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(1=author & date in parentheses)	Citation	Pages	Common Name	Synonyms	Type Locality	Distribution	CITES	IUCN	Comments	column3781	column3791	subtribe	CONCAT_ALTNAMES
line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L706	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	Tadarida bemmeleni	Chaerephon bemmeleni	Tadarida bemmelini	Chaerephon bemmeleni	Chaerephon bemmelini	Chaerephon bemmeleni	Chaerephon bemmeleni	Mops bemmeleni	Mops bemmeleni	Chaerephon bemmeleni	Mops bemmeleni	Mops bemmeleni	Mops bemmeleni	Mops bemmeleni	Mops bemmeleni		[MSW2] Includes cistura; see Koopman (1975:425).; [MSW3] bivittatus species group. Includes cistura; see Koopman (1975) and Peterson (1971). Revised by Peterson (1971).; [HMW] Nyctinomus bemmeleni Jentink, 1879 , “ Liberia .” Two subspecies are recognized.; [batnames2022]  bivittatus species group. Includes cistura; see Koopman (1975) and Peterson (1971). Revised by Peterson (1971).; [MDD2022] moved from Chaerephon to Mops, although further research is needed to confirm this placement; [IUCN] This may be a complex of two distinct species, one in the east of the range and one in the west. Further taxonomic studies are needed to resolve this.; [batnames2023]  bivittatus species group. Includes cistura; see Koopman (1975) and Peterson (1971). Revised by Peterson (1971).; [MDD2023] moved from Chaerephon to Mops, although further research is needed to confirm this placement; [MDD2025_2.0] moved from Chaerephon to Mops, although further research is needed to confirm this placement; [batnames2025_1.7] bivittatusspecies group. Includes cistura; see Koopman (1975) and Peterson (1971). Revised by Peterson (1971).; [MDD2025_2.2] moved from Chaerephon to Mops, although further research is needed to confirm this placement						cistura.	bemmelini, cisturus	bemmeleni , cistura		bemmeleni, cistura		bemmeleni, cistura		bemmeleni, cisturus	This may be a complex of two distinct species, one in the east of the range and one in the west. Further taxonomic studies are needed to resolve this.	bemmeleni, cistura		bemmeleni, cisturus	bemmeleni, cisturus	bemmeleni, cistura		bemmeleni (Jentink, 1879)|cisturus (O. Thomas, 1903)|bemmelini (Freeman, 1981) [incorrect subsequent spelling]		Corbet, G.B. and Hill, J.E. 1980. A World List of Mammalian Species. British Museum (Natural History), London, 226 pp.	Gland-tailed free-tailed bat	Liberia – S Sudan – N Tanzania; ref. 4.115	Honacki, J.H., Kinman, K.E. and Koeppl, J.W. 1982. Mammal Species of the World: A Taxonomic and Geographic Reference. Allen Press, Lawrence, 694 pp.	Chaerephon bemmeleni	Liberia.	Jentink	1879	Notes Leyden Mus., 1:125.	Distribution: Ranging from Liberia to southern Sudan and south to northeastern Zaire and northern Tanzania.		Corbet, G.B. and Hill, J.E. 1991. A World List of Mammalian Species. Third edition. Oxford University Press, London, 243 pp. ISBN 0-19-854017-5	Gland-tailed free-tailed bat	Liberia – S Sudan – N Tanzania	Koopman, K.F. 1993. Order Chiroptera. Pp. 137–242 in Wilson, D.E. and Reeder, D.M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference. Second edition. Smithsonian Institution Press, Washington, 1206 pp.	Jentink	1879	Notes Leyden Mus., 1:125.	Includes cistura; see Koopman (1975:425).	Liberia, Cameroon, Sudan, Zaire, Uganda, Kenya, Tanzania.	"Liberia", but see Kuhn (1965).		JENTINCK	1879	Anterior palatal emargination narrow but definitely present. Basisphenoid pits relatively shallow. Anterior upper premolar relatively large. Size relatively small (forearm length, 40-47 mm). Upper lip wrinkles relatively few.	Distribution: Ranging from Liberia to southern Sudan and south to northeastern Zaire and northern Tanzania.	Two subspecies are recognized:	C. b. bemmelini (known from Liberia and Cameroon), C. b. cisturus (eastern African range).	139	species	C. bemmelini	JENTINCK	1879	Chaerephon	genus	Chaerephon bemmelini				Anterior palatal emargination narrow but definitely present. Basisphenoid pits relatively shallow. Anterior upper premolar relatively large. Size relatively small (forearm length, 40-47 mm). Upper lip wrinkles relatively few.	Two subspecies are recognized:		1. C. bemmelini (JENTINCK 1879) [bivittata group],	1	_M. b. bemmeleni_ (Jentink, 1879); _M. b. cisturus_ (Thomas, 1903)			Don E. Wilson & DeeAnn M. Reeder (editors). 2005. Mammal Species of the World. A Taxonomic and Geographic Reference (3rd ed), Johns Hopkins University Press, 2,142 pp. (Available from Johns Hopkins University Press, 1-800-537-5487 or (410) 516-6900, or at http://www.press.jhu.edu).	CHIROPTERA	Molossidae	Molossinae		Chaerephon bemmeleni	Chaerephon		bemmeleni	Jentink	y	1879		Notes Leyden Mus.	1		125		Gland-tailed Free-tailed Bat	Liberia.	Sierra Leone, Liberia, Cameroon, Sudan, Dem. Rep. Congo, Uganda, Kenya, Tanzania.	IUCN 2003 and IUCN/SSC Action Plan (2001) – Lower Risk (lc).	cistura Thomas, 1903.	bivittatus species group. Includes cistura; see Koopman (1975) and Peterson (1971). Revised by Peterson (1971).	194287C9FFA6BA0AB16FF2ACBB80F3BA	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Molossidae_598.pdf.imf	hash://md5/e57bffb1ffbcba10b412f760b226ffce	645	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/19/42/87/194287C9FFA6BA0AB16FF2ACBB80F3BA.xml	Chaerephon bemmeleni	Molossidae	Chaerephon	bemmeleni		1879	Tadaride de van Bemmelen @fr | Van-Bemmelen-Bulldogfledermaus @de | Caerepon de van Bemmelen @es | Gland-tailed Wrinkle-lipped Bat @en | Van Bemmelen's Wrinkle-lipped Bat @en	Nyctinomus bemmeleni Jentink, 1879 , “ Liberia .” Two subspecies are recognized.	C.b.bemmeleniJentink,1879—Guinea,SierraLeone,Liberia,IvoryCoast,andCameroon. C. b. cistura Thomas, 1903 — SC South Sudan , Uganda , E DR Congo , SW Kenya , and N Tanzania .	Head-body ¢.60-70 mm, tail 34—40 mm , ear 14-20 mm, hindfoot 9-13 mm, forearm 41-48 mm; weight 11-14 g. The Gland-tailed Free-tailed Bat is distinguished from other molossids by having a pair of sac-like glands in both sexes just lateral to base of tail that open ventrally and may be evident as dorsal swellings. Pelage is short, covering rump but not extending onto tail membrane. Pelage is dark brown with slight frosting but no spots or stripes above, paler below, with flanks darker than belly; there are no mid-ventral markings, and ventral flank-stripes are whitish, contrasting with dark flanks. Upper lip has 6-8 well-defined wrinkles on each side and comparatively few spoon-hairs. Ears are blackish brown, joined by V-shaped band of skin between inner margins, very broad and square, just reaching snout when laid forward. Interaural band has no distinct crest, sometimes having dense hairs, sometimes being almost naked. Tragus is small and concealed by large antitragus. Wing and tail membranes are pale gray. Anterior emargination of palate is narrow and basisphenoid pits are shallow. M? has third ridge about three-quarters as long as second ridge. Chromosomal complement has 2n = 48 and FNa = 54.	Both moist and dry lowland tropical forests, montane grasslands (at 1600 m on Mount Nimba ), as well as drier and moist savannas in the eastern population, including Acacia ( Fabaceae )—-Commiphora ( Burseraceae ) bushland, Isoberlinia ( Fabaceae ) woodland, and forest-savanna mosaic.	No information.	No information.	Gland-tailed Free-tailed Bats are nocturnal and may roost in hollow trees or crevices in rocky areas.	Gland-tailed Free-tailed Bats probably roost communally in small groups, which they may share with Ansorge’s Freetailed Bats ( Chaerephon ansorgei ).	Classified as Least Concern on The IUCN Red Lust.	Fahr (2013v) | Monadjem, Fahr, Bergmans, Mickleburgh, Hutson & Cotterill (2017a)	https://zenodo.org/record/6418571/files/figure.png	60. Gland-tailed Free-tailed Bat Chaerephon bemmeleni French: Tadaride de van Bemmelen / German: Van-Bemmelen-Bulldogfledermaus / Spanish: Caerepon de van Bemmelen Other common names: Gland-tailed Wrinkle-lipped Bat , Van Bemmelen's Wrinkle-lipped Bat Taxonomy. Nyctinomus bemmeleni Jentink, 1879 , “ Liberia .” Two subspecies are recognized. Subspecies and Distribution. C.b.bemmeleniJentink,1879—Guinea,SierraLeone,Liberia,IvoryCoast,andCameroon. C. b. cistura Thomas, 1903 — SC South Sudan , Uganda , E DR Congo , SW Kenya , and N Tanzania . Descriptive notes. Head-body ¢.60-70 mm, tail 34—40 mm , ear 14-20 mm, hindfoot 9-13 mm, forearm 41-48 mm; weight 11-14 g. The Gland-tailed Free-tailed Bat is distinguished from other molossids by having a pair of sac-like glands in both sexes just lateral to base of tail that open ventrally and may be evident as dorsal swellings. Pelage is short, covering rump but not extending onto tail membrane. Pelage is dark brown with slight frosting but no spots or stripes above, paler below, with flanks darker than belly; there are no mid-ventral markings, and ventral flank-stripes are whitish, contrasting with dark flanks. Upper lip has 6-8 well-defined wrinkles on each side and comparatively few spoon-hairs. Ears are blackish brown, joined by V-shaped band of skin between inner margins, very broad and square, just reaching snout when laid forward. Interaural band has no distinct crest, sometimes having dense hairs, sometimes being almost naked. Tragus is small and concealed by large antitragus. Wing and tail membranes are pale gray. Anterior emargination of palate is narrow and basisphenoid pits are shallow. M? has third ridge about three-quarters as long as second ridge. Chromosomal complement has 2n = 48 and FNa = 54. Habitat. Both moist and dry lowland tropical forests, montane grasslands (at 1600 m on Mount Nimba ), as well as drier and moist savannas in the eastern population, including Acacia ( Fabaceae )—-Commiphora ( Burseraceae ) bushland, Isoberlinia ( Fabaceae ) woodland, and forest-savanna mosaic. Food and Feeding. No information. Breeding. No information. Activity patterns. Gland-tailed Free-tailed Bats are nocturnal and may roost in hollow trees or crevices in rocky areas. Movements, Home range and Social organization. Gland-tailed Free-tailed Bats probably roost communally in small groups, which they may share with Ansorge’s Freetailed Bats ( Chaerephon ansorgei ). Status and Conservation. Classified as Least Concern on The IUCN Red Lust. Bibliography. Fahr (2013v), Monadjem, Fahr, Bergmans, Mickleburgh, Hutson & Cotterill (2017a).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Molossidae	Mops bemmeleni	Mops		bemmeleni	Jentink	1879	1	Notes Leyden Mus.	0.1285	Gland-tailed Free-tailed Bat	<b> cistura </b>Thomas, 1903.	Liberia.	Sierra Leone, Liberia, Cameroon, Sudan, Dem. Rep. Congo, Uganda, Kenya, Tanzania.	Not listed.	Least Concern as Chaerephon bemmeleni 	 bivittatus species group. Includes cistura; see Koopman (1975) and Peterson (1971). Revised by Peterson (1971).	Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Mops bemmeleni	23	Gland-tailed Free-tailed Bat	Gland-tailed Wrinkle-lipped Bat|Van Bemmelen's Wrinkle-lipped Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	MOLOSSIDAE	MOLOSSINAE	NA	Mops	Chaerephon	bemmeleni	Jentink	1879	1	Nyctinomus_Bemmeleni	Jentink, F. A. (1879). On a new bat, Nyctinomus bemmeleni, from Liberia. Notes from the Royal Zoological Museum of the Netherlands at Leyden, 1, 125.	https://www.biodiversitylibrary.org/item/37312#page/137/mode/1up	RMNH MAM.21649		"Liberia."			bemmeleni (Jentink, 1879)|cisturus (O. Thomas, 1903)	moved from Chaerephon to Mops, although further research is needed to confirm this placement	Gregorin, R., & Cirranello, A. (2016). Phylogeny of Molossidae Gervais (Mammalia: Chiroptera) inferred by morphological data. Cladistics, 32(1), 2-35.|Simmons, N. B. and A. L. Cirranello (2020). https://batnames.org/explore.html. Accessed: 10 May 2020. URL: https://batnames.org/explore.html.	Guinea|Sierra Leone|Liberia|Burkina Faso?|Côte d'Ivoire|Cameroon|Central African Republic?|South Sudan|Uganda|Democratic Republic of the Congo|Kenya|Tanzania|Rwanda?	Africa	Afrotropic	LC	0	0	0	Chaerephon_bemmeleni	1	oldname match	Chaerephon_bemmeleni	0	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	4307	Chaerephon bemmeleni	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	MOLOSSIDAE	Chaerephon	bemmeleni	(Jentink, 1879)	This may be a complex of two distinct species, one in the east of the range and one in the west. Further taxonomic studies are needed to resolve this.	20000000	Chaerephon bemmeleni	Least Concern		2017	2016-08-31 00:00:00 UTC	3.1	English	Listed as Least Concern in view of its wide distribution, presumed large population, it occurs in a number of protected areas, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.	In the west of its range it is associated with montane tropical moist forest. The eastern populations are generally found in savanna and grassland savanna, Acacia , Combretum and Acacia–Commiphora communities. Roost in rocks, rock caves and fissures (Freeman 1981).	The species is threatened by habitat loss, largely through logging activities and the conversion of land to agricultural use.	The abundance of this species is unclear, in parts of the range it seems to be a rare species that is found in small colonies consisting of a few individuals. In East African this species exhibits a clumped distribution, yet is relatively abundant locally (Weballa et al. 2004).	Unknown	This African species is widely distributed from Sierra Leone in the west, then patchily through West and Central Africa to southern Sudan, Uganda, Kenya and Tanzania in the east. It has been recorded up to 1,700 m Asl on Mount Nimba along the border of Côte d'Ivoire and Guinea (J. Fahr pers. comm. 2004).		Terrestrial	It is present in Meru National Park, Kenya, and is presumably present in several more protected areas. Further studies are needed into the taxonomy, natural history, threats and distribution of this species.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Molossidae	Mops		bemmeleni	Jentink	1879	1	Notes Leyden Mus.	0.128472	Gland-tailed Free-tailed Bat	<b> cistura </b>Thomas, 1903.	Liberia.	Sierra Leone, Liberia, Cameroon, Sudan, Dem. Rep. Congo, Uganda, Kenya, Tanzania.	Not listed.	Least Concern as Chaerephon bemmeleni 	 bivittatus species group. Includes cistura; see Koopman (1975) and Peterson (1971). Revised by Peterson (1971).	Mops bemmeleni	1005147	23	Gland-tailed Free-tailed Bat	Gland-tailed Wrinkle-lipped Bat|Van Bemmelen's Wrinkle-lipped Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Molossidae	MOLOSSINAE	NA	Mops	Chaerephon	bemmeleni	Jentink	1879	1	Nyctinomus_Bemmeleni	Jentink, F. A. (1879). On a new bat, Nyctinomus bemmeleni, from Liberia. Notes from the Royal Zoological Museum of the Netherlands at Leyden, 1, 125.	https://www.biodiversitylibrary.org/item/37312#page/137/mode/1up	RMNH MAM.21649		"Liberia."			bemmeleni (Jentink, 1879)|cisturus (O. Thomas, 1903)	moved from Chaerephon to Mops, although further research is needed to confirm this placement	Gregorin, R., & Cirranello, A. (2016). Phylogeny of Molossidae Gervais (Mammalia: Chiroptera) inferred by morphological data. Cladistics, 32(1), 2-35.|Simmons, N. B. and A. L. Cirranello (2020). https://batnames.org/explore.html. Accessed: 10 May 2020. URL: https://batnames.org/explore.html.				Guinea|Sierra Leone|Liberia|Burkina Faso?|Côte d'Ivoire|Cameroon|Central African Republic?|South Sudan|Uganda|Democratic Republic of the Congo|Kenya|Tanzania|Rwanda?	Africa	Afrotropic	LC	0	0	0	Chaerephon_bemmeleni	1	oldname match	Chaerephon_bemmeleni	0	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Mops_bemmeleni	1005147	23	Gland-tailed Free-tailed Bat	Gland-tailed Wrinkle-lipped Bat|Van Bemmelen's Wrinkle-lipped Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Molossidae	Molossinae	NA	Mops	Chaerephon	bemmeleni	Jentink	1	Nyctinomus Bemmeleni	Jentink, F.A. 1879. On a new bat, _Nyctinomus bemmeleni_, from Liberia. Notes from the Royal Zoological Museum of the Netherlands at Leyden 1:125-127.	https://www.biodiversitylibrary.org/page/9631004	RMNH.MAM.21649	holotype	https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.21649.a | https://data.biodiversitydata.nl/naturalis/specimen/RMNH.MAM.21649.b	"Liberia."			moved from Chaerephon to Mops, although further research is needed to confirm this placement	Gregorin, R., & Cirranello, A. (2016). Phylogeny of Molossidae Gervais (Mammalia: Chiroptera) inferred by morphological data. Cladistics, 32(1), 2-35.|Simmons, N. B. and A. L. Cirranello (2020). https://batnames.org/explore.html. Accessed: 10 May 2020. URL: https://batnames.org/explore.html.				Guinea|Sierra Leone|Liberia|Burkina Faso?|Cote d'Ivoire|Cameroon|Central African Republic?|South Sudan|Uganda|Democratic Republic of the Congo|Kenya|Tanzania|Rwanda?	Africa	Afrotropic	LC	0	0	0	Chaerephon_bemmeleni	1	oldname match	Chaerephon_bemmeleni	0	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Molossidae	Mops		bemmeleni	Jentink	1879	1	Notes Leyden Mus.	0.128472	Gland-tailed Free-tailed Bat	cistura Thomas, 1903.	Liberia.	Sierra Leone, Liberia, Cameroon, Sudan, Dem. Rep. Congo, Uganda, Kenya, Tanzania.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/4307/22020379/' target='_blank'>Least Concern</a>	bivittatusspecies group. Includes cistura; see Koopman (1975) and Peterson (1971). Revised by Peterson (1971).		Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Chaerephon bemmeleni; Chaerephon bemmeleni; Mops bemmeleni; Mops bemmeleni; Chaerephon bemmeleni; Mops bemmeleni; bemmeleni ; cistura; bemmeleni; cistura; cistura; bemmeleni; cisturus; Tadaride de van Bemmelen; Van-Bemmelen-Bulldogfledermaus; Caerepon de van Bemmelen; Gland-tailed Wrinkle-lipped Bat; Van Bemmelen's Wrinkle-lipped Bat; Gland-tailed Free-tailed Bat; Gland-tailed Wrinkle-lipped Bat; Van Bemmelen's Wrinkle-lipped Bat; Gland-tailed Free-tailed Bat; Gland-tailed Free-tailed Bat; M. bemmeleni