to 22,000 individuals were reported in Cathedral Cave in Ankarana Special Reserve, Madagascar .	Classified as Least Concern on The IUCNRed List. Cave colonies of Peters’s Little Mastiff Bat might be threatened by unsustainable hunting practices (e.g. an estimated 70,000 -140,000 insectivorous bats, including Peters’s Little Mastiff Bats, were removed from caves annually in one region of south-western Madagascar ), but its ability to readily colonize human habitations means thatit is not at risk of extinction.	Andrianaivoarivelo et al. (2006) | Goodman (2011) | Goodman, Andriafidison et al. (2005) | Monadjem, Razafimanahaka et al. (2017a) | Peterson et al. (1995) | Rasoma & Goodman (2007) | Ratrimomanarivo, Goodman, Taylor et al. (2009) | Richards et al. (2010) | Russ, Bennett et al. (2003) | Volleth et al. (2002)	https://zenodo.org/record/6772244/files/figure.png	4. Peters’s Little Mastff Bat Mormopterus jugularis French: Molosse de Madagascar / German: Peters-Mastino-Fledermaus / Spanish: Morméptero de Madagascar Other common names: Peters’s Goblin Bat , Peters’s Little Free-tailed Bat , Peters’s Wrinkle-lipped Bat Taxonomy. Nyctinomus (Mormopterus) jugularis Peters in Sclater, 1865, “ Antananarivo ,” Madagascar . This species is monotypic. Distribution. Madagascar except most W region. Descriptive notes. Head—body 60-70 mm, tail 20-38 mm, ear 14-17 mm, hindfoot 5-6 mm, forearm 30-39 mm; weight 8: 5—17 g . Males are notably larger than females in some external and cranial measurements. Peters’s Little Mastiff Bat has flattened head, no facial ornamentation, and tail protruding beyond hind margin of uropatagium. Fur is short, soft, and uniformly brown to reddish brown above, becoming pale brown to dirty white on abdomen and throat. Wings and uropatagium are dark brown. Ears are small, subtriangular, and separated at bases, without complex folds or sickle-shaped emargination below tip. Latter feature allows distinction of Peters’s Little Mastiff Bat from the Reunion Little Mastiff Bat (M. francoismoutour) and the Mauritian Little Mastiff Bat ( M. acetabulosus ). Tragus is small but not concealed by antitragus. Adult males have distinct throat gland that is absent in females. Upper lips have several wrinkles, a few spoon-hairs, and many fine hairs. Hindclaws have fine hairs and many stout bristles. It has greater skull lengths (15-4=17-2 mm) than the Reunion Little Mastiff Bat (14-1-15-3 mm). Braincase is low but not particularly flattened, and anterior palatal emargination is wide. M* has third ridge equal or longer than second ridge. Dental formulais11/3,C1/1,P1/2,M 3/3 (x2) = 30. Chromosomal complement has 2n = 48 and FN = 54. Habitat. Wide variety of bioclimatic zones from humid to arid forests, open woodlands, and degraded habitats from sea levelto elevations of ¢. 1750 m . Food and Feeding. In eastern Madagascar , Peters’s Little Mastiff Bats ate Coleoptera (c.61% by volume), Hemiptera (c.20%), Lepidoptera (c.11%), and Diptera (c.3%). Breeding. In late January 2001, a nursery colony with more than 1000 young Peters’s Little Mastiff Bats was observed in Ankarana Special Reserve, northern Madagascar . Activity patterns. Peters’s Little Mastiff Bats are nocturnal. They roost in association with human habitations but also in natural settings such as crevices, roofs of caves, and cliff faces. They have narrowband,relatively long duration echolocation calls with average peak frequency of 32 kHz. Predators include common barn-owls (7yto alba). Movements, Home range and Social organization. Peters’s Little Mastiff Bats are colonial, living in groups of usually less than 100 individuals, but up to 22,000 individuals were reported in Cathedral Cave in Ankarana Special Reserve, Madagascar . Status and Conservation. Classified as Least Concern on The IUCNRed List. Cave colonies of Peters’s Little Mastiff Bat might be threatened by unsustainable hunting practices (e.g. an estimated 70,000 -140,000 insectivorous bats, including Peters’s Little Mastiff Bats, were removed from caves annually in one region of south-western Madagascar ), but its ability to readily colonize human habitations means thatit is not at risk of extinction. Bibliography. Andrianaivoarivelo et al. (2006), Goodman (2011), Goodman, Andriafidison et al. (2005), Monadjem, Razafimanahaka et al. (2017a), Peterson et al. (1995), Rasoma & Goodman (2007), Ratrimomanarivo, Goodman, Taylor et al. (2009), Richards et al. (2010), Russ, Bennett et al. (2003), Volleth et al. (2002).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Molossidae	Mormopterus jugularis	Mormopterus	Mormopterus	jugularis	Peters	1865	1	In Sclater, Proc. Zool. Soc. Lond.	1872:48:00	Peters' Wrinkle-lipped Bat	 albiventer Dobson, 1877.	Madagascar, Tananarive (= Antananarivo).	Madagascar.	Not listed.	Least Concern	 acetabulosus species group. Reviewed by Peterson et al. (1995).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Mormopterus jugularis	23	Peters's Little Mastiff Bat	Peters's Goblin Bat|Peters's Little Free-tailed Bat|Peters's Wrinkle-lipped Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	MOLOSSIDAE	MOLOSSINAE	NA	Mormopterus	NA	jugularis	W. Peters	1865	1	Nyctinomus_(Mormopterus)_jugularis	Peters, W. C .H. (1865.) In Sclater, P. L. Report on a collection of animals from Madagascar, transmitted to the Society by M.J. Caldwell. Proceedings of the Zoological Society London, 1865, 468.	https://www.biodiversitylibrary.org/item/90412#page/548/mode/1up	ZMB 2977		"Antananarivo," Madagascar.			jugularis (W. Peters, 1865)|albiventer (Dobson, 1877)	NA	NA	Madagascar	Africa	Afrotropic	LC	0	0	0	Mormopterus_jugularis	0	sciname match	Mormopterus_jugularis	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	13882	Mormopterus jugularis	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	MOLOSSIDAE	Mormopterus	jugularis	(Peters in Sclater, 1865)		20000000	Mormopterus jugularis	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its widespread distribution across Madagascar and because it is not thought to be facing any major threats across its range.	It is usually found roosting in synanthropic settings but also uses caves and rock fissures (Goodman et al. 2005; Andrianaivoarivelo et al . 2006). A nursery colony with over 1,000 young bats was observed in Réserve Spéciale d’Ankarana (Goodman and Cardiff 2004). In eastern Madagascar it feeds mainly on coleopterans and hemipterans (Andrianaivoarivelo et al . 2006) and forages in open areas (Randrianandriananina et al . 2006). In the south-east it has been netted near water in gallery and spiny forest (Goodman 1999). It frequently shares synanthropic roosts with other molossid species (Andrianaivoarivelo et al . 2006).	This species is not threatened by loss of native forests because it forages in open areas associated with agriculture. It is locally persecuted when roosting in buildings although this is not yet a major threat. Some colonies are subject to hunting and although this is though to have resulted in abandonment in at least two cases (Goodman et al. 2008), harvest levels do not appear a major threat.	There are no quantitative data available on populations of this species; however, it is assumed to have a high population size because it is widespread and roosts in large colonies in buildings.	Unknown	This species is endemic to Madagascar (Peterson et al. 1995) and very widespread across the island with a known distribution extending from the coast up to the high plateau at 1,400 m in Anjozorobe (J. Ranivo pers. comm.).		Terrestrial	As this species is more commonly found roosting in buildings than natural settings it is not found in many of the protected areas in Madagascar (Goodman et al. 2005). However, there is no need for any conservation measures at present as this species appears to be widely distributed and is locally abundant.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Molossidae	Mormopterus	Mormopterus	jugularis	Peters	1865	1	In Sclater, Proc. Zool. Soc. Lond.	1872:48:00	Peters' Wrinkle-lipped Bat	 albiventer Dobson, 1877.	Madagascar, Tananarive (= Antananarivo).	Madagascar.	Not listed.	Least Concern	 acetabulosus species group. Reviewed by Peterson et al. (1995).	Mormopterus jugularis	1005227	23	Peters's Little Mastiff Bat	Peters's Goblin Bat|Peters's Little Free-tailed Bat|Peters's Wrinkle-lipped Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Molossidae	MOLOSSINAE	NA	Mormopterus	NA	jugularis	W. Peters	1865	1	Nyctinomus_(Mormopterus)_jugularis	Peters, W. C .H. (1865.) In Sclater, P. L. Report on a collection of animals from Madagascar, transmitted to the Society by M.J. Caldwell. Proceedings of the Zoological Society London, 1865, 468.	https://www.biodiversitylibrary.org/item/90412#page/548/mode/1up	ZMB 2977		"Antananarivo," Madagascar.			jugularis (W. Peters, 1865)|albiventer (Dobson, 1877)	NA	NA				Madagascar	Africa	Afrotropic	LC	0	0	0	Mormopterus_jugularis	0	sciname match	Mormopterus_jugularis	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Mormopterus_jugularis	1005227	23	Peters's Little Mastiff Bat	Peters's Goblin Bat|Peters's Little Free-tailed Bat|Peters's Wrinkle-lipped Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Molossidae	Molossinae	NA	Mormopterus	NA	jugularis	W. C. H. Peters in P. L. Sclater	1	Nyctinomus (Mormopterus) jugularis	Sclater, P.L. 1865-10. Report on a collection of animals from Madagascar, transmitted to the Society. Proceedings of the Zoological Society of London 1865(2):467-470.	https://www.biodiversitylibrary.org/page/28497439	ZMB 2977	holotype		"Antananarivo," Madagascar.			NA	NA				Madagascar	Africa	Afrotropic	LC	0	0	0	Mormopterus_jugularis	0	sciname match	Mormopterus_jugularis	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Molossidae	Mormopterus	Mormopterus	jugularis	Peters	1865	1	In Sclater, Proc. Zool. Soc. Lond.	1872:48:00	Peters' Wrinkle-lipped Bat	albiventer Dobson, 1877.	Madagascar, Tananarive (= Antananarivo).	Madagascar.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/13882/22083579/' target='_blank'>Least Concern</a>	acetabulosusspecies group. Reviewed by Peterson et al. (1995).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Mormopterus jugularis; Mormopterus jugularis; Mormopterus jugularis; Mormopterus jugularis; Mormopterus jugularis; Mormopterus jugularis; albiventer; albiventer; jugularis; albiventer; Molosse de Madagascar; Peters-Mastino-Fledermaus; Morméptero de Madagascar; Peters’s Goblin Bat; Peters’s Little Free-tailed Bat; Peters’s Wrinkle-lipped Bat; Peters's Little Mastiff Bat; Peters's Goblin Bat; Peters's Little Free-tailed Bat; Peters's Wrinkle-lipped Bat; Peters's Wrinkle-lipped Bat; Peters' Wrinkle-lipped Bat; M. jugularis