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line:xlsx:hash://sha256/181a039844a33e66a35a457b7ece741051086608e425a040051b79581d606b97!/Sheet1!/L704	application/vnd.openxmlformats-officedocument.spreadsheetml.sheet	N/A	N/A	N/A	N/A	N/A	N/A	Chaerephon atsinanana	Mops atsinanana	Mops atsinanana	Chaerephon atsinanana	Mops atsinanana	Mops atsinanana	Mops atsinanana	Mops atsinanana	Mops atsinanana		[HMW] Chaerephon atsinanana Goodman et al, 2010 , “ Madagascar : Province de Fianarantsoa , Farafangana , Colleges d’Enseignement Général (CEG) Fenoarivo , 22°49.275’S , 47°49.860’E , 10 m . ” The populations that comprise Chaerephon atsinanana were previously included in C. pumilus but have been shown to be distinct on genetic and morphological grounds. Monotypic.; [MDD2022] recently described; moved from Chaerephon to Mops, although further research is needed to confirm this placement; [IUCN] For many decades, it was cited in the literature that the widespread species Chaerephon pumilus was a common synanthropic species on Madagascar (Petersen et al . 1995). On the basis of detailed analyses, particularly with molecular data, it became clear that amongst different Afro-Malagasy populations assigned to this taxon, several different species are involved. ; Using this information, as well as some morphological characters, the population on Madagascar was named as new to science (Goodman et al. 2010).; [MDD2023] recently described; moved from Chaerephon to Mops, although further research is needed to confirm this placement; [MDD2025_2.0] recently described; moved from Chaerephon to Mops, although further research is needed to confirm this placement; [MDD2025_2.2] recently described; moved from Chaerephon to Mops, although further research is needed to confirm this placement														atsinanana	For many decades, it was cited in the literature that the widespread species Chaerephon pumilus was a common synanthropic species on Madagascar (Petersen et al . 1995). On the basis of detailed analyses, particularly with molecular data, it became clear that amongst different Afro-Malagasy populations assigned to this taxon, several different species are involved. ; Using this information, as well as some morphological characters, the population on Madagascar was named as new to science (Goodman et al. 2010).			atsinanana	atsinanana			atsinanana (S. M. Goodman, Buccas, Naidoo, F. Ratrimomanarivo, P. J. Taylor, & J. Lamb, 2010)						N/A																																								NA																											194287C9FFA2BA0EB498FA6BBB40F839	Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions	978-84-16728-19-0	hbmw_9_Molossidae_598.pdf.imf	hash://md5/e57bffb1ffbcba10b412f760b226ffce	649	zip:hash://sha256/ec5fd314a06aba1a7b0b72f23e54ac625ae272bd98f82f1d01f4c09627d9e8e0!/treatments-xml-main/data/19/42/87/194287C9FFA2BA0EB498FA6BBB40F839.xml	Chaerephon atsinanana	Molossidae	Chaerephon	atsinanana	Goodman et al.	2010	Tadaride d @fr | Atsinanana @en | Madagaskar-Bulldogfledermaus @de | Caerepon de Atsinanana @es | Eastern Madagascar Free-tailed Bat @en	Chaerephon atsinanana Goodman et al, 2010 , “ Madagascar : Province de Fianarantsoa , Farafangana , Colleges d’Enseignement Général (CEG) Fenoarivo , 22°49.275’S , 47°49.860’E , 10 m . ” The populations that comprise Chaerephon atsinanana were previously included in C. pumilus but have been shown to be distinct on genetic and morphological grounds. Monotypic.	A narrow zone along E seaboard of Madagascar , from c. 16° S to Vangaindrano (at 23° S ).	Head—body 61-67 mm, tail 27-39 mm, ear 15-18 mm, hindfoot 5-7 mm, forearm 37-42 mm; weight 9-17 g. Pelage of the Madagascar Free-tailed Bat is short, blackish brown above, brown on throat, and dark brown below, rarely with small white mid-ventral patch; typically, there is distinct whitish or beige stripe of longer hairs at flanks, as found in the Little Free-tailed Bat ( C. pumilus ), but this is variable. Upperlip is wrinkled, and ears are similar in color and shape to those of the Little Free-tailed Bat. However, antitragus is notably broad, with angular anterior edge, and terminating with rectangular-blunt tip, while in the Little Free-tailed Bat the structureis less thickset, with less angular anterior margin, more angular posterior margin, and slightly rounded tip. Anterior palate is emarginated but narrowly open, and basisphenoid pits are deep. M* has third ridge almost as long as second ridge.	Common in urban and rural settlements from near sea level up to elevations over 1100 m .	In a study in the Andasibe region, Madagascar , Free-tailed Bats fed mostly on Coleoptera , Hemiptera , Lepidoptera , and Diptera . Diptera were most prevalent in the cold season, compared to Coleoptera in the hot season.	No information.	The Madagascar Free-tailed Bat is nocturnal. Day roosts have only been located in synanthropic settings such as roofs of houses, churches, and schools. Occupied houses are usually of a typically architectural type, 4-6 m in height, with metal roof, and with or without ventilation gaps in the gables. No natural roosts of the species have been found. Presumably, before human settlement they would have roosted in crevices in rocks and cliffs in mountainous areas. According to J. M. Russ and colleagues in 2003, in Madagascar their narrow FM echolocation calls recorded in an open environment had a low peak frequency (frequency with most energy) of 28 kHz, a bandwidth of 19 kHz, and a duration of 8 milliseconds.	Madagascar Free-tailed Bats roost communally in small to medium-sized groups.	Classified as Least Concern on The IUCN Red Lust.	Bouchard (2001b) | Andrianaivoarivelo et al. (2006) | Goodman (2011, 2017i) | Goodman, Buccas et al. (2010) | Happold, M. (2013ak) | Monadjem, Taylor et al. (2010) | Russ, Bennett et al. (2003)	https://zenodo.org/record/6567816/files/figure.png	71. Madagascar Free-tailed Bat Chaerephon atsinanana French: Tadaride d/Atsinanana / German: Madagaskar-Bulldogfledermaus / Spanish: Caerepon de Atsinanana Other common names: Eastern Madagascar Free-tailed Bat Taxonomy. Chaerephon atsinanana Goodman et al, 2010 , “ Madagascar : Province de Fianarantsoa , Farafangana , Colleges d’Enseignement Général (CEG) Fenoarivo , 22°49.275’S , 47°49.860’E , 10 m . ” The populations that comprise Chaerephon atsinanana were previously included in C. pumilus but have been shown to be distinct on genetic and morphological grounds. Monotypic. Distribution. A narrow zone along E seaboard of Madagascar , from c. 16° S to Vangaindrano (at 23° S ). Descriptive notes. Head—body 61-67 mm, tail 27-39 mm, ear 15-18 mm, hindfoot 5-7 mm, forearm 37-42 mm; weight 9-17 g. Pelage of the Madagascar Free-tailed Bat is short, blackish brown above, brown on throat, and dark brown below, rarely with small white mid-ventral patch; typically, there is distinct whitish or beige stripe of longer hairs at flanks, as found in the Little Free-tailed Bat ( C. pumilus ), but this is variable. Upperlip is wrinkled, and ears are similar in color and shape to those of the Little Free-tailed Bat. However, antitragus is notably broad, with angular anterior edge, and terminating with rectangular-blunt tip, while in the Little Free-tailed Bat the structureis less thickset, with less angular anterior margin, more angular posterior margin, and slightly rounded tip. Anterior palate is emarginated but narrowly open, and basisphenoid pits are deep. M* has third ridge almost as long as second ridge. Habitat. Common in urban and rural settlements from near sea level up to elevations over 1100 m . Food and Feeding. In a study in the Andasibe region, Madagascar , Free-tailed Bats fed mostly on Coleoptera , Hemiptera , Lepidoptera , and Diptera . Diptera were most prevalent in the cold season, compared to Coleoptera in the hot season. Breeding. No information. Activity patterns. The Madagascar Free-tailed Bat is nocturnal. Day roosts have only been located in synanthropic settings such as roofs of houses, churches, and schools. Occupied houses are usually of a typically architectural type, 4-6 m in height, with metal roof, and with or without ventilation gaps in the gables. No natural roosts of the species have been found. Presumably, before human settlement they would have roosted in crevices in rocks and cliffs in mountainous areas. According to J. M. Russ and colleagues in 2003, in Madagascar their narrow FM echolocation calls recorded in an open environment had a low peak frequency (frequency with most energy) of 28 kHz, a bandwidth of 19 kHz, and a duration of 8 milliseconds. Movements, Home range and Social organization. Madagascar Free-tailed Bats roost communally in small to medium-sized groups. Status and Conservation. Classified as Least Concern on The IUCN Red Lust. Bibliography. Bouchard (2001b), Andrianaivoarivelo et al. (2006), Goodman (2011, 2017i), Goodman, Buccas et al. (2010), Happold, M. (2013ak), Monadjem, Taylor et al. (2010), Russ, Bennett et al. (2003).	Simmons, N.B. and A.L. Cirranello. 2022B. Bat Species of the World: A taxonomic and geographic database. Accessed on 10/11/2022.	Molossidae	Mops atsinanana	Mops		atsinanana	Goodman, Buccas, Naidoo, Ratrimomanirivo, Taylor & Lamb	2010	1	Zootaxa	######	Eastern Free-tailed Bat	None.		Madagascar.	Not listed.	Least Concern as Chaerephon atsinanana 		Mammal Diversity Database. (2023). Mammal Diversity Database (Version 1.11) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.7830771 released 15 April 2023	Mops atsinanana	23	Madagascar Free-tailed Bat	Eastern Madagascar Free-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	MOLOSSIDAE	MOLOSSINAE	NA	Mops	Chaerephon	atsinanana	Goodman, Buccas, Naidoo, Ratrimomanarivo, P. J. Taylor, & Lamb	2010	1	Chaerephon_atsinanana	Goodman, S. M., Buccas, W., Naidoo, T., Ratrimomanarivo, F., Taylor, P. J., & Lamb, J. (2010). Patterns of morphological and genetic variation in western Indian Ocean members of the Chaerephon 'pumilus' complex (Chiroptera: Molossidae), with the description of a new species from Madagascar. Zootaxa, 2551(1), 21.	https://www.biotaxa.org/Zootaxa/article/view/zootaxa.2551.1.1	FMNH 185259		"Madagascar: Province de Fianarantsoa, Farafangana, Collèges d'Enseignement Général (CEG) Fenoarivo, 22°49.275'S, 47°49.860'E, 10 m."	-22.82	47.831	atsinanana (Goodman, Buccas, Naidoo, Ratrimomanarivo, P. J. Taylor, & Lamb, 2010)	recently described; moved from Chaerephon to Mops, although further research is needed to confirm this placement	Goodman, S. M., Buccas, W., Naidoo, T., Ratrimomanarivo, F., Taylor, P. J., & Lamb, J. (2010). Patterns of morphological and genetic variation in western Indian Ocean members of the Chaerephon'pumilus' complex (Chiroptera: Molossidae), with the description of a new species from Madagascar. Zootaxa, 2551(1), 1-36.|Gregorin, R., & Cirranello, A. (2016). Phylogeny of Molossidae Gervais (Mammalia: Chiroptera) inferred by morphological data. Cladistics, 32(1), 2-35.|Simmons, N. B. and A. L. Cirranello (2020). https://batnames.org/explore.html. Accessed: 10 May 2020. URL: https://batnames.org/explore.html.	Madagascar	Africa	Afrotropic	LC	0	0	0	Chaerephon_atsinanana	1	unmatched	NA	1	IUCN. 2022. The IUCN Red List of Threatened Species. Version 2022-1. https://www.iucnredlist.org. Accessed on [28 September, 2022].	70000000	Chaerephon atsinanana	ANIMALIA	CHORDATA	MAMMALIA	CHIROPTERA	MOLOSSIDAE	Chaerephon	atsinanana	Goodman, Buccas, Naidoo, Ratrimomanarivo, Taylor &; Lamb, 2010	For many decades, it was cited in the literature that the widespread species Chaerephon pumilus was a common synanthropic species on Madagascar (Petersen et al . 1995). On the basis of detailed analyses, particularly with molecular data, it became clear that amongst different Afro-Malagasy populations assigned to this taxon, several different species are involved. ; Using this information, as well as some morphological characters, the population on Madagascar was named as new to science (Goodman et al. 2010).	70000000	Chaerephon atsinanana	Least Concern		2017	2016-05-02 00:00:00 UTC	3.1	English	This newly described species is known from numerous sites in the eastern portion of the island, particularly in synanthropic settings, where it can be very common (Goodman and Ramasindrazana, 2013; Goodman et al . 2010).	To date, all known day roost sites are in synanthropic settings, but it is presumed to also occur in caves and rock crevices in the eastern portions of the island. Suitable habitats are lowland and lower montane regions of eastern Madagascar, including urban areas, agricultural zones, and the forest-open area ecotone.	None known.	Not estimated, but can be common as a synanthropic species in villages and towns of eastern Madagascar. ;On the basis of inference from genetic data, this species has witnessed a recent population expansion (Lamb et al. 2012).	Increasing	Known to occur across much of the eastern length of Madagascar within the elevational range from near sea level to 1000 m (Goodman et al. , 2010).		Terrestrial	None suggested.	Afrotropical		FALSE	FALSE	Global	Simmons, N. B., & Cirranello, A. L. (2023). Batnames.org Species List Version 1.4 (1.4). Zenodo. https://doi.org/10.5281/zenodo.8136157 	Molossidae	Mops		atsinanana	Goodman, Buccas, Naidoo, Ratrimomanirivo, Taylor & Lamb	2010	1	Zootaxa	########	Eastern Free-tailed Bat	None.		Madagascar.	Not listed.	Least Concern as Chaerephon atsinanana 		Mops atsinanana	1005146	23	Madagascar Free-tailed Bat	Eastern Madagascar Free-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	CHIROPTERA	VESPERTILIONIFORMES	NA	NA	VESPERTILIONOIDEA	Molossidae	MOLOSSINAE	NA	Mops	Chaerephon	atsinanana	Goodman, Buccas, Naidoo, Ratrimomanarivo, P. J. Taylor, & Lamb	2010	1	Chaerephon_atsinanana	Goodman, S. M., Buccas, W., Naidoo, T., Ratrimomanarivo, F., Taylor, P. J., & Lamb, J. (2010). Patterns of morphological and genetic variation in western Indian Ocean members of the Chaerephon 'pumilus' complex (Chiroptera: Molossidae), with the description of a new species from Madagascar. Zootaxa, 2551(1), 21.	https://www.biotaxa.org/Zootaxa/article/view/zootaxa.2551.1.1	FMNH 185259		"Madagascar: Province de Fianarantsoa, Farafangana, Collèges d'Enseignement Général (CEG) Fenoarivo, 22°49.275'S, 47°49.860'E, 10 m."	-22.8213	47.831	atsinanana (Goodman, Buccas, Naidoo, Ratrimomanarivo, P. J. Taylor, & Lamb, 2010)	recently described; moved from Chaerephon to Mops, although further research is needed to confirm this placement	Goodman, S. M., Buccas, W., Naidoo, T., Ratrimomanarivo, F., Taylor, P. J., & Lamb, J. (2010). Patterns of morphological and genetic variation in western Indian Ocean members of the Chaerephon'pumilus' complex (Chiroptera: Molossidae), with the description of a new species from Madagascar. Zootaxa, 2551(1), 1-36.|Gregorin, R., & Cirranello, A. (2016). Phylogeny of Molossidae Gervais (Mammalia: Chiroptera) inferred by morphological data. Cladistics, 32(1), 2-35.|Simmons, N. B. and A. L. Cirranello (2020). https://batnames.org/explore.html. Accessed: 10 May 2020. URL: https://batnames.org/explore.html.				Madagascar	Africa	Afrotropic	LC	0	0	0	Chaerephon_atsinanana	1	unmatched	NA	1	Burgin, C. J., Zijlstra, J. S., Becker, M. A., Handika, H., Alston, J. M., Widness, J., Liphardt, S., Huckaby, D. G., and Upham, N. S. (2025). How many mammal species are there now? Updates and trends in taxonomic, nomenclatural, and geographic knowledge. Journal of Mammalogy in revision: TBD. https://doi.org/10.1101/2025.02.27.640393	Mops_atsinanana	1005146	23	Madagascar Free-tailed Bat	Eastern Madagascar Free-tailed Bat	Theria	Placentalia	Boreoeutheria	Laurasiatheria	Chiroptera	Yangochiroptera	NA	NA	Vespertilionoidea	Molossidae	Molossinae	NA	Mops	Chaerephon	atsinanana	S. M. Goodman, Buccas, Naidoo, F. Ratrimomanarivo, P. J. Taylor, & J. Lamb	1	Chaerephon atsinanana	Goodman, S.M., Buccas, W., Naidoo, T., Ratrimomanarivo, F., Taylor, P.J. and Lamb, J. 2010-07-28. Patterns of morphological and genetic variation in western Indian Ocean members of the _Chaerephon 'pumilus'_ complex (Chiroptera: Molossidae), with the description of a new species from Madagascar. Zootaxa 2551:1-36.	https://doi.org/10.11646/zootaxa.2551.1.1	FMNH:Mamm:185259	holotype	https://collections-zoology.fieldmuseum.org/catalogue/2684027	"Madagascar: Province de Fianarantsoa, Farafangana, Collèges d'Enseignement Général (CEG) Fenoarivo, 22°49.275'S, 47°49.860'E, 10 m."	-22.8213	47.831	recently described; moved from Chaerephon to Mops, although further research is needed to confirm this placement	Goodman, S. M., Buccas, W., Naidoo, T., Ratrimomanarivo, F., Taylor, P. J., & Lamb, J. (2010). Patterns of morphological and genetic variation in western Indian Ocean members of the Chaerephon'pumilus' complex (Chiroptera: Molossidae), with the description of a new species from Madagascar. Zootaxa, 2551(1), 1-36.|Gregorin, R., & Cirranello, A. (2016). Phylogeny of Molossidae Gervais (Mammalia: Chiroptera) inferred by morphological data. Cladistics, 32(1), 2-35.|Simmons, N. B. and A. L. Cirranello (2020). https://batnames.org/explore.html. Accessed: 10 May 2020. URL: https://batnames.org/explore.html.				Madagascar	Africa	Afrotropic	LC	0	0	0	Chaerephon_atsinanana	1	unmatched	NA	1	Simmons, N. B., & Cirranello, A. L. (2025). Batnames.org Species List Version 1.7 (1.7). Zenodo. https://doi.org/10.5281/zenodo.14796586	Molossidae	Mops		atsinanana	Goodman, Buccas, Naidoo, Ratrimomanirivo, Taylor & Lamb	2010	1	Zootaxa	########	Eastern Free-tailed Bat	None.		Madagascar.	<a href='https://cites.org/eng/app/appendices.php' target='_blank'>Not Listed</a>	<a href='https://www.iucnredlist.org/species/67360705/67360707/' target='_blank'>Least Concern</a>			Mammal Diversity Database. (2025). Mammal Diversity Database (Version 2.2) [Data set]. Zenodo. https://doi.org/10.5281/zenodo.15007505	NA	Chaerephon atsinanana; Mops atsinanana; Mops atsinanana; Chaerephon atsinanana; Mops atsinanana; atsinanana; Tadaride d; Atsinanana; Madagaskar-Bulldogfledermaus; Caerepon de Atsinanana; Eastern Madagascar Free-tailed Bat; Madagascar Free-tailed Bat; Eastern Madagascar Free-tailed Bat; Eastern Free-tailed Bat; M. atsinanana