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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L72	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus	Artibeus fraterculus		[MSW2] Subgenus Artibeus. Considered a subspecies of jamaicensis by Jones and Carter (1976:2); but see Koopman (19781>:14).; [MSW3] Subgenus Artibeus. Considered a subspecies of jamaicensis by Jones and Carter (1976), but see Koopman (1978b) and Marques-Aguiar (1994).; [HMW] Artibeus fraterculus Anthony, 1924 , “Portovelo, Provincia del Oro, Ecuador ;altitude, 2000 ft. [= 610 m ].” Artibeus fraterculus is in subgenus Artibeus . Molecular analyses suggest thatit is sister to A. hirsutus from central Mexico . A few authors listed A. fraterculus as a subspecies of A. jamaicensis . Monotypic.; [batnames2022] Subgenus Artibeus .  Considered a subspecies of jamaicensis by Jones and Carter (1976), but see Koopman (1978 b ) and Marques-Aguiar (1994).; [IUCN] This used to be a subspecies of A. jamaicensis .; [batnames2023] Subgenus Artibeus .  Considered a subspecies of jamaicensis by Jones and Carter (1976), but see Koopman (1978 b ) and Marques-Aguiar (1994).; [batnames2025_1.7] Subgenus Artibeus. Considered a subspecies of jamaicensis by Jones and Carter (1976), but see Koopman (1978 b) and Marques-Aguiar (1994).														fraterculus	This used to be a subspecies of A. jamaicensis .			fraterculus	fraterculus			fraterculus H. E. Anthony, 1924		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.		S Ecuador, N Peru; refs 4.6,71	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Artibeus fraterculus	Ecuador, El Oro, Portovelo, 2000 ft. (610 m).	Anthony	1924	Am. Mus. Novit., 114:5.	Distribution: Confined to arid areas in western Ecuador and western Peru.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5		S Ecuador, N Peru	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Anthony	1924	Am. Mus. Novit., 114:5.	Subgenus Artibeus. Considered a subspecies of jamaicensis by Jones and Carter (1976:2); but see Koopman (19781>:14).	Ecuador, Peru.	Ecuador, El Oro, Portovelo, 2,000 ft.(610 m).		ANTHONY	1924	Size fairly large (forearm length, 51 -61 mm; greatest length of skull, 26-29 mm). Preorbital and postorbital processes poorly developed. Last upper molar absent, last lower normally present. Medial facial stripe fairly well developed but lateral absent or poorly developed. Uropatagium virtually hairless.	Distribution: Confined to arid areas in western Ecuador and western Peru.	No subspecies.		91	species	A. fraterculus	ANTHONY	1924	Artibeus	subgenus	Artibeus fraterculus				Size fairly large (forearm length, 51 -61 mm; greatest length of skull, 26-29 mm). Preorbital and postorbital processes poorly developed. Last upper molar absent, last lower normally present. Medial facial stripe fairly well developed but lateral absent or poorly developed. Uropatagium virtually hairless.	No subspecies.		9. A.fraterculus ANTHONY 1924.	9	NA			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Phyllostomidae	Stenodermatinae	Stenodermatini	Artibeus fraterculus	Artibeus	Artibeus	fraterculus	Anthony		1924		Am. Mus. Novit.	114		5		Fraternal Fruit-eating Bat	Ecuador, El Oro, Portovelo, 2,000 ft. (610 m).	Ecuador, Peru.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Vulnerable.		Subgenus Artibeus. Considered a subspecies of jamaicensis by Jones and Carter (1976), but see Koopman (1978b) and Marques-Aguiar (1994).	03A687BCFFE6FFE916B8FAC1FBFEF11F	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Phyllostomidae_444.pdf.imf	hash://md5/ff9fffc4ffb1ffb1133cffbaffe0f244	571	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/03/A6/87/03A687BCFFE6FFE916B8FAC1FBFEF11F.xml	Artibeus fraterculus	Phyllostomidae	Artibeus	fraterculus	Anthony	1924	Artibée d Anthony @fr | Peru-Fruchtvampir @de | Artibeo de Anthony @es | Fraternal Artibeus @en	Artibeus fraterculus Anthony, 1924 , “Portovelo, Provincia del Oro, Ecuador ;altitude, 2000 ft. [= 610 m ].” Artibeus fraterculus is in subgenus Artibeus . Molecular analyses suggest thatit is sister to A. hirsutus from central Mexico . A few authors listed A. fraterculus as a subspecies of A. jamaicensis . Monotypic.	Endemic to Pacific coast from W Ecuador ( Manabi Province ) to W Peru ( Lima Department).	Head—body 64-76 (tailless), ear 15-21 mm, hindfoot 12-16 mm, forearm 52-59 mm; weight 30-55 g. The Fraternal Fruit-eating Bat is the smallest and palest species in subgenus Artibeus in South America. Pelageis soft and velvety. Dorsal fur is dark gray to grayish brown and has dichromatic appearance, occurring in grayish brown and yellowish brown morphs. Dorsalfuris short (less or equal to 8 mm ). Ventral pelage is pale, with tips of hairs appearing silvery gray, giving a frosted appearance. Basally, dorsal and ventral surfaces of forearm are hairy. Horseshoe of noseleat is free. Wing membraneis blackish gray and extends to near metatarsals. Uropatagium is dark brown and broad (greater or equal to 10 mm ), with central notch and slightly hairy and fringed medially. Dental formulais12/2,C1/1,P 2/2, M 2/3 ( x2 ) = 30. Crowns of I' are bilobed. Shape of M' is almost triangular in occlusal view, and hypocone is well developed—a character unique to the Fraternal Fruit-eating Bat. Chromosomal complement has 2n = 30 and FN = 56, with XX/XY sex-determination system.	Coastal mangroves, desert scrubland, deciduous and semideciduous forests, disturbed habitats, croplands, and urban areas from sea level to elevations of ¢. 2150 m . The Fraternal Fruit-eating Bat is more common in drier areas (tropical dry forests) and less abundant in wet areas (mangroves).	The Fraternal Fruit-eating Bat is a generalist frugivore; its diet is varied and includes fruits of Mangifera indica ( Anacardiaceae ); Brosimum alicastrum and Ficus spp. ( Moraceae ); Muntingia calabura ( Muntingiaceae ); Psidium guajava and Syzygiumjambos ( Myrtaceae ); Eriobotryajaponica ( Rosaceae ); Acnistus arborescens and Solanum crinitipes ( Solanaceae ); Styrax subargentea ( Styracaceae ); and Cecropia spp. ( Urticaceae ). Insect remains have been found in feces.	The Fraternal Fruit-eating Bat exhibits seasonal bimodal polyestry, with parturition occurring during dry and wet seasons, with peaks in February and May. In Ecuador , pregnant females were found in October-November and lactating females in July and November.	The Fraternal Fruit-eating Bat is nocturnal and appears to be active throughout the night without distinctive peaks. It inhabits natural roosts such as hollow trees, shrubs, caves, termite mounds, and artificial structureslike bridges, churches, houses, gardens, and mines. Parameters of search-phase FM echolocation call taken in Loja , Ecuador , included starting frequency of 95-8 kHz (range 89-4-104 kHz), ending frequency of 62-3 kHz (59-2-68 kHz), frequency of maximum energy 74-4 kHz (67-9-84-1 kHz), and duration 1-6 milliseconds; it had three harmonics. On following pages: 190. Hairy Fruit-eating Bat ( Artibeus hirsutus ); 191. Fringed Fruit-eating Bat ( Artibeus fimbriatus ); 192. Ecuadorian Fruit-eating Bat ( Artibeus aequatorialis ); 193. Jamaican Fruit-eating Bat ( Artibeus jamaicensis ); 194. Dark Fruit-eating Bat ( Artibeus obscurus ); 195. Schwartz's Fruit-eating Bat ( Artibeus schwartz); 196. Great Fruit-eating Bat ( Artibeus lituratus ); 197. Large Fruit-eating Bat ( Artibeus amplus ); 198. Flat-faced Fruit-eating Bat ( Artibeus planirostris ); 199. Rosenberg’s Fruit-eating Bat ( Artibeus rosenbergi ); 200. Thomas's Fruit-eating Bat ( Artibeus watson ); 201. Toltec Fruit-eating Bat ( Artibeus toltecus ); 202. Pygmy Fruit-eating Bat ( Artibeus phaeotis ); 203. Gervais's Fruit-eating Bat ( Artibeus cinereus ); 204. Andersen's Fruit-eating Bat ( Artibeus anderseni ); 205.Little Fruit-eating Bat (Artibeusravus); 206. Aztec Fruit-eating Bat ( Artibeus aztecus ); 207. Bogota Fruit-eating Bat ( Artibeus bogotensis ); 208. Silvery Fruit-eating Bat ( Artibeus glaucus ); 209. Dwarf Fruit-eating Bat ( Artibeus gnomus ); 210. Jamaican Fig-eating Bat ( Ariteus flavescens ); 211. Tree Bat ( Ardops nicholls); 212. Red Fruit Bat ( Stenoderma rufum ); 213. Wrinkle-faced Bat ( Centurio senex ): 214. Ipanema Broad-nosed Bat ( Pygoderma bilabiatum ); 215. Visored Bat ( Sphaeronycteris toxophyllum ); 216. Little White-shouldered Bat ( Ametrida centurio ); 217. Cuban Fig-eating Bat ( Phyllops falcatus ).	Fraternal Fruit-eating Bats have been found in groups ranging from nine individuals in a termite nest to 709 individuals in an active coalmine.	Classified as Least Concern on The IUCN Red List. The Fraternal Fruit-eating Bat has a wide distribution and presumably large population, and itis unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	Albuja (1999) | Anthony (1924a) | Baker (1979) | Carrera et al. (2010) | Koopman (1978) | Larsen, PA. et al. (2007) | Marques-Aguiar (2008a) | Mena & Williams de Castro (2002) | Novoa et al. (2011) | Patterson et al. (1992) | Pinto et al. (2013) | Redondo et al. (2008) | Salas et al. (2018) | Tirira (2012c)	https://zenodo.org/record/6458975/files/figure.png	189. Fraternal Fruit-eating Bat Artibeus fraterculus French: Artibée dAnthony / German: Peru-Fruchtvampir / Spanish: Artibeo de Anthony Other common names: Fraternal Artibeus Taxonomy. Artibeus fraterculus Anthony, 1924 , “Portovelo, Provincia del Oro, Ecuador ;altitude, 2000 ft. [= 610 m ].” Artibeus fraterculus is in subgenus Artibeus . Molecular analyses suggest thatit is sister to A. hirsutus from central Mexico . A few authors listed A. fraterculus as a subspecies of A. jamaicensis . Monotypic. Distribution. Endemic to Pacific coast from W Ecuador ( Manabi Province ) to W Peru ( Lima Department). Descriptive notes. Head—body 64-76 (tailless), ear 15-21 mm, hindfoot 12-16 mm, forearm 52-59 mm; weight 30-55 g. The Fraternal Fruit-eating Bat is the smallest and palest species in subgenus Artibeus in South America. Pelageis soft and velvety. Dorsal fur is dark gray to grayish brown and has dichromatic appearance, occurring in grayish brown and yellowish brown morphs. Dorsalfuris short (less or equal to 8 mm ). Ventral pelage is pale, with tips of hairs appearing silvery gray, giving a frosted appearance. Basally, dorsal and ventral surfaces of forearm are hairy. Horseshoe of noseleat is free. Wing membraneis blackish gray and extends to near metatarsals. Uropatagium is dark brown and broad (greater or equal to 10 mm ), with central notch and slightly hairy and fringed medially. Dental formulais12/2,C1/1,P 2/2, M 2/3 ( x2 ) = 30. Crowns of I' are bilobed. Shape of M' is almost triangular in occlusal view, and hypocone is well developed—a character unique to the Fraternal Fruit-eating Bat. Chromosomal complement has 2n = 30 and FN = 56, with XX/XY sex-determination system. Habitat. Coastal mangroves, desert scrubland, deciduous and semideciduous forests, disturbed habitats, croplands, and urban areas from sea level to elevations of ¢. 2150 m . The Fraternal Fruit-eating Bat is more common in drier areas (tropical dry forests) and less abundant in wet areas (mangroves). Food and Feeding. The Fraternal Fruit-eating Bat is a generalist frugivore; its diet is varied and includes fruits of Mangifera indica ( Anacardiaceae ); Brosimum alicastrum and Ficus spp. ( Moraceae ); Muntingia calabura ( Muntingiaceae ); Psidium guajava and Syzygiumjambos ( Myrtaceae ); Eriobotryajaponica ( Rosaceae ); Acnistus arborescens and Solanum crinitipes ( Solanaceae ); Styrax subargentea ( Styracaceae ); and Cecropia spp. ( Urticaceae ). Insect remains have been found in feces. Breeding. The Fraternal Fruit-eating Bat exhibits seasonal bimodal polyestry, with parturition occurring during dry and wet seasons, with peaks in February and May. In Ecuador , pregnant females were found in October-November and lactating females in July and November. Activity patterns. The Fraternal Fruit-eating Bat is nocturnal and appears to be active throughout the night without distinctive peaks. It inhabits natural roosts such as hollow trees, shrubs, caves, termite mounds, and artificial structureslike bridges, churches, houses, gardens, and mines. Parameters of search-phase FM echolocation call taken in Loja , Ecuador , included starting frequency of 95-8 kHz (range 89-4-104 kHz), ending frequency of 62-3 kHz (59-2-68 kHz), frequency of maximum energy 74-4 kHz (67-9-84-1 kHz), and duration 1-6 milliseconds; it had three harmonics. On following pages: 190. Hairy Fruit-eating Bat ( Artibeus hirsutus ); 191. Fringed Fruit-eating Bat ( Artibeus fimbriatus ); 192. Ecuadorian Fruit-eating Bat ( Artibeus aequatorialis ); 193. Jamaican Fruit-eating Bat ( Artibeus jamaicensis ); 194. Dark Fruit-eating Bat ( Artibeus obscurus ); 195. Schwartz's Fruit-eating Bat ( Artibeus schwartz); 196. Great Fruit-eating Bat ( Artibeus lituratus ); 197. Large Fruit-eating Bat ( Artibeus amplus ); 198. Flat-faced Fruit-eating Bat ( Artibeus planirostris ); 199. Rosenberg’s Fruit-eating Bat ( Artibeus rosenbergi ); 200. Thomas's Fruit-eating Bat ( Artibeus watson ); 201. Toltec Fruit-eating Bat ( Artibeus toltecus ); 202. Pygmy Fruit-eating Bat ( Artibeus phaeotis ); 203. Gervais's Fruit-eating Bat ( Artibeus cinereus ); 204. Andersen's Fruit-eating Bat ( Artibeus anderseni ); 205.Little Fruit-eating Bat (Artibeusravus); 206. Aztec Fruit-eating Bat ( Artibeus aztecus ); 207. Bogota Fruit-eating Bat ( Artibeus bogotensis ); 208. Silvery Fruit-eating Bat ( Artibeus glaucus ); 209. Dwarf Fruit-eating Bat ( Artibeus gnomus ); 210. Jamaican Fig-eating Bat ( Ariteus flavescens ); 211. Tree Bat ( Ardops nicholls); 212. Red Fruit Bat ( Stenoderma rufum ); 213. Wrinkle-faced Bat ( Centurio senex ): 214. Ipanema Broad-nosed Bat ( Pygoderma bilabiatum ); 215. Visored Bat ( Sphaeronycteris toxophyllum ); 216. Little White-shouldered Bat ( Ametrida centurio ); 217. Cuban Fig-eating Bat ( Phyllops falcatus ). Movements, Home range and Social organization. Fraternal Fruit-eating Bats have been found in groups ranging from nine individuals in a termite nest to 709 individuals in an active coalmine. Status and Conservation. Classified as Least Concern on The IUCN Red List. The Fraternal Fruit-eating Bat has a wide distribution and presumably large population, and itis unlikely to be declining at nearly the rate required to qualify for listing in a threatened category. Bibliography. Albuja (1999), Anthony (1924a), Baker (1979), Carrera et al. (2010), Koopman (1978), Larsen, PA. et al. (2007), Marques-Aguiar (2008a), Mena & Williams de Castro (2002), Novoa et al. (2011), Patterson et al. (1992), Pinto et al. (2013), Redondo et al. (2008), Salas et al. (2018), Tirira (2012c).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Phyllostomidae	Artibeus fraterculus	Artibeus		fraterculus	Anthony	1924	0	Am. Mus. Novitates	######	Fraternal Fruit-eating Bat	None.	Ecuador, El Oro, Portovelo, 2,000 ft. (610 m).	Ecuador, Peru.	Not listed.	Least Concern	Subgenus Artibeus .  Considered a subspecies of jamaicensis by Jones and Carter (1976), but see Koopman (1978 b ) and Marques-Aguiar (1994).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Artibeus fraterculus	23	Fraternal Fruit-eating Bat	Fraternal Artibeus	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	PHYLLOSTOMIDAE	STENODERMATINAE	STENODERMATINI	Artibeus	NA	fraterculus	Anthony	1924	0	Artibeus_fraterculus	Anthony, H. E. (1924). Preliminary report on Ecuadorean mammals. No. 4. Americam Museum Novitates, 114, 5.	http://digitallibrary.amnh.org/bitstream/handle/2246/4332//v2/dspace/ingest/pdfSource/nov/N0114.pdf?sequence=1&isAllowed=y	AMNH 47248		"Portovelo, Provincia del Oro, Ecuador; altitude, 2000 ft. [= 610 m]."			fraterculus Anthony, 1924	NA	NA	Ecuador|Peru	South America	Neotropic	LC	0	0	0	Artibeus_fraterculus	0	sciname match	Artibeus_fraterculus	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	2127	Artibeus fraterculus	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	PHYLLOSTOMIDAE	Artibeus	fraterculus	Anthony, 1924	This used to be a subspecies of A. jamaicensis .	20000000	Artibeus fraterculus	Least Concern		2015	2015-07-20 00:00:00 UTC	3.1	English	This species is listed as Least Concern in view of its wide distribution, presumed large population, and because it is unlikely to be declining at nearly the rate required to qualify for listing in a threatened category.	This species is strongly frugivorous. This species is common in dry deserts and arid zones (Burneo pers. comm.).	There are no threats to this species.	In Ecuador this bat is locally common (Tirira in prep.), as in Peru (Solari pers. comm.). Sometimes it is considered to be a pest to crops.	Decreasing	This species occurs in west Ecuador, and northwest and central Peru (Simmons 2005).	This species is not used.	Terrestrial	There are no conservation actions in place or recommended for this species.	Neotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Phyllostomidae	Artibeus		fraterculus	Anthony	1924	0	Am. Mus. Novitates	114:05:00	Fraternal Fruit-eating Bat	None.	Ecuador, El Oro, Portovelo, 2,000 ft. (610 m).	Ecuador, Peru.	Not listed.	Least Concern	Subgenus Artibeus .  Considered a subspecies of jamaicensis by Jones and Carter (1976), but see Koopman (1978 b ) and Marques-Aguiar (1994).	Artibeus fraterculus	1005006	23	Fraternal Fruit-eating Bat	Fraternal Artibeus	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	NOCTILIONOIDEA	Phyllostomidae	STENODERMATINAE	STENODERMATINI	Artibeus	NA	fraterculus	Anthony	1924	0	Artibeus_fraterculus	Anthony, H. E. (1924). Preliminary report on Ecuadorean mammals. No. 4. Americam Museum Novitates, 114, 5.	http://digitallibrary.amnh.org/bitstream/handle/2246/4332//v2/dspace/ingest/pdfSource/nov/N0114.pdf?sequence=1&isAllowed=y	AMNH 47248		"Portovelo, Provincia del Oro, Ecuador; altitude, 2000 ft. [= 610 m]."			fraterculus Anthony, 1924	NA	NA				Ecuador|Peru	South America	Neotropic	LC	0	0	0	Artibeus_fraterculus	0	sciname match	Artibeus_fraterculus	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Artibeus_fraterculus	1005006	23	Fraternal Fruit-eating Bat	Fraternal Artibeus	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Noctilionoidea	Phyllostomidae	Stenodermatinae	Stenodermatini	Artibeus	NA	fraterculus	H. E. Anthony	0	Artibeus fraterculus	Anthony, H.E. 1924-05-29. Preliminary report on Ecuadorean mammals. No. 4. American Museum Novitates 114:1-6.	https://www.biodiversitylibrary.org/item/170246	AMNH M-47248	holotype	http://portal.vertnet.org/o/amnh/mammals?id=urn-catalog-amnh-mammals-m-47248	"Portovelo, Provincia del Oro, Ecuador; altitude, 2000 ft. [= 610 m]."			NA	NA				Ecuador|Peru	South America	Neotropic	LC	0	0	0	Artibeus_fraterculus	0	sciname match	Artibeus_fraterculus	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Phyllostomidae	Artibeus		fraterculus	Anthony	1924	0	Am. Mus. Novitates	114:05:00	Fraternal Fruit-eating Bat	None.	Ecuador, El Oro, Portovelo, 2,000 ft. (610 m).	Ecuador, Peru.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/2127/21998872/' target='_blank'>Least Concern</a>	Subgenus Artibeus. Considered a subspecies of jamaicensis by Jones and Carter (1976), but see Koopman (1978 b) and Marques-Aguiar (1994).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	Artibeina	Artibeus fraterculus; Artibeus fratercula; fraterculus; Artibée d Anthony; Peru-Fruchtvampir; Artibeo de Anthony; Fraternal Artibeus; Fraternal Fruit-eating Bat; Fraternal Artibeus; Fraternal Fruit-eating Bat; Fraternal Fruit-eating Bat; Artibeus fraterculus fraterculus; A. fraterculus